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Sexual Di(Et)Morphism: Using Experimental Evolution To Test Whether Resource Partitioning Promotes Divergence Of The Sexes In Drosophila Melanogaster, Owen A. Jamieson 2027 Wilfrid Laurier University

Sexual Di(Et)Morphism: Using Experimental Evolution To Test Whether Resource Partitioning Promotes Divergence Of The Sexes In Drosophila Melanogaster, Owen A. Jamieson

Theses and Dissertations (Comprehensive)

Sexual dimorphism has been a long-studied phenomenon in the field of evolutionary biology. Usually, these differences in morphology, physiology and/or behaviour between the sexes are studied through the parsimonious lens of sexual selection. However, there are natural history cases where this lens does not seem applicable. Some species show sex specific morphological traits whose presence may not be due to a direct benefit to reproduction, but a difference in how each sex interacts with the environment and differing selective pressures from the environment on either sex. Darwin even brought up examples of this kind of dimorphism and pondered their existence, …


Genomic Analysis Reveals New Subspecies Of Cercyonis Scudder, 1875 (Nymphalidae: Satyrinae: Satyrini), Jing Zhang, Qian Cong, Jinhui Shen, Leina Song, Todd L. Stout, Michael S. Fisher, Robb Hannawacker, Nick V. Grishin 2026 University of Texas Southwestern Medical Center

Genomic Analysis Reveals New Subspecies Of Cercyonis Scudder, 1875 (Nymphalidae: Satyrinae: Satyrini), Jing Zhang, Qian Cong, Jinhui Shen, Leina Song, Todd L. Stout, Michael S. Fisher, Robb Hannawacker, Nick V. Grishin

The Taxonomic Report of the International Lepidoptera Survey

Genomic analysis of Cercyonis (Cercyonis) Scudder, 1875 (Nymphalidae Rafinesque, 1815) partitions the subgenus into three species groups and reveals five new subspecies from the USA (type localities in parentheses): Cercyonis (Cercyonis) pegala koosharem Stout and Grishin, ssp. n. (Utah, Sevier Co.), Cercyonis (Cercyonis) pegala lasala Grishin, M. Fisher, and Hannawacker ssp. n. (Utah, Grand Co.), Cercyonis (Cercyonis) pegala hortoni Grishin, ssp. n. (Arizona, Gila Co.), Cercyonis (Cercyonis) meadii maculana Grishin, ssp. n. (Colorado, Montezuma Co.), and Cercyonis (Cercyonis) meadii rimana Grishin, ssp. n. (Arizona, Yavapai Co.). A neotype is designated for Satyrus boopis Behr, 1864 (USA, California, Contra Costa Co.). …


Organic Pesticides: A General Overview, Marion Murray, Ann Mull 2026 Utah State University

Organic Pesticides: A General Overview, Marion Murray, Ann Mull

All Current Publications

Organic pesticides are naturally occurring, nonsynthetic, or minimally processed substances that control unwanted organisms in the environment. Categories include insecticides, herbicides, fungicides, and miticides.


Coming Soon To A Cellar Near You: Novel Range Expansion Of A Short-Bodied Cellar Spider, Physocyclus Enaulus (Araneae: Pholcidae), In A Building In Central Illinois, Usa, Ricky Gieser, Mark Davis 2026 Illinois Natural History Survey, Prairie Research Institute, University of Illinois Urbana-Champaign

Coming Soon To A Cellar Near You: Novel Range Expansion Of A Short-Bodied Cellar Spider, Physocyclus Enaulus (Araneae: Pholcidae), In A Building In Central Illinois, Usa, Ricky Gieser, Mark Davis

The Great Lakes Entomologist

A reproductive population of Physocyclus enaulus Crosby, 1926 (Pholcidae: Araneae), a short-bodied cellar spider native to the American southwest and Mexico, is documented for the first time in a single building in Illinois, far outside of their known native range. The means of introduction is unknown but is likely influenced by either accidental hitchhiking spiders or a deliberate introduction to control brown recluse populations. Cellar spiders were purchased and released in the same building to predate on brown recluse spiders, but the first P. enaulus individual was found a year prior to the release. These cellar spiders have a wide-ranging …


New Wisconsin State Records For Predatory Fungus Gnats (Diptera: Bibionomorpha: Keroplatidae), Daniel K. Young 2026 University of Wisconsin

New Wisconsin State Records For Predatory Fungus Gnats (Diptera: Bibionomorpha: Keroplatidae), Daniel K. Young

The Great Lakes Entomologist

In Wisconsin, no published literature species records for the keroplatid genera Keroplatus and Proceroplatus exist (Diptera: Bibionomorpha: Keroplatidae). New state records are presented for Keroplatus carbonarius Bosc, Keroplatus clausus Coquillett, Keroplatus militaris Johannsen and Proceroplatus elegans (Coquillett). The new records are associated largely with Malaise trap samples from Hemlock Draw (The Nature Conservancy), in the Baraboo Hills of Sauk County, and a Malaise trap at Kemp Natural Resources Station, in Oneida County, Wisconsin. Two specimens of K. clausus and a single specimen of K. militaris discovered within the research collection at the University of Wisconsin-Madison are also reported. The specimens …


First Records Of The Rare Nearctic Water Boatman Palmacorixa Janeae (Hemiptera: Corixidae) In The United States, Kurt L. Schmude, Gerald Shepard, Adam Frankiewicz 2026 University of Wisconsin, Superior

First Records Of The Rare Nearctic Water Boatman Palmacorixa Janeae (Hemiptera: Corixidae) In The United States, Kurt L. Schmude, Gerald Shepard, Adam Frankiewicz

The Great Lakes Entomologist

We report the first record of the rare water boatman, Palmacorixa janeae Brooks (Hemiptera: Corixidae), from the United States. Two hundred and forty specimens were collected in Wisconsin from Chequamegon Bay in Lake Superior. Palmacorixa gillettei Abbott, a similar species known from the area, was also identified among the samples. Most specimens, all of which were adults, were collected at night using a neuston net towed along the surface for approximately 5 minutes. The adults were found in areas that were 34 meters to 4.2 kilometers from the shore that had depths ranging from 1.0 to 5.5 meters. All representatives …


Seasonal Emergence, Egg Gallery Characteristics, And Natural Enemies Of The Eastern Ash Bark Beetle, Hylesinus Aculeatus (Coleoptera: Curculionidae: Scolytinae), In Michigan, Robert A. Haack 2026 USDA Forest Service

Seasonal Emergence, Egg Gallery Characteristics, And Natural Enemies Of The Eastern Ash Bark Beetle, Hylesinus Aculeatus (Coleoptera: Curculionidae: Scolytinae), In Michigan, Robert A. Haack

The Great Lakes Entomologist

Hylesinus aculeatus Say (Coleoptera: Curculionidae: Scolytinae) commonly infests the trunks and limbs of recently dead, dying, or cut ash (Fraxinus spp.) trees in eastern North America. In the Great Lakes region, H. aculeatus is univoltine, overwinters as adults, breeds in late spring and early summer, and constructs transverse biramous egg galleries. I sampled trunk and branch sections from three small-diameter white ash (F. americana L.) trees that were cut in April 1986 in Ingham County, Michigan, and then naturally infested by H. aculeatus and associates. Bolts cut from these trees and placed in an outdoor rearing cage in …


Phenology Of Ground-Dwelling Beetle Assemblages (Coleoptera: Carabidae, Staphylinidae) From Three Fir Stands Of Southern Québec (Canada), With Comparison Among Selected Species Between 1985 And 2006-2010, Claire Levesque, Gilles-Yvon Levesque 2026 independent

Phenology Of Ground-Dwelling Beetle Assemblages (Coleoptera: Carabidae, Staphylinidae) From Three Fir Stands Of Southern Québec (Canada), With Comparison Among Selected Species Between 1985 And 2006-2010, Claire Levesque, Gilles-Yvon Levesque

The Great Lakes Entomologist

A total of 5,303 adult epigaeic beetles (Coleoptera) were collected using pitfall traps across three balsam fir (Abies balsamea (Linnaeus) Miller) stands in southern Québec (Canada). Sampling occurred during May–September 1985 in sites representing three distinct understory vegetation types. Three dominant families (Carabidae, Leiodidae and Staphylinidae) together represented at least 93% of catches at each site. Staphylinidae were less abundant than Carabidae at two sites, whereas Carabidae and Staphylinidae were collected in similar numbers at a third stand. Associated with increased abundance and heterogeneity of the understory vegetation, we observed 1) the decrease of specialist forest carabids, often active …


Caddisfly (Trichoptera) Assemblages From The Southern Laurentian Highlands, Québec (Canada), David C. Houghton 2026 Hillsdale College

Caddisfly (Trichoptera) Assemblages From The Southern Laurentian Highlands, Québec (Canada), David C. Houghton

The Great Lakes Entomologist

The caddisflies (Trichoptera) represent an important taxon for the natural functioning of aquatic ecosystems and as biological monitoring organisms; however, basic information on their assemblages is still lacking in Canada. In this study, adult caddisflies from 30 lakes and streams in the southern Laurentian Highlands of Québec were sampled during 2023–2025. A total of 186 caddisfly species were determined, including 22 species not previously reported from Québec and eight not previously reported from anywhere in Canada. Two species of Cernotina also represented the first collection of that genus within the province. Using Indicator Species Analysis, six species indicated small rivers, …


Paper Wasp Species Polistes Fuscatus (Hymenoptera: Vespidae) Tracks Urban Lead (Pb) Levels More Than Polistes Dominula, Lindsey R. Kemmerling, Anna L. Lovat, Allison A. Hanka, Emilie C. Snell-Rood 2026 University of Minnesota, Twin Cities

Paper Wasp Species Polistes Fuscatus (Hymenoptera: Vespidae) Tracks Urban Lead (Pb) Levels More Than Polistes Dominula, Lindsey R. Kemmerling, Anna L. Lovat, Allison A. Hanka, Emilie C. Snell-Rood

The Great Lakes Entomologist

Lead (Pb) is a persistent, toxic pollutant with widespread but inequitably distributed impacts, especially for humans in urban areas. The impacts of lead pollution on other organisms is largely unknown, even as they commonly experience exposure to lead alongside humans. We tested hypotheses regarding the lead concentrations of paper wasps across a gradient of pollution relative to soil and human lead levels, and correlations with wasp body condition. We collected two species of paper wasp across an urban gradient of lead pollution: Polistes fuscatus (Fabricius) (Hymenoptera: Vespidae), native to our study location, and Polistes dominula (Christ), nonnative to our study …


Long-Term Monitoring And Response Of Papaipema Sciata Bird (Lepidoptera: Noctuidae) To Prescribed Fire In Michigan (Usa), Gregory J. Norwood, Logan M. Rowe, Brett A. DeGregorio, David L. Cuthrell, Aaron D. Meredith, Steven M. Gray 2026 Michigan Department of Natural Resources, State Park Stewardship Unit

Long-Term Monitoring And Response Of Papaipema Sciata Bird (Lepidoptera: Noctuidae) To Prescribed Fire In Michigan (Usa), Gregory J. Norwood, Logan M. Rowe, Brett A. Degregorio, David L. Cuthrell, Aaron D. Meredith, Steven M. Gray

The Great Lakes Entomologist

The Culver’s root borer moth (Papaipema sciata) (Lepidoptera: Noctuidae) is a rare Noctuid that requires Culver’s root (Veronicastrum virginicum) to complete its lifecycle within remnant native grasslands and open woodlands in the midwest and northeast United States. We report on 26 years of monitoring this species’ colonization of Culver’s root stems at Algonac State Park (ASP) in Michigan, where prairies have been maintained by prescribed fire since 1990. P. sciata is considered fire sensitive because eggs spend the dormant season in the combustible prairie litter and duff near host plants. The objectives were to determine the …


Could The European Thanasimus Formicarius Outcompete The North American Thanasimus Dubius (Coleoptera: Cleridae) If Introduced To North America?, Robert A. Haack, Robert K. Lawrence 2026 USDA Forest Service

Could The European Thanasimus Formicarius Outcompete The North American Thanasimus Dubius (Coleoptera: Cleridae) If Introduced To North America?, Robert A. Haack, Robert K. Lawrence

The Great Lakes Entomologist

A federal quarantine was enacted in the United States (US) in the fall of 1992 in response to discovery of the Eurasian pine shoot beetle, Tomicus piniperda (L.) (Coleoptera: Scolytinae), in July 1992 in Ohio. The quarantine regulated movement of pine products (e.g., Christmas trees, nursery stock, and logs) at the county level from infested to uninfested US states. A classical biological program was considered for T. piniperda starting in 1995, focusing on the Eurasian clerid beetle Thanasimus formicarius (L.), an important predator of T. piniperda. Concerns were raised in 1996 that if T. formicarius were released in the …


Fecundity Of Thanasimus Dubius (Coleoptera: Cleridae) Rebounds After A Period Of Prey Scarcity, Robert A. Haack, Robert K. Lawrence 2026 USDA Forest Service

Fecundity Of Thanasimus Dubius (Coleoptera: Cleridae) Rebounds After A Period Of Prey Scarcity, Robert A. Haack, Robert K. Lawrence

The Great Lakes Entomologist

We field-collected Thanasimus dubius (Fabricius) (Coleoptera: Cleridae) adults during spring, maintained them separately in the laboratory, and initially gave them four bark beetle adults [Ips grandicollis (Eichhoff) or Tomicus piniperda (L.) (Coleoptera: Curculionidae: Scolytinae)] daily as prey. Using a subset of the female clerids (N = 41), we assigned them to one of four feeding groups that varied in the number of prey provided daily. For the first four weeks of a 12–week study, the four diets were: four prey daily, two prey daily, one prey daily, and one prey every 2 days. After four weeks, all females were …


Distribution And Status Of Silphium Herb Gall Wasps (Cynipoidea: Aulacideini: Antistrophus) And Their Associated Endophytic Communities In Wisconsin, Richard A. Henderson, Louis F. Nastasi, Jay Watson, Scott Sauer 2026 Research Ecologist (retired), Wisconsin Department of Natural Resources, Office of Applied Science

Distribution And Status Of Silphium Herb Gall Wasps (Cynipoidea: Aulacideini: Antistrophus) And Their Associated Endophytic Communities In Wisconsin, Richard A. Henderson, Louis F. Nastasi, Jay Watson, Scott Sauer

The Great Lakes Entomologist

At least six described species of the herb gall wasp genus Antistrophus Walsh (Hymenoptera: Cynipoidea: Aulacideini) are dependent on the plant genus Silphium L., a plant group emblematic of now-endangered Midwestern tallgrass prairies. The distribution and conservation status of Antistrophus gall wasps is mostly a mystery, and these insects have only recently been recorded from Wisconsin. We surveyed the state for Antistrophus gall wasps on four Silphium species, whole-leaf rosinweed (S. integrifolium Michx.), compass plant (S. laciniatum L.), cup plant (S. perfoliatum L.), and prairie dock (S. terebinthinaceum Jacq.) to determine their distribution and abundance in Wisconsin. We report new …


Cover Art For Tgle Vol. 59 Nos. 1 & 2, 2026 Valparaiso University

Cover Art For Tgle Vol. 59 Nos. 1 & 2

The Great Lakes Entomologist

Cover Art for TGLE Vol. 59 Nos. 1 & 2


Cover Pages For Tgle Vol. 59 Nos. 1 & 2, 2026 Valparaiso University

Cover Pages For Tgle Vol. 59 Nos. 1 & 2

The Great Lakes Entomologist

Cover Pages for TGLE Vol. 59 Nos. 1 & 2


Full Issue For Tgle Vol. 59 Nos. 1 & 2, 2026 Valparaiso University

Full Issue For Tgle Vol. 59 Nos. 1 & 2

The Great Lakes Entomologist

Full issue for TGLE Vol. 59 Nos. 1 & 2


Spider Biodiversity And Insecticide Sensitivity Across Active And Restored Cranberry Bogs On Nantucket, Tyler J. Kulak 2026 University of Massachusetts Boston

Spider Biodiversity And Insecticide Sensitivity Across Active And Restored Cranberry Bogs On Nantucket, Tyler J. Kulak

Graduate Masters Theses

Spiders are one of the most diverse groups of organisms on the planet with over 53,000 species described as of 2026. They provide a suite of ecosystem services including their regulation of problematic pests in agricultural systems, appearing in high biomasses across a breadth of ecological niches. While beneficial as pest regulators, they are also susceptible to the same management techniques as pest organisms and are especially vulnerable to chemical control. Cranberry production in Massachusetts likely benefits from spider ecosystem services, but diversity research on spiders in these systems is limited. This study compares the diversity of spider populations from …


American Plum Borer In Utah Orchards (Euzophera Semifuneralis), Emilie Demard, Marion Murray, Claudia Nischwitz 2026 Utah State University

American Plum Borer In Utah Orchards (Euzophera Semifuneralis), Emilie Demard, Marion Murray, Claudia Nischwitz

All Current Publications

This fact sheet provides information on American plum borer in Utah orchards, also known as APB. This pest primarily damages fruit trees. APB moths were detected in traps in 2026, but to date, no tree damage has been found. This fact sheet discusses hosts, identification, and management methods.


Clara S. Ludlow: Mississippi A & M Graduate And American Military Entomology Pioneer, Bradley Brazzeal 2026 Mississippi State University

Clara S. Ludlow: Mississippi A & M Graduate And American Military Entomology Pioneer, Bradley Brazzeal

Midsouth Entomologist

The systematic study of the mosquitoes of the world did not begin until the early years of the 20th century. Among the early entomologists to specialize in the field was Clara S. Ludlow, who graduated from Mississippi Agricultural and Mechanical College. Her research began in the Philippines in 1901, while American troops were still fighting in the Philippine-American War, and for over 20 years she researched mosquitoes in connection with the U.S. Army. This article explores Ludlow’s scientific career and the legacy of her research.


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