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Articles 5251 - 5280 of 6307
Full-Text Articles in Terrestrial and Aquatic Ecology
Leadership In Wolf, Canis Lupus, Packs, L. David Mech
Leadership In Wolf, Canis Lupus, Packs, L. David Mech
United States Geological Survey, Northern Prairie Wildlife Research Center: Publications
I examine leadership in Wolf (Callis lupus) packs based on published observations and data gathered during summers from 1986 to 1998 studying a free-ranging pack of Wolves on Ellesmere Island that were habituated to my presence. The breeding male tended to initiate activities associated with foraging and travel, and the breeding female to initiate, and predominate in, pup care and protection. However, there was considerable overlap and interaction during these activities such that leadership could be considered a joint function. In packs with multiple breeders, quantitative information about leadership is needed.
Do Wolves Affect White-Tailed Buck Harvest In Northeastern Minnesota?, L. David Mech, Michael E. Nelson
Do Wolves Affect White-Tailed Buck Harvest In Northeastern Minnesota?, L. David Mech, Michael E. Nelson
United States Geological Survey, Northern Prairie Wildlife Research Center: Publications
We used simple linear regression to analyze 8-23 years of data on a wolf (Canis lupus) population and human harvest of white-tailed deer (Odocoileus virginianus) bucks in northeastern Minnesota to determine any effects of wolves on buck harvesting. Over the long term, wolves accounted for at least 14-22% inter-year variation in buck harvest in the region, but an unknown amount of variation in hunter effort have obscured any more precise estimate. For part of the area with poorest habitat, we found strong relationships (r2 = 0.66-0.84) between annual wolf numbers and buck harvests from 1988 …
The Relationship Of Animal Protection Interests To Animal Damage Management: Historic Paths, Contemporary Concerns And The Uncertain Future, John Hadidian
Conservation Biology and Animal Welfare Collection
More than a decade ago Schmidt (1989) called for consideration of animal welfare to become a "firstorder" decision rule in wildlife management concerns, including animal damage control. Although there has been movement in that direction, this clearly has not yet come to pass. This paper takes a brief look at the interests we call animal damage management, animal welfare and protection, animal rights, and environmentalism in order to speculate about their shared concerns and the uncertain future before them. Since animal damage and the management of that damage cannot be abstracted from the environmental context in which they occur, this …
Dc Birdscape: A Program For Monitoring Neotropical Migratory Birds In Washington, Dc, John Sauer, John Hadidian, Sam Droege, Paul Handly, Carolyn Williams, Christopher Swarth, George Didden, Jane Huff
Dc Birdscape: A Program For Monitoring Neotropical Migratory Birds In Washington, Dc, John Sauer, John Hadidian, Sam Droege, Paul Handly, Carolyn Williams, Christopher Swarth, George Didden, Jane Huff
Biogeography and Ecological Opportunity Collection
Urban and suburban habitats often contain a variety of Neotropical migratory birds, but are poorly sampled by programs such as the North American Breeding Bird Survey. DC Birdscape was developed to inventory and monitor birds in Washington, DC. Birds were surveyed using a systematic sample of point counts during 1993-1995. Results indicate that species richness of Neotropical migratory birds varied among land-use categories, and that maximum species richness occurred in parkland habitats. Although DC Birdscape has provided relevant information on bird distribution and species richness, it is unclear whether the information is of sufficient management interest to support its continuation …
Assessing Factors That May Predispose Minnesota Farms To Wolf Depredations On Cattle, L. David Mech, Elizabeth K. Harper, Thomas J. Meier, William J. Paul
Assessing Factors That May Predispose Minnesota Farms To Wolf Depredations On Cattle, L. David Mech, Elizabeth K. Harper, Thomas J. Meier, William J. Paul
United States Geological Survey, Northern Prairie Wildlife Research Center: Publications
Wolf (Canis lupus) depredations on livestock cause considerable conflict and expense in Minnesota. Furthermore, claims are made that such depredations are fostered by the type of animal husbandry practiced. Thus, we tried to detect factors that might predispose farms in Minnesota to wolf depredations. We compared results of interviews with 41 cattle farmers experiencing chronic cattle losses to wolves (chronic farms) with results from 41 nearby "matched" farms with no wolf losses to determine farm characteristics or husbandry practices that differed and that therefore might have affected wolf depredations. We also used a Geographic Information System (GIS) to …
A New Elfin Butterfly (Lycaenidae: Eumaeini) From Northern China With Comments On The Nomenclature Of Palaearctic Elfins, Kurt Johnson
A New Elfin Butterfly (Lycaenidae: Eumaeini) From Northern China With Comments On The Nomenclature Of Palaearctic Elfins, Kurt Johnson
The Taxonomic Report of the International Lepidoptera Survey
Ahlbergia hsui, new species, is described from two specimens recently collected in China’s Nan Shan area of endemism located along the boundary of northern China with Mongolia. In discussing the new species, the historical literature concerning Palaearctic elfin butterflies is reviewed and compared to nomenclatorial usages in D’Abrera’s widely disseminated 1993 treatment of Palaearctic butterflies; 25 errors or omissions in the latter treatment are corrected.
Mortality Of Adult White-Tailed Deer On Fort Chaffee, Arkansas, Gregory G. Humphreys, Thomas A. Nelson
Mortality Of Adult White-Tailed Deer On Fort Chaffee, Arkansas, Gregory G. Humphreys, Thomas A. Nelson
Journal of the Arkansas Academy of Science
We conducted a telemetry study on Fort Chaffee, Arkansas, to estimate the extent, timing, and causes of mortality among resident adult white-tailed deer {Odocoileus virginianus). Twenty-seven deer were captured, radio-collared and monitored for 1 yr to assess seasonal mortality. Annual mortality rates differed considerably between sexes, with males exhibiting a much higher rate (86.1%) than females (5.3%). The primary causes of death among males were poaching, predation by coyotes, and legal harvest. Legal harvest by a gun-hunter accounted for the only female mortality. No mortalities were attributed to military exercises, which occurred frequently on the study area. Our results suggest …
Feeding Relationship Between Two Synoptic, Morphologically Similar Fishes, The Western Mosquitofish (Gambusia Affinis) And The Blackspotted Topminnow (Fundulus Olivaceus), Allyson R. Neely, Edmund J. Pert
Feeding Relationship Between Two Synoptic, Morphologically Similar Fishes, The Western Mosquitofish (Gambusia Affinis) And The Blackspotted Topminnow (Fundulus Olivaceus), Allyson R. Neely, Edmund J. Pert
Journal of the Arkansas Academy of Science
We sampled fish during the summer of 1999, in Caney Bayou on the campus of the University of Arkansas at Pine Bluff, Jefferson County, in order to study the feeding relationship between the western mosquitofish (Gambusia ajfinis) and the blackspotted topminnow (Fundulus olivaceus). We identified and enumerated the diet items of 56 blackspotted topminnows and 28 mosquitofish that were captured using electroshocking, seining, and dipnetting. Pooled diet items for each species were analyzed using a variety of approaches: percent of diet composition, frequency of occurrence, Levins' measure of diet breadth, and Morisita's measure of diet overlap. Twenty-one percent of mosquitofish …
Characteristics Of Some Fruiting Plant Species In Northwest Arkansas, And The Avian Assemblages That Feed On Them, John W. Prather, Kimberly G. Smith, Michael A. Mlodinow, Cecilia M. Riley
Characteristics Of Some Fruiting Plant Species In Northwest Arkansas, And The Avian Assemblages That Feed On Them, John W. Prather, Kimberly G. Smith, Michael A. Mlodinow, Cecilia M. Riley
Journal of the Arkansas Academy of Science
Fruits continue to be recognized as an important food source for birds in temperate areas, particularly during the fall migration period. More than 20 species of plants producing fleshy fruits are found in the Arkansas Ozarks. However, only a few of these appear to be important resources for birds during the fall migration period (August - October). Among those are sassafras (Sassafras albidum), gray-backed grape (Vitis cinerea), black cherry (Prunus serotina), hercules club (Araliaspinosa) and pokeweed (Phytolacca americana). Over the past 4 years, we have documented the physical and nutritional characteristics of those fruits and taken observational data on the …
Record Of The Lake Sturgeon, Acipenser Fulvescens Rafinesque, From The Caddo River (Ouachita River Drainage), Arkansas, Betty G. Crump, Henry W. Robison
Record Of The Lake Sturgeon, Acipenser Fulvescens Rafinesque, From The Caddo River (Ouachita River Drainage), Arkansas, Betty G. Crump, Henry W. Robison
Journal of the Arkansas Academy of Science
No abstract provided.
Impacts Of White-Tailed Deer Grazing On Spring Wildflower Communities, Faye Frankland
Impacts Of White-Tailed Deer Grazing On Spring Wildflower Communities, Faye Frankland
Masters Theses
High deer densities on state forests and nature preserves throughout Illinois have prompted concerns that deer are impacting the growth, survival, and reproduction of spring-flowering herbaceous plants. I investigated the impacts of white-tailed deer (Odocoileus virginianus) on the herbaceous vegetation, particularly spring wildflowers, at Beall Woods Nature Preserve (BWNP). BWNP was selected as the site for this study because it supports a diverse community of spring wildflowers and a large deer population. The objectives for this study were to: (1) quantify the extent of deer foraging on representative spring-flowering plant species at Beall Woods Nature Preserve, (2) investigate the effects …
Effects Of Leafy Spurge (Euphorbia Esula) Infestation On Breeding Birds Of The Sheyenne National Grassland, Nd, Daniel M. Scheiman
Effects Of Leafy Spurge (Euphorbia Esula) Infestation On Breeding Birds Of The Sheyenne National Grassland, Nd, Daniel M. Scheiman
Masters Theses
Leafy spurge (Euphorbia esula) is an exotic invasive weed in the northern Great Plains. We examined the effects of leafy spurge infestation on densities and nest success of breeding birds in grasslands on the Sheyenne National Grassland (SNG), ND. We categorized spurge-infested grasslands into three levels of infestation, based on the area covered by spurge patches: (a) low (0-20%), (b) medium (20-60%) and, (c) high (> 60%). We surveyed 60 100-m radius circular plots (20 in each category), and searched for nests in three 16-ha plots (one in each category). There were no statistically significant differences in mean species richness …
Characterization And Improvement Of Eos Land Products Using Measurements At Ameriflux Grassland And Wheat Sites In The Arm/Cart Region: Research Annual Performance Report For Period March 1, 2000- February 28, 2001, E. A. Walter-Shea, S. B. Verma
Characterization And Improvement Of Eos Land Products Using Measurements At Ameriflux Grassland And Wheat Sites In The Arm/Cart Region: Research Annual Performance Report For Period March 1, 2000- February 28, 2001, E. A. Walter-Shea, S. B. Verma
School of Natural Resources: Documents and Reviews
No abstract provided.
Proximity Of White-Tailed Deer, Odocoileus Virginianus, Ranges To Wolf, Canis Lupus, Pack Homesites, Michael E. Nelson, L. David Mech
Proximity Of White-Tailed Deer, Odocoileus Virginianus, Ranges To Wolf, Canis Lupus, Pack Homesites, Michael E. Nelson, L. David Mech
United States Geological Survey, Northern Prairie Wildlife Research Center: Publications
Seven adult female White-tailed Deer (Odocoileus virginianus) in northeastern Minnesota lived within 1.8 km of Wolf pack (Canis lupus) homesites without vacating their home ranges. Six of these deer and at least three of their fawns survived through the Wolf homesite period.
Growth Curves Of Four Species Of Commercially Valuable Freshwater Mussels (Bivalva: Unionidae), Alan D. Christian, Chris L. Davidson, William R. Posey Ii, Peter J. Rust, Jerry L. Farris, John L. Harris, George L. Harp
Growth Curves Of Four Species Of Commercially Valuable Freshwater Mussels (Bivalva: Unionidae), Alan D. Christian, Chris L. Davidson, William R. Posey Ii, Peter J. Rust, Jerry L. Farris, John L. Harris, George L. Harp
Journal of the Arkansas Academy of Science
North American freshwater mussels (Bivalvia: Unionidae) have been exploited commercially for over 100 years and have been regulated using shell size limits and/or harvest seasons. Presently, freshwater mussels are considered a threatened faunal group in North America due to the large numbers of endangered, threatened or special concern species. Therefore, management of this fauna should emphasize their long-term sustainability. The objectives of this study were 1) to construct von Bertalanffy growth curves for selected "commercially-most-valuable" species, Fusconaia ebena, Megalonaias nervosa, Amblema plicata and Quadrula quadrula, from five rivers and two reservoirs, 2) to compare species-specific von Bertalanffy growth curves from …
Curly-Tail Malformity In Hatchlings Of The Alligator Snapping Turtle, Macroclemys Temminckii (Testudines: Chelydridae) In Arkansas, Malcolm L. Mccallum, Stanley E. Trauth
Curly-Tail Malformity In Hatchlings Of The Alligator Snapping Turtle, Macroclemys Temminckii (Testudines: Chelydridae) In Arkansas, Malcolm L. Mccallum, Stanley E. Trauth
Journal of the Arkansas Academy of Science
No abstract provided.
Breeding Mortality In The Wood Frog, Rana Sylvatica (Anura: Ranidae), From Northcentral Arkansas, Stanley E. Trauth, Malcolm L. Mccallum, Michael E. Cartwright
Breeding Mortality In The Wood Frog, Rana Sylvatica (Anura: Ranidae), From Northcentral Arkansas, Stanley E. Trauth, Malcolm L. Mccallum, Michael E. Cartwright
Journal of the Arkansas Academy of Science
No abstract provided.
Home Ranges And Movements Of Adult Deer On Fort Chaffee, Arkansas, Gregory G. Humphreys, Thomas A. Nelson
Home Ranges And Movements Of Adult Deer On Fort Chaffee, Arkansas, Gregory G. Humphreys, Thomas A. Nelson
Journal of the Arkansas Academy of Science
We radio-tracked 27 white-tailed deer (Odocoileus virginianus) weekly for one year on Fort Chaffee, Arkansas, to investigate their seasonal home ranges and movements between hunted areas and refuges on this military base. This work resulted in2,123 separate radiolocations, of which 85% (1,799) were suitable for use in home range analyses. We used the McPAAL computer package to estimate home range using the Harmonic Mean and minimum convex polygon (MCP) methods. Harmonic mean estimates were based on 95% contour lines. Home range size differed between the sexes and methods. Male home ranges were larger than those of females (t= 3.32, P<0.01; harmonic mean) (t=2.07, P<0.05; MCP). Average home range sizes for males and females based on the harmonic mean method were estimated to be 483 ha and 181 ha, respectively, whereas home range estimates for males and females using the MCP method were 636 ha and 289 ha, respectively. The average home range size for all deer was 259 ha (harmonic mean) and 379 ha (MCP). We found no evidence that females restricted their home ranges during the fawning period. However, females' home ranges expanded during the breeding season, perhaps to find mates. Few deer moved to refuge areas that were off-limits to hunters during the hunting season.
Winter Breeding As A Common Occurrence In The Ringed Salamander, Ambystoma Annulatum (Caudata: Ambystomatidae), In The Ozark National Forest Of Northcentral Arkansas, Stanley E. Trauth
Winter Breeding As A Common Occurrence In The Ringed Salamander, Ambystoma Annulatum (Caudata: Ambystomatidae), In The Ozark National Forest Of Northcentral Arkansas, Stanley E. Trauth
Journal of the Arkansas Academy of Science
No abstract provided.
Salinity And Shade Preferences Result In Ovipositional Differences Between Sympatric Tiger Beetle Species, W. Wyatt Hoback, Douglas A. Golick, Tina Marie Svatos, Stephen M. Spomer, Leon G. Higley
Salinity And Shade Preferences Result In Ovipositional Differences Between Sympatric Tiger Beetle Species, W. Wyatt Hoback, Douglas A. Golick, Tina Marie Svatos, Stephen M. Spomer, Leon G. Higley
Department of Entomology: Faculty Publications
1. Adult tiger beetles of the genus Cicindela often co-occur within a habitat but larvae do not. Larvae are sedentary and form usually permanent burrows at the site of oviposition where they require 1-3 years for development.
2. To test niche partitioning based on ovipositional preference, the behavior of two sympatric salt marsh tiger beetles, Cicindela circumpicta and C. togata (Coleoptera: Cicindelidae), were examined.
3. In laboratory studies, female C. circumpicta and C. togata distinguished between experimental salinities, with the former preferring 4 parts per thousand (ppt) and the latter preferring 12 ppt. In the field, C. circumpicta larvae were …
A Record Large Wolf, Canis Lupus, Pack In Minnesota, L. David Mech
A Record Large Wolf, Canis Lupus, Pack In Minnesota, L. David Mech
United States Geological Survey, Northern Prairie Wildlife Research Center: Publications
This report documents a pack of 22-23 Wolves (Canis lupus) in central Minnesota. This is larger than the largest pack previously observed on the mainland in the midwestern U.s. during 650 wolf pack-years. Because this record-large pack preyed on White-tailed Deer (Odocoileus virginianus), one of the Wolfs smaller prey, it is evidence that pack size and prey size are not tightly related. It also indicates the size that Wolf packs can attain in the area if fully protected from human persecution.
Lack Of Reproduction In Muskoxen And Arctic Hares Caused By Early Winter?, L. David Mech
Lack Of Reproduction In Muskoxen And Arctic Hares Caused By Early Winter?, L. David Mech
United States Geological Survey, Northern Prairie Wildlife Research Center: Publications
A lack of young muskoxen (Ovibos moschatus) and arctic hares (Lepus arcticus) in the Eureka area of Ellesmere Island, Northwest Territories (now Nunavut), Canada, was observed during summer 1998, in contrast to most other years since 1986. Evidence of malnourished muskoxen was also found. Early winter weather and a consequent 50% reduction of the 1997 summer replenishment period appeared to be the most likely cause, giving rise to a new hypothesis about conditions that might cause adverse demographic effects in arctic herbivores.
Durant l’été 1998, et ce, à la différence de la plupart des années depuis …
Accuracy And Precision Of Estimating Age Of Gray Wolves By Tooth Wear, Philip S. Gipson, Warren B. Ballard, Ronald M. Nowak, L. David Mech
Accuracy And Precision Of Estimating Age Of Gray Wolves By Tooth Wear, Philip S. Gipson, Warren B. Ballard, Ronald M. Nowak, L. David Mech
United States Geological Survey, Northern Prairie Wildlife Research Center: Publications
We evaluated the accuracy and precision of tooth wear for aging gray wolves (Canis lupus) from Alaska, Minnesota, and Ontario based on 47 known-age or known-minimum-age skuIIs. Estimates of age using tooth wear and a commercial cementum annuli-aging service were useful for wolves up to 14 years old. The precision of estimates from cementum annuli was greater than estimates from tooth wear, but tooth wear estimates are more applicable in the field. We tended to overestimate age by 1-2 years and occasionaIIy by 3 or 4 years. The commercial service aged young wolves with cementum annuli to within …
Prolonged Winter Undernutrition And The Interpretation Of Urinary Allantoin:Creatinine Ratios In White-Tailed Deer, Glenn D. Delgiudice, Ken D. Kerr, L. David Mech, Ulysses S. Seal
Prolonged Winter Undernutrition And The Interpretation Of Urinary Allantoin:Creatinine Ratios In White-Tailed Deer, Glenn D. Delgiudice, Ken D. Kerr, L. David Mech, Ulysses S. Seal
United States Geological Survey, Northern Prairie Wildlife Research Center: Publications
The urinary allantoin:creatinine (A:C) ratio (expressed in micromoles of allantoin to micromoles of creatinine) has shown potential as an index of recent winter energy intake in preliminary controlled studies of elk (Cervus elaphus) involving mild condition deterioration (up to 11% loss of body mass). To ensure reliable nutritional assessments of free-ranging cervids by measuring A:C ratios of urine in snow, it is essential to extend this work. We assessed the effect of moderate and severe winter nutritional restriction on urinary A:C ratios of captive white-tailed deer (Odocoileus virginianus) that lost up to 32% body mass and …
Clonal Growth Of Lithospermum Caroliniense (Boraginaceae) In Contrasting Sand Dune Habitats, Stephen G. Weller, Kathleen H. Keeler, Barbara A. Thomson
Clonal Growth Of Lithospermum Caroliniense (Boraginaceae) In Contrasting Sand Dune Habitats, Stephen G. Weller, Kathleen H. Keeler, Barbara A. Thomson
School of Biological Sciences: Faculty Publications
The occurrence of clonal growth of distylous Lithospermum caroliniense was investigated in a population in the Nebraska Sandhills, an area where sand dunes have been relatively stable for at least 1,500–3,000 yr, and compared to a population occurring at the Indiana Dunes, an area of active sand dune formation. Spatial autocorrelation analysis indicated the occurrence of significant clonal propagation of genetically based floral morphs at Arapaho Prairie, but not for the Indiana Dunes. Apparent clonal growth in the Sandhills population had no overall negative effect on pollen deposition or fecundity relative to the Indiana population, although in some large clones …
Prairie Analysis At Prairie Ridge State Natural Area, Jasper County, Il, Annette Kessler
Prairie Analysis At Prairie Ridge State Natural Area, Jasper County, Il, Annette Kessler
Masters Theses
Five restored prairie sites at Prairie Ridge State Natural Area in Jasper County were studied during the growing season of 1998. These sites varied in age from two years to thirty years and in size from 4 acres to 10 acres. Key areas of data recorded were an overall and site specific vascular species lists, phenology of various species, and cover analysis of each site. Frequency cover and importance values of the taxa were determined using randomly located quadrats along line transects. Prairie quality of each site was determined, as well as a similarity index that compared the sites to …
Origin And Fate Of Organic Matter In South-West Australian Wetlands, Darren Stuart Ryder
Origin And Fate Of Organic Matter In South-West Australian Wetlands, Darren Stuart Ryder
Theses: Doctorates and Masters
The development and local distribution of organic soils in Australia have been poorly documented. Within Western Australia, conditions conducive to the accumulation of organic matter are geographically restricted and generally occur in coastal and/or forested landscapes. An extensive system of wetlands with peal soils occurs in the Muir-Unicup region in the far south west of Western Australia. Bokarup Swamp, Kodjinup Swamp and Noobijup Lake are representative of the wetlands occurring in this region. They arc shallow (
The Role Of Benthic Macroalgae In Sediment-Water Nutrient Cycling In The Swan-Canning Estuarine System, Western Australia, Helen Lee Astill
The Role Of Benthic Macroalgae In Sediment-Water Nutrient Cycling In The Swan-Canning Estuarine System, Western Australia, Helen Lee Astill
Theses: Doctorates and Masters
This study documented the macroalgal assemblages of the Swan-Canning Estuarine System (SCES) over a two year period, and the influences of several environmental parameters on the assemblages. In addition, the Impacts of unattached macroalgal accumulations on benthic nutrient fluxes and microbial communities were investigated. Benthic macroalgal assemblages and physico-chemical regimes were monitored in the SCES, to determine temporal and spatial changes in macroalgal communities and the influence of environmental factors in these changes. Physico-chemical regimes demonstrated strong seasonal changes, which revolved around the onset and cessation of freshwater flows in winter (May to September). In the months after freshwater flows, …
Changes To The Distribution Of Posidonia Seagrass Communities Of James Point, In Response To The Development Of Cockburn Sound, Western Australia, Anthony Muscara
Changes To The Distribution Of Posidonia Seagrass Communities Of James Point, In Response To The Development Of Cockburn Sound, Western Australia, Anthony Muscara
Theses: Doctorates and Masters
Australia’s oceans provide many economic and environmental benefits both nationally and regionally, and are of particular social, recreational and cultural importance. Western Australia's most intensively used marine embayment is Cockburn Sound, is Cockburn Sound, it supports one of the most extensive Posidoni seagrass communities in Western Australia. The protected coastal waters off the southern metropolitan coastline of Perth arc utilised intensively for industrial, commercial and recreational purposes. Over the past 50 years, wastes have been routinely discharged into Cockburn Sound causing extensive phytoplankton and epiphyte blooms, particularly during the 1960's and 1970's. The development of industrial and naval facilities and …
Three New Hesperioidae (Hesperiinae) From South Carolina: New Subspecies Of Euphyes Bimacula, Poanes Aaroni, And Hesperia Attalus, Ronald R. Gatrelle
Three New Hesperioidae (Hesperiinae) From South Carolina: New Subspecies Of Euphyes Bimacula, Poanes Aaroni, And Hesperia Attalus, Ronald R. Gatrelle
The Taxonomic Report of the International Lepidoptera Survey
Euphyes bimacula arbogasti is described as a new subspecies from Berkeley County, South Carolina, United States. It is known from only a few widely scattered colonies in the coastal swamp forests of the southeastern United States from Georgia to southeastern North Carolina. It is darker then E. b. bimacula and E. b. illinois. Poanes aaroni minimus is described as a new subspecies from Bull Swamp, Orangeburg County, South Carolina. This unique inland subspecies is presently known only from the type locality. It is darker then P. a. aaroni and P. a. …