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Full-Text Articles in Terrestrial and Aquatic Ecology

Beyond Better Conditions: Seaweed Farming And The Number Of Sentient Animals In Aquaculture, Shrilaxmi Patil, Matthew Unerman, Wasseem Emam Aug 2026

Beyond Better Conditions: Seaweed Farming And The Number Of Sentient Animals In Aquaculture, Shrilaxmi Patil, Matthew Unerman, Wasseem Emam

Animal Sentience

Oldfield & Bonano (2024) (O&B) document how far  fish welfare research lags behind welfare research in other animals. We share O&B’s feelings-based concept of welfare and agree that aquaculture seeded this field; but research volume has not scaled with animal numbers: Welfare attention per farmed fish or decapod remains negligible. Along the lines of preceding commentaries (Maia 2024, Mather 2024), we treat farmed aquatic animals as individuals rather than tonnage. We then raise a point so far absent from the commentary on O&B so far: reducing the number of sentient animals entering intensive systems. Coastal seaweed farming, which cultivates non-sentient …


Dimensional Welfare Beyond Pain: Extending The Precautionary Principle In Birch’S “The Edge Of Sentience”, Willa M. Lane, Nicola S. Clayton Jun 2026

Dimensional Welfare Beyond Pain: Extending The Precautionary Principle In Birch’S “The Edge Of Sentience”, Willa M. Lane, Nicola S. Clayton

Animal Sentience

Birch's (2024) The Edge of Sentience identifies sentience candidates primarily through the pain-focused LSE Criteria. Yet, as Birch himself notes in Proposal 17, sentience is not pain. We argue that statutory welfare protections must therefore reach beyond pain, and that the right tool for doing so is Birch et al.’s (2020) dimensionality framework, which characterises conscious experience across five domains. Used together, the LSE Criteria establish that welfare is at stake, while the dimensionality framework maps what is at stake beyond pain to provide a taxon-specific profile of where captivity impoverishes experience. We apply this approach to invertebrate welfare and …


On The Edge Of Science?, Mark Briffa, Lynne U. Sneddon Jun 2026

On The Edge Of Science?, Mark Briffa, Lynne U. Sneddon

Animal Sentience

In The Edge of Sentience, Jonathan Birch (2024) provides a thought-provoking synthesis of current perspectives on animal (and artificial) sentience. He addresses the question from conceptual, ethical, regulatory and empirical viewpoints. Here we focus on some of the scientific issues raised, including defining and operationalising the concept of sentience and the implications for policy and regulation. Despite unresolved conceptual challenges, the book makes a compelling case for the continued scientific investigation of animal sentience.


Natural Behavior Vs. Subjective Experience In Fish Welfare Assessments, Ronald G. Oldfield Jun 2026

Natural Behavior Vs. Subjective Experience In Fish Welfare Assessments, Ronald G. Oldfield

Animal Sentience

Several commentaries on the Oldfield & Bonano (2024) target article proposed that natural behavior should play a larger role in assessing the welfare of fishes in zoos and aquariums. However, the natural-behavior concept of animal welfare has been widely criticized because natural behaviors do not necessarily improve welfare, and unnatural behaviors do not necessarily reduce it. I review the three major concepts of animal welfare — physical health, natural behavior, and affect/feelings — and argue that behavior patterns, whether natural or otherwise, should be evaluated according to their effects on health and subjective experience. Using the affect/feelings concept and the …


Aggression And Fish Welfare: Challenges And Opportunities, Christine M. Soper, Peter D. Dijkstra Apr 2026

Aggression And Fish Welfare: Challenges And Opportunities, Christine M. Soper, Peter D. Dijkstra

Animal Sentience

Oldfield & Bonano’s (2024) [O&B] target article discusses the wellbeing and sociality of bony fishes. Many fish species, such as cichlids, are not only social but territorial. Aggression is normal, but territorial aggression in a captive environment can have a negative impact on welfare. Aggression-related injuries and deaths are prevalent in pet stores as well as research settings. We discuss the need to develop effective methods to minimize the negative consequences of aggression.


Both Studying And Protecting Octopuses “On Their Own Terms”, Jennifer Jacquet, Janelle Kaz, Becca Franks, Christine Webb Nov 2025

Both Studying And Protecting Octopuses “On Their Own Terms”, Jennifer Jacquet, Janelle Kaz, Becca Franks, Christine Webb

Animal Sentience

Cephalopods – indeed, all animals – “must be seen on their own terms,” with respect for their autonomy and their ecological imperatives. This will provide greater scientific insight along with a more ethical and accurate understanding of minds beyond our own.


Cephalopod Consciousness On “Their Own Terms”: Who Are “They”?, Willa M. Lane, Victor Ajuwon, Nicola S. Clayton Nov 2025

Cephalopod Consciousness On “Their Own Terms”: Who Are “They”?, Willa M. Lane, Victor Ajuwon, Nicola S. Clayton

Animal Sentience

Mather’s call to consider cephalopods’ consciousness “on their own terms” raises the question of the extent to which cephalopods can be considered a homogenous group. While coloeoid cephalopods share traits that strongly distinguish them from other molluscs and invertebrates, they show vast ecological and morphological diversity, with implications for the design of empirical investigations of consciousness. Extending Mather’s argument, we apply Birch et al.’s (2020) five-dimensional model to compare octopus, cuttlefish, squid, and nautilus. This comparison shows that even within a graded rather than categorical framework, putative consciousness profiles depend on which taxa are included in discussions of “cephalopod consciousness.” …


The Term Consciousness And Impressive Evidence For Cephalopod Awareness, Donald M. Broom Oct 2025

The Term Consciousness And Impressive Evidence For Cephalopod Awareness, Donald M. Broom

Animal Sentience

Mather presents comprehensive evidence of cephalopod cognitive ability and of a high level of awareness. The importance of taking account of the biological functioning and adaptations to the living environment is emphasised. An argument is presented that it is better to limit the use of the word conscious to ‘not unconscious’ and to use awareness in relation to the various levels of cognitive functioning.


More Equal Than Others: Taxonomic Bias In Animal Management, Barrett L. Christie Jun 2025

More Equal Than Others: Taxonomic Bias In Animal Management, Barrett L. Christie

Animal Sentience

Oldfield & Bonano (2024) point out not only the scarcity of research on fish welfare but also significant taxonomic bias in broader research on these species. This bias extends to the management practices of aquaria and zoos, where the focus is on higher vertebrates. More resources need to be allocated to research to address this disparity. We also need to develop welfare constructs that recognize the inherent differences between aquatic and terrestrial species, taking into account the unique ecological contexts and behavioral patterns of fishes.


Précis Of The Edge Of Sentience: Risk And Precaution In Humans, Other Animals, And Ai, Jonathan Birch Jan 2025

Précis Of The Edge Of Sentience: Risk And Precaution In Humans, Other Animals, And Ai, Jonathan Birch

Animal Sentience

We often face grave practical decisions that seem to hinge on whether a system is sentient. This family of cases includes invertebrate animals, people who are unresponsive after brain injury, fetuses, neural organoids, and now AI technologies. We must decide what to do despite ongoing disagreement about the nature of sentience. In our state of uncertainty, we should pragmatically transform the question from “Is it sentient?” to “Is it a sentience candidate, an investigation priority, or neither?”. When a system is a sentience candidate, it is negligent to fail to consider precautions. We should instead evaluate precautions for their proportionality …


Social Learning And Cultural Enrichment For Fish Welfare, Chiawen Chiang, Becca Franks Dec 2024

Social Learning And Cultural Enrichment For Fish Welfare, Chiawen Chiang, Becca Franks

Animal Sentience

This commentary examines how understanding animal culture—shared behaviors and knowledge transmitted through social learning—plays a role in advancing welfare practices for captive fishes. Despite extensive evidence of fishes' complex sociocultural capacities, their psycho-social needs remain underestimated and unmet in zoos and aquaria. We explore what considering fish culture might mean for welfare practices: diverse social learning opportunities, cultural development, and thoughtful group management. More knowledge about wild fishes' lives will help bridge the gap between their full, complex lives and current captive welfare practices.


Understanding The Welfare Of Aquarium Fish, Paul Rose Nov 2024

Understanding The Welfare Of Aquarium Fish, Paul Rose

Animal Sentience

Bony fishes (Osteichthyes) are prevalent in public and domestic aquaria, yet evidence-based welfare practices exist for only a limited number of species. Oldfield & Bonano’s (O&B’s) target article highlights critical questions about the wellbeing and sociality of Osteichthyes. Advancing research on their biology, physiology, psychology, and behavior is essential to develop and validate welfare measures, ensuring guidelines for welfare-focused housing and husbandry.


Can We Use Judgement Bias To Assess The Welfare Of Fish In Aquariums And Zoos?, Culum Brown Oct 2024

Can We Use Judgement Bias To Assess The Welfare Of Fish In Aquariums And Zoos?, Culum Brown

Animal Sentience

This commentary examines the potential of tests of judgment bias to assess fish welfare in aquariums and zoos. Judgment bias offers a non-invasive approach to interpreting affective states in animals, using cognitive responses to ambiguous stimuli as indicators of mental wellbeing. Despite the constraints of captive environments—such as limited handling and species diversity in exhibits—recent adaptations of judgment-bias protocols make them suitable across taxa, including fishes. Collaborations with progressive zoos and aquariums could facilitate this research, based on the shared goal of enhancing animal welfare and promoting public education.


Psychological And Social Well-Being Of Bony Fishes In Zoos And Aquariums, Ronald G. Oldfield, Patrick Bonano Oct 2024

Psychological And Social Well-Being Of Bony Fishes In Zoos And Aquariums, Ronald G. Oldfield, Patrick Bonano

Animal Sentience

Research on the behaviour of animals in zoos has been conducted for decades, and observations have provided information that has improved the psychological and social well-being of animals. However, research on fishes in zoos and aquariums seems to be lacking. Here, we assess the current state of research on fishes in zoos and aquariums by surveying the peer-reviewed literature. Our assessment differs from previous surveys in that we examine the taxonomic classes Chondrichthyes (sharks and rays) and Osteichthyes (bony fishes) separately. Our survey finds that bony fishes have been drastically underrepresented in zoo journals, more drastically than chondrichthyans, revealing an …


Complexities Of Consistent Individual Behavioral Differences In Effective Wildlife Management, Nathalie R. Sommer Jan 2024

Complexities Of Consistent Individual Behavioral Differences In Effective Wildlife Management, Nathalie R. Sommer

Animal Sentience

The study of animal personality, or consistent individual behavioral differences, has faced numerous challenges since its inception, including terminological disputes, labor-intensive methodologies, and notable retractions. This commentary explores the mechanisms behind personality traits, questioning the link between personality and sentience, and highlights the bias towards megafauna in wildlife management. It underscores the ecological and ethical significance of considering arthropod personalities and the complexities of integrating personality into wildlife reintroductions. Caution is advised in generalizing personality effects, and the limitations of current measurement techniques are discussed, suggesting that personality traits should be one of many factors in comprehensive management strategies.


Not Just Bony Fish: All Nonmammalian And Nonavian Animals Are Neglected In Both Welfare And General Research, Jennifer A. Mather Ph.D. Jan 2024

Not Just Bony Fish: All Nonmammalian And Nonavian Animals Are Neglected In Both Welfare And General Research, Jennifer A. Mather Ph.D.

Animal Sentience

The lack of research on fish welfare highlighted by Oldfield & Bonano (2024) is only part of a more general problem: a relative neglect of animals other than mammals and birds. The causes are our lack of knowledge, our negative view of what we consider to be “lower” animals and our underestimation of their intelligence and sensitivity. Our estimation will increase as our knowledge increases. As a broader view spreads to the public through the media, it should gradually increase the demand to expand both research and welfare, especially beyond mammals and birds.


Plants Lack The Functional Neurotransmitters And Signaling Pathways Required For Sentience In Animals, David G. Robinson, Michael R. Blatt, Andreas Draguhn, Lincoln Taiz, Jon Mallatt Apr 2023

Plants Lack The Functional Neurotransmitters And Signaling Pathways Required For Sentience In Animals, David G. Robinson, Michael R. Blatt, Andreas Draguhn, Lincoln Taiz, Jon Mallatt

Animal Sentience

We cannot agree with Segundo-Ortin and Calvo that plants are sentient organisms. We have critically examined several aspects of their target article, and find their claims are not supported by the published evidence. We address these claims in sections on whether plants have a ‘neurobiology’ analogous to that of animal nervous systems, including neurotransmitters and synaptic receptors that respond to anesthetics; and whether plant signaling resembles neural transmission. For the latter, we especially consider the unique way plants signal their responses to wounding. Although the plant vascular system has been compared to the animal nervous system, animal blood vessels would …


Plant Sentience? Between Romanticism And Denial: Science, Miguel Segundo-Ortin, Paco Calvo Jan 2023

Plant Sentience? Between Romanticism And Denial: Science, Miguel Segundo-Ortin, Paco Calvo

Animal Sentience

A growing number of non-human animal species are being seriously considered as candidates for sentience, but plants are either forgotten or explicitly excluded from these debates. In our view, this is based on the belief that plant behavior is hardwired and inflexible and on an underestimation of the role of plant electrophysiology. We weigh such assumptions against the evidence to suggest that it is time to take seriously the hypothesis that plants, too, might be sentient. We hope this target article will serve as an invitation to investigate sentience in plants with the same rigor as in non-human animals.


Pain Sentience Criteria And Their Grading, Eva Jablonka, Simona Ginsburg Jan 2022

Pain Sentience Criteria And Their Grading, Eva Jablonka, Simona Ginsburg

Animal Sentience

On the basis of the target article by Crump and colleagues, we suggest a more parsimonious scheme for evaluating the evidence for sentience. Since some of the criteria used by Crump et al. are not independent and some are uninformative we exclude some criteria and amalgamate others. We propose that evidence of flexible learning and prioritization, in conjunction with relevant data on brain organization, is sufficient for assigning pain-sentience to an animal and we suggest a scoring scheme based on four criteria.


Truly Minimal Criteria For Animal Sentience, Mark Solms Jan 2022

Truly Minimal Criteria For Animal Sentience, Mark Solms

Animal Sentience

The criteria for determining animal sentience proposed in the target article are sensible but they lack an explicit functional justification for the focus on pain. This commentary provides an abbreviated account of the most basic functional principles that underpin animal sentience and articulates some minimal criteria for determining its presence.


Crustacean Pain, Michael Tye Jan 2022

Crustacean Pain, Michael Tye

Animal Sentience

This commentary discusses the target article’s methodology, the relevance of the claim that crustaceans lack a neocortex to the thesis that they feel pain, and the evaluation of the results of some trade-off experiments done with hermit crabs.


No Need For Certainty In Animal Sentience, Yew Kwang Ng Jan 2022

No Need For Certainty In Animal Sentience, Yew Kwang Ng

Animal Sentience

This commentary supports Crump et al.’s (2022) point that where risks to welfare are severe, strong evidence of sentience is sufficient to warrant protecting welfare. Crump et al.’s eight criteria for sentience are also useful. Flexible decision-making (5) and flexible behaviour (6) are consistent with Ng (1995). The concession that the “no-need-for-sentience” proposition is unnecessary also strengthens the importance of the target article’s conclusions.


What Might Decapod Sentience Mean For Policy, Practice, And Public?, Richard Gorman Jan 2022

What Might Decapod Sentience Mean For Policy, Practice, And Public?, Richard Gorman

Animal Sentience

Crump et al. provide eight criteria for evaluating sentience in decapods, with scope for for application to other taxa. Their work has attracted the interest of policymakers. This commentary discusses the limitations of conceptual and legal acknowledgement of sentience in chainging practice and public attitudes. More work is needed. Social science may be able to help.


Does The Sentience Framework Imply All Animals Are Sentient?, Kristin Andrews Jan 2022

Does The Sentience Framework Imply All Animals Are Sentient?, Kristin Andrews

Animal Sentience

The eight criteria proposed in Crump et al.’s framework for evaluating pain sentience in decapod crustaceans are just the tip of the iceberg when it comes to markers that could increase confidence in an animal’s sentience more generally. Some of the commentaries have already pointed out that pain is only one kind of sentience (Souza Valente). It has also already been pointed out that there are other criteria for pain that could be usefully added to the framework’s eight (Burrell). This expansive thinking about criteria that can be used to increase confidence in sentience raisess the question: in an expansive …


Distinguishing Epistemic And Moral Grounds For Legal Protection, Carlos Montemayor Jan 2022

Distinguishing Epistemic And Moral Grounds For Legal Protection, Carlos Montemayor

Animal Sentience

The criteria proposed by Crump et al. are based on various cognitive roles associated with sentience. A subset of them may be sufficient for certain kinds of welfare, but the presence of all of them should be considered as clearly sufficient for substantial kinds of legal protection based on their relation to capacities that we consider essential for moral standing in human beings.


Decapod Sentience: Broadening The Framework, Cecilia De Souza Valente Jan 2022

Decapod Sentience: Broadening The Framework, Cecilia De Souza Valente

Animal Sentience

A framework for studying sentience in decapods is of great value, but how high a cost (in suffering) to each individual decapod (or any animal) is warranted for collecting scientific evidence of sentience? The lack of evidence for some of the target article’s proposed criteria surely results from the fact that research is focused mainly on biomedical studies, ecotoxicology, and commercial production, with decapod sentience and welfare seen as only a secondary research topic. I draw attention also to the possibility of a wider framework that includes all felt experiences, from suffering to pleasure.


Sentience In Decapods: Difficulties To Surmount, Michael L. Woodruff Jan 2022

Sentience In Decapods: Difficulties To Surmount, Michael L. Woodruff

Animal Sentience

In the target article Crump et al. present 8 criteria to assess whether decapods experience pain. Four of these -- sensory integration, motivational trade-offs, flexible self-protection, and associative learning -- could be used to assess sentience in general. In this commentary I discuss difficulties with using these criteria to provide evidence of sentience in decapods, particularly if this evidence is to change public opinion and policies. These difficulties are lack of evidence, the potential to eventually explain the neurobiological basis of the behaviors chosen as criteria, thereby eliminating any explanatory work for sentience, and the reluctance to bring animals that …


Pain In Pleocyemata, But Not In Dendrobranchiata?, Gary Comstock Jan 2022

Pain In Pleocyemata, But Not In Dendrobranchiata?, Gary Comstock

Animal Sentience

Crump et al.’s contribution to assessing whether decapods feel pain raises an important question: Is pain distributed unevenly across the order? The case for pain appears stronger in Pleocyemata than in Dendrobranchiata. Some studies report pain avoidance behaviors in Dendrobranchiata (Penaeidae) shrimp, but further studies are needed to determine whether the chemicals used are acting as analgesics to relieve pain, or as soporifics to reduce overall alertness. If the latter, the most farmed shrimp species may not require the same level of protection as crabs, crayfish, and lobsters.


Decapod Sentience: Promising Framework And Evidence, Jon Mallatt, Todd E. Feinberg Md Jan 2022

Decapod Sentience: Promising Framework And Evidence, Jon Mallatt, Todd E. Feinberg Md

Animal Sentience

Strong points of the target article by Crump et al. are that it offers clear criteria for judging whether decapods are sentient, an effective semi-quantitative grading system for this purpose, and an astute, critical review of the literature. It concludes plausibly that major subgroups of decapods are sentient. A minor problem is that it includes classical, Pavlovian learning as a marker of sentience along with the more valid marker of complex (e.g., operant) learning. Another minor problem is that it does not distinguish results that are negative because of likely absence of sentience from results that are negative because they …


Decapods As Food, Companions And Research Animals: Legal Impact Of Ascribing Sentience, Jonathan J. Cooper, Ambrose Tinarwo, Beth A. Ventura Jan 2022

Decapods As Food, Companions And Research Animals: Legal Impact Of Ascribing Sentience, Jonathan J. Cooper, Ambrose Tinarwo, Beth A. Ventura

Animal Sentience

This commentary provides an overview of the practical implications of attributing sentience to protect decapods as food, companion and research animals in the UK context. Recognising their capacity to suffer has implications for humane slaughter in farming and fishing sectors. It should also place a greater duty of care on owners of captive decapods, considering their needs and avoiding unnecessary suffering. The recognition of decapod sentience should also have an impact on their protection as research animals, although research with a potential to cause suffering may be needed to better understand decapods’ capacity to suffer.