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Full-Text Articles in Population Biology

2023 (35th) Report Of The Nou Records Committee, Mark A. Brogie Sep 2024

2023 (35th) Report Of The Nou Records Committee, Mark A. Brogie

Nebraska Bird Review

The functions and methods of the Nebraska Ornithologists’ Union Records Committee (NOURC) are described in its bylaws (NOURC 2010). The committee’s purpose is to provide a procedure for documenting unusual bird sightings and to establish a list of all documented birds for Nebraska. Species for which the NOURC seeks documentation (NOURC Review List) can be found at the NOU website https://noubirds.org/Birds/ReviewList.aspx. Starting in 2020, the NOURC decided to deal mainly with those species deemed as Accidental on the “Official List of the Birds of Nebraska”. Casual species and out-of- range species are now summarized in the “Seasonal Reports” of the …


The Nebraska Bird Review (September 2024) 92(3), Whole Issue Sep 2024

The Nebraska Bird Review (September 2024) 92(3), Whole Issue

Nebraska Bird Review

Summer Field Report, June–July 2024, by W. Ross Silcock

Correcting the Record: Subspecific Identification of Nebraska’s Lone Common Eider (Somateria mollissima), by Sam Manning, Joel G. Jorgensen, Stephen J. Brenner, and W. Ross Silcock

2023 (35th) Report of the NOU Records Committee, by Mark A. Brogie

Subscription and Organization Information


Summer Field Report, June–July 2024, W. Ross Silcock Sep 2024

Summer Field Report, June–July 2024, W. Ross Silcock

Nebraska Bird Review

This was an intriguing summer in several ways. I am far from being a knowledgeable ecologist, but, like most of us, as a layman I wonder whether and how weather affects birds, and if and how local weather is reflecting the conventional wisdom of climate change. Climate change seems increasingly to be expressed as various extreme or rapidly changing weather events that have differing effects on vegetation and consequently insect and larger prey populations. Every species responds differently it seems, depending on its biological needs, but food supply is the most important issue for birds. It is tempting to assign …


From Turf To Turf: Fungal Invasion Of Ascocoryne Turficola In A Northern Michigan Bog, Evan L. Coscarelli-Trott Sep 2024

From Turf To Turf: Fungal Invasion Of Ascocoryne Turficola In A Northern Michigan Bog, Evan L. Coscarelli-Trott

All NMU Master's Theses

This is the first paper to report Ascocoryne turficola Boud. 1905 (Helotiales: Ascomycota) within the United States. It is an obligate bog fungus native to Europe and Siberia, where it is extremely rare across most of its range. A. turficola appeared at Lily Pond Bog in Marquette, Michigan (87.47º W, 46.59º N) in 2019, almost contemporaneous with anecdotal reports in Minnesota and Ontario. Since 2019, the Lily Pond population has exploded and become the dominant fungus in this bog but has yet to spread to similar bogs nearby. This species is adapted to long distance dispersal and carries the potential …


The Influence Of Environmental Change On Genetic Diversity Across Spatial And Taxonomic Scales, Connor M. French Sep 2024

The Influence Of Environmental Change On Genetic Diversity Across Spatial And Taxonomic Scales, Connor M. French

Dissertations, Theses, and Capstone Projects

The spatial distribution of genetic diversity is of interest to biodiversity scientists and conservationists and is a fundamental metric of biodiversity. Genetic diversity patterns across spatial and taxonomic scales contain information about population and assemblage dynamics that can convey their resilience to environmental change. Ectotherms are especially linked to their environments and may be especially sensitive to fluctuations in the environment over time. Herein, I investigate global and regional patterns of genetic diversity in two groups of ectotherms, insects and lizards, to understand the relationship between environmental change and genetic diversity, from populations to assemblages. Overall, my research aims to …


Impact Of Thermal-Hydric Stress Of Physiological Performance And Mating Behavior Of Fiddler Crabs, Talene G. Yeghissian Aug 2024

Impact Of Thermal-Hydric Stress Of Physiological Performance And Mating Behavior Of Fiddler Crabs, Talene G. Yeghissian

Dissertations

Fiddler crabs live on the edge of their thermal tolerance limits, balancing the need to thermoregulate with the need to mate. Previous research suggests that increased temperature reduces fiddler crab surface activity time, thus decreasing mating opportunities, throughout the range species with greatest impacts to low latitude populations. The less studied missed mating opportunities associated with behavioral compensation for thermal stress are crucial to understanding species vulnerability to climate change. In this dissertation, I describe a combination of experimental and observational approaches in the field and in the laboratory to investigate the physiological and behavioral responses of fiddler crabs to …


Effects Of A Protection Zone In A Reaction-Diffusion Model With Shifting Bounded Habitat And Allee Effect., Davis Henderson Aug 2024

Effects Of A Protection Zone In A Reaction-Diffusion Model With Shifting Bounded Habitat And Allee Effect., Davis Henderson

Electronic Theses and Dissertations

We consider a reaction-diffusion equation that models a population in a shifting bounded habitat with a protection zone and the Allee effect. We assume that the population growth function exhibits the strong Allee effect and has a positive integral within the protection zone (indicating population persistence) and a negative integral in the surrounding patches (indicating population decay). We prove that the existence of steady-state solutions to this system depends on the length of the protection zone. It is demonstrated that there exists some value H* such that, for a protection zone of size H*, there exists a solution and, for …


The Role Of Sexual Network Core Groups In Disease Epidemics, Jason Janeaux Jul 2024

The Role Of Sexual Network Core Groups In Disease Epidemics, Jason Janeaux

LSU Doctoral Dissertations

Sexually transmitted infections (STIs) remain an economic and social burden, non-discriminately affecting sexually active individuals worldwide. The unique transmission of STIs is dependent upon host and pathogen life histories, as well as the underlying network structure of the host population. Within a population there is a group of individuals, the core group, that maintains STI presence and helps sustain ongoing transmission both within the core group and in the surrounding population. While the presence of these core groups is well documented, the criteria for inclusion into a sexual core group is not. In this dissertation, I explore common factors that …


Spatial Ecology Of Mountain Lions Throughout Their Distribution In California, Kyle Duncan Dougherty Jul 2024

Spatial Ecology Of Mountain Lions Throughout Their Distribution In California, Kyle Duncan Dougherty

Dissertations and Doctoral Documents, University of Nebraska-Lincoln, 2023–

Mountain lions (Puma concolor) have been studied extensively throughout their wide geographic distribution in the Western Hemisphere, but most research has been conducted at relatively small spatial scales. We used GPS telemetry data contributed by researchers in 16 study areas throughout California to advance understanding of mountain lion spatial ecology. First, we investigated multi-scale resource selection of mountain lions relative to spatially varying risk of intentional human-caused mortality associated with livestock depredation management. We show that individuals that proactively avoided mortality risk when selecting home ranges were able to relax their avoidance of risk when making resource selection …


Hybridization Between The Rare Gray-Headed Chickadee And The Abundant Boreal Chickadee In The Midst Of Shifting Climate, Matthew R. Armstrong Jul 2024

Hybridization Between The Rare Gray-Headed Chickadee And The Abundant Boreal Chickadee In The Midst Of Shifting Climate, Matthew R. Armstrong

School of Natural Resources: Dissertations, Theses, and Student Research

As species respond to changing climate, distributions and abundances may shift and alter species interactions. Hybridization, a relatively widespread phenomenon becoming more common with climate change, can have beneficial and detrimental effects on population growth rates and genetic integrity. Beneficial effects due to the introduction of advantageous alleles and increased genetic diversity may result from hybridization. Species may also accrue fitness costs associated with changing climates if mismatches occur between environmental variables and phenotypes. The gray-headed chickadee, Poecile cinctus lathami, is an extremely rare songbird that has experienced marked declines in recent decades within its restricted distribution in Alaska …


New Insights On Hybridization In Potamogeton Floridanus (The Florida Pondweed), Kaitlyn R. Sampson Jul 2024

New Insights On Hybridization In Potamogeton Floridanus (The Florida Pondweed), Kaitlyn R. Sampson

Graduate Theses and Dissertations (2019 - present)

Freshwater ecosystems are some of the most important and highest threatened habitats in the world, and aquatic plants play an important, but often-overlooked, role in maintaining them. Potamogeton is a diverse and ecologically important aquatic plant genus well known for taxonomic difficulty and rampant hybridization. lbis study aimed to 1) test the hypothesis that Potamogeton jloridanus (Florida pondweed) is a hybrid between P. oakesianus and P. pulcher, and 2) to investigate correlations in ecological conditions for the focal species. This study revealed the discovery of a new population of P. jloridanus in Big Coldwater Creek in Santa Rosa Co., FL, …


Spatial And Abiotic Effects Of Urbanization On Small Mammal Communities, Bri Casement Jul 2024

Spatial And Abiotic Effects Of Urbanization On Small Mammal Communities, Bri Casement

Master's Theses

Urbanization, the phenomenon of increasing human population density and artificial structures, is known to decrease the richness, abundance, and diversity of native wildlife populations, fragment natural areas, and increase human and human-commensal animal interactions with wildlife. Dynamics of species persistence in habitat fragments is often understood in the framework of island biogeography theory (IBT), with habitat fragments being analogous to oceanic islands. Despite the conceptual similarities between oceanic islands in IBT and urbanized habitat fragments (fragment size, fragment age, and distance to immigration sources), there are important differences. In urban fragments, the matrix is not completely impassible, habitats within fragments …


Removal As A Control Method For Trachemys Scripta Elegans And The Response Of Chrysemys Picta Bellii, Jacob Swanson Jun 2024

Removal As A Control Method For Trachemys Scripta Elegans And The Response Of Chrysemys Picta Bellii, Jacob Swanson

University Honors Theses

The red-eared slider is considered one of the most ubiquitous freshwater turtles globally. Several key ecological advantages and life history traits of this species jointly impact growth rates and survivorship of native freshwater turtles where sympatry occurs. Evaluation of current management actions for red-eared slider turtles is limited, resulting in a paucity of information to guide management for the species and conservation of declining native freshwater turtles. We used previously collected mark-recapture and removal sampling data from sites with sympatric populations of red-eared slider and western painted turtles. We examined the effects of control (trapping and removal) of red-eared sliders …


The Effects Of Wildfire Burn Severity On Bee And Wasp Communities Two Decades Post-Fire, Alaina K. Smith Jun 2024

The Effects Of Wildfire Burn Severity On Bee And Wasp Communities Two Decades Post-Fire, Alaina K. Smith

Electronic Theses and Dissertations

While fire is a natural disturbance in many global ecosystems, anthropogenic changes are affecting fire frequency, severity, and seasonality. Disturbances like wildfire affect bee and wasp communities, but recent meta-analyses are conflicted about whether fire harms or benefits insect pollinators. We investigated the lasting effects of wildfire burn severity on wild bee communities and floral resource availability across different burn severities in two-mixed severity fires that burned >20 years ago. We found that high severity fire has positive effects on bee diversity that are associated with severity-mediated changes to the floral community. We also studied the effects of wildfire burn …


A Multispecies Hierarchical Model To Integrate Count And Distance-Sampling Data, Neil A. Gilbert, Caroline M. Blommel, Matthew T. Farr, David S. Green, Kay E. Holekamp, Elise F. Zipkin Jun 2024

A Multispecies Hierarchical Model To Integrate Count And Distance-Sampling Data, Neil A. Gilbert, Caroline M. Blommel, Matthew T. Farr, David S. Green, Kay E. Holekamp, Elise F. Zipkin

Institute for Natural Resources Publications

Integrated community models—an emerging framework in which multiple data sources for multiple species are analyzed simultaneously—offer opportunities to expand inferences beyond the single-species and single-data-source approaches common in ecology. We developed a novel integrated community model that combines distance sampling and single-visit count data; within the model, information is shared among data sources (via a joint likelihood) and species (via a random-effects structure) to estimate abundance patterns across a community. Parameters relating to abundance are shared between data sources, and the model can specify either shared or separate observation processes for each data source. Simulations demonstrated that the model provided …


Nou Spring Field Days And Annual Meeting, 125th Anniversary Of The Nebraska Ornithologists’ Union, Birding Bowl Kickoff Weekend, Lincoln, Mary 3–5, 2024, Janis Paseka Jun 2024

Nou Spring Field Days And Annual Meeting, 125th Anniversary Of The Nebraska Ornithologists’ Union, Birding Bowl Kickoff Weekend, Lincoln, Mary 3–5, 2024, Janis Paseka

Nebraska Bird Review

The NOU Annual Meeting and Spring Field Days were held in Lincoln on May 3-5, 2024, with 80 in attendance. This year is the 125th anniversary of the founding of the Nebraska Ornithologists’ Union, and this meeting also served as the kickoff of the Nebraska Birding Bowl.

Field trips were led by Olivia DaRugna, Thane Dinsdale, Joe Gubanyi, Thomas Labedz, Jason St. Sauver, T. J. Walker, and Michael Willison. Destinations included Wilderness Park, Bluestem Lake SRA, Killdeer WMA, Cottontail WMA, Pawnee Lake SRA, Branched Oak Lake, Butler County Prairie-Chicken lek, Marsh Wren Saline Wetland, Spring Creek Prairie, and Conestoga Lake …


The Early Years Of The Nebraska Ornithologists’ Union: People And Publications, Daniel W. Leger Jun 2024

The Early Years Of The Nebraska Ornithologists’ Union: People And Publications, Daniel W. Leger

Nebraska Bird Review

Nebraska once boasted that it had the greatest avian biodiversity of any state. Today, such a claim seems preposterous, but when it was made, back in 1899, it was serious. The antecedents of that claim underlie the objective of this article, namely to describe the early history of the Nebraska Ornithologists’ Union (NOU), its founding members, and its publications, one of which, the Nebraska Bird Review, you are reading today.

. . .

To say that many changes have occurred in ornithology and birding since the NOU started in 1899 would be a severe understatement. Most of these changes have …


The Nebraska Bird Review (June 2024) 92(2), Whole Issue Jun 2024

The Nebraska Bird Review (June 2024) 92(2), Whole Issue

Nebraska Bird Review

Spring Field Report, March–May 2024, by W. Ross Silcock

The Early Years of the Nebraska Ornithologists’ Union: People and Publications, by Daniel W. Leger

NOU Spring Field Days and Annual Meeting, 125th Anniversary of the Nebraska Ornithologists’ Union, Birding Bowl Kickoff Weekend, Lincoln, May 3–5, 2024, by Janis Paseka

Subscription and Organization Information


Spring Field Report, March–May 2024, W. Ross Silcock Jun 2024

Spring Field Report, March–May 2024, W. Ross Silcock

Nebraska Bird Review

A major concern this spring was the widely lamented shortfall in numbers of migrant warblers and the shortened duration of the migration, with most sightings relatively early. Numbers of migrant warblers east of the 100th meridian were investigated by Joel Jorgensen and Stephen Brenner and the scarcity was indeed found to be supported by eBird data; see the “News and Updates” section in Birds of Nebraska-Online (BONO). At Oliver Reservoir in the Panhandle, however, Steven Mlodinow found that migration was at least normal for a range of common migrants, with some migrants above the levels of the last four years. …


Multi-Type Branching Processes In Time-Varying Environments, Arash Jamshidpey May 2024

Multi-Type Branching Processes In Time-Varying Environments, Arash Jamshidpey

Biology and Medicine Through Mathematics Conference

No abstract provided.


Mathematically Modeling Stoichiometric Drivers Of Nitrogen Fixation, Rebecca Everett, Corday Selden, Mohamed Hatha Abdulla, Jabir Thajudeen, James Powell, Edwin Cruz-Rivera, Luca Schenone, Renn Schipper, Megan Berberich, Halvor Halvorson, Robinson Fulweiler, Amy Marcarelli, Thad Scott May 2024

Mathematically Modeling Stoichiometric Drivers Of Nitrogen Fixation, Rebecca Everett, Corday Selden, Mohamed Hatha Abdulla, Jabir Thajudeen, James Powell, Edwin Cruz-Rivera, Luca Schenone, Renn Schipper, Megan Berberich, Halvor Halvorson, Robinson Fulweiler, Amy Marcarelli, Thad Scott

Biology and Medicine Through Mathematics Conference

No abstract provided.


Maturation Of An Off-Channel Habitat Concept To Conserve Native Fishes In The Lower Colorado River, Paul C. Marsh, Thomas E. Dowling, Thomas F. Turner, Megan J. Osborne, Brian R. Kesner May 2024

Maturation Of An Off-Channel Habitat Concept To Conserve Native Fishes In The Lower Colorado River, Paul C. Marsh, Thomas E. Dowling, Thomas F. Turner, Megan J. Osborne, Brian R. Kesner

Monographs of the Western North American Naturalist

Two endemic, “large river” fishes of the Colorado River basin of western North America, bonytail Gila elegans and razorback sucker Xyrauchen texanus, are among several critically endangered species in the system. Wild populations of bonytail are gone, and there are no self-sustaining populations of razorback sucker anywhere; reproduction occurs but recruitment does not. Both species have been under intensive management in the Lower Colorado River since the 1980s. Today, with the single exception of Lake Mead, remaining populations are composed entirely of repatriated individuals that depend on stocking for their continued existence. In 2003, a conceptual off-channel habitat (OCH) …


Omani Camels From A Cultural And Genomics Perspective, Al Muatasim Al Zadjali May 2024

Omani Camels From A Cultural And Genomics Perspective, Al Muatasim Al Zadjali

Electronic Theses and Dissertations

The Dromedarian camel, Camelus dromedarius, is native to the Arabian Peninsula, including the Sultanate of Oman. These camels are used for food, milk, as well as show and racing competitions. Despite their economic and cultural importance research on camels in Oman is limited. The goal of this study was to examine their genomic variation, relationship with camels in other parts of the Arabian Peninsula, and to determine if selective breeding has led to the establishment of distinct breeds in Oman. Information was compiled from multiple sources to produce a comprehensive review on the breeding, management, economic and cultural use, …


Soil Bacterial Community Shifts In A Montane Longleaf Pine Restoration Site: Effects Of Dominant Herbaceous Species Removal, Prescribed Burn, And Changes In Season, Isabella K. Vahle May 2024

Soil Bacterial Community Shifts In A Montane Longleaf Pine Restoration Site: Effects Of Dominant Herbaceous Species Removal, Prescribed Burn, And Changes In Season, Isabella K. Vahle

Master's Theses

Longleaf pine (Pinus palustris Mill.) ecosystems are one of the most endangered and ecologically diverse systems in North America. The native herbaceous species serve as fuel for low-intensity, recurring fires this system relies on. The herbaceous species play major roles in maintaining ecosystem structure and diversity. Soil microorganisms like bacteria are known to aid greatly in many different plant and ecosystem processes. However, little is known about the dominant herbaceous species and soil bacterial compositions within montane longleaf pine ecosystems, especially before and after a prescribed fire. We completed a multiyear, seasonal analysis of the soil bacterial community in …


Researching Covid To Enhance Recovery (Recover) Pediatric Study Protocol: Rationale, Objectives And Design, Rachel S. Gross, Tanayott Thaweethai, Erika B. Rosenzweig, James Chan, Lori B. Chibnik, Mine S. Cicek, Amy J. Elliott, Valerie J. Flaherman, Andrea S. Foulkes, Margot Gage Witvliet, Richard Gallagher, Maria Laura Gennaro, Terry L. Jernigan, Elizabeth W. Karlson, Stuart D. Katz, Patricia A. Kinser, Lawrence C. Kleinman, Michelle F. Lamendola-Essel, Joshua D. Milner, Sindhu Mohandas, Praveen C. Mudumbi, Jane W. Newburger, Kyung E. Rhee, Amy L. Salisbury, Jessica N. Snowden, Cheryl R. Stein, Melissa S. Stockwell, Kelan G. Tantisira, Tamara Bradford, Et Al May 2024

Researching Covid To Enhance Recovery (Recover) Pediatric Study Protocol: Rationale, Objectives And Design, Rachel S. Gross, Tanayott Thaweethai, Erika B. Rosenzweig, James Chan, Lori B. Chibnik, Mine S. Cicek, Amy J. Elliott, Valerie J. Flaherman, Andrea S. Foulkes, Margot Gage Witvliet, Richard Gallagher, Maria Laura Gennaro, Terry L. Jernigan, Elizabeth W. Karlson, Stuart D. Katz, Patricia A. Kinser, Lawrence C. Kleinman, Michelle F. Lamendola-Essel, Joshua D. Milner, Sindhu Mohandas, Praveen C. Mudumbi, Jane W. Newburger, Kyung E. Rhee, Amy L. Salisbury, Jessica N. Snowden, Cheryl R. Stein, Melissa S. Stockwell, Kelan G. Tantisira, Tamara Bradford, Et Al

School of Medicine Faculty Publications

Importance The prevalence, pathophysiology, and long-term outcomes of COVID-19 (post-acute sequelae of SARS-CoV-2 [PASC] or “Long COVID”) in children and young adults remain unknown. Studies must address the urgent need to define PASC, its mechanisms, and potential treatment targets in children and young adults. Observations We describe the protocol for the Pediatric Observational Cohort Study of the NIH’s REsearching COVID to Enhance Recovery (RECOVER) Initiative. RECOVER-Pediatrics is an observational meta-cohort study of caregiver-child pairs (birth through 17 years) and young adults (18 through 25 years), recruited from more than 100 sites across the US. This report focuses on two of …


Impacts Of Urbanization On Demographics And The Stress Level Of White-Tailed Deer In The Piedmont Of South Carolina, Cameron Bramhall May 2024

Impacts Of Urbanization On Demographics And The Stress Level Of White-Tailed Deer In The Piedmont Of South Carolina, Cameron Bramhall

Graduate Theses

Population demographics and density studies are a common way of determining number of individuals in location. Understanding how many individuals and sex ratios can help estimate total populations for future potential management practices. For white-tailed deer (Odocoileus virginianus), commonly referred to as whitetail deer, like other cervids, having too many or too few individuals in an area can cause issues. Some of the issues that may arise with too many individuals include higher disease potential, and a general lack of resources leading to an increase in competition. As areas that were once habitat for whitetail deer become more and more …


West Coast Rock Lobster Resource Harvest Strategy, Department Of Primary Industries And Regional Development, Western Australia May 2024

West Coast Rock Lobster Resource Harvest Strategy, Department Of Primary Industries And Regional Development, Western Australia

Fisheries Management Papers

Harvest strategies for Western Australia’s (WA) aquatic resources are formal documents developed by the Department of Primary Industries and Regional Development (DPIRD, the Department) to support decision-making processes that ensure the outcomes are consistent with the principles of Ecologically Sustainable Development (ESD; Fletcher 2002a) and Ecosystem Based Fisheries Management (EBFM; Fletcher et al. 2012). Harvest strategies are a key component of all contemporary fishery management systems and a requirement for certification under the Marine Stewardship Council (MSC). The objectives of ESD are reflected in the objectives of the Fish Resources Management Act 1994 (FRMA) and the Aquatic Resources Management Act …


Migration And Breeding Ecology Of King Rails (Rallus Elegans) In Southeast Arkansas, Jessica Schmit May 2024

Migration And Breeding Ecology Of King Rails (Rallus Elegans) In Southeast Arkansas, Jessica Schmit

Graduate Theses and Dissertations

King Rails (Rallus elegans) are a secretive marsh bird species of conservation concern in Arkansas. Freddie Black Choctaw West WMA in southeast Arkansas has multiple confirmed pairs of breeding King Rails and offered an opportunity to investigate King Rail migration ecology, as well as nesting ecology and nest site selection. In Chapter 1, we determined migratory status (resident vs. migratory) and described seasonal movement. To do this, we captured and outfitted 23 birds with Argos GPS tags to track migration patterns throughout the season. We found that most of the population did not make long distance migrations but instead were …


Survey Of The Bolctes From Fish River Nature Preserve, Noah T. Nelson May 2024

Survey Of The Bolctes From Fish River Nature Preserve, Noah T. Nelson

Honors Theses

Boletes are a special kind of mushroom inside of the Kingdom Fungi that are distinguished from other mushrooms by their uniquely pored hymenium, as opposed to gills. Many boletes are ectomycorrhizal, meaning they are mutualists with vascular plants, and thus beneficial to the overall health of the terrestrial ecosystem. Some are well-known for their edibility and are of economic importance. In the United States 59 genus-level clades comprising 290 operational taxonomic units have been reported. The southeastern US and Gulf Coast regions however are less researched and recorded for their mushroom ecology. In Alabama, only 56 bolete species have been …


Thamnophis And Their Toxic Relationship: Testing For Intraspecific Venom Variation In The Common Garter Snake, Thamnophis Sirtalis, Grace E. Coppinger May 2024

Thamnophis And Their Toxic Relationship: Testing For Intraspecific Venom Variation In The Common Garter Snake, Thamnophis Sirtalis, Grace E. Coppinger

Poster Presentations

Project Objective - Estimate intraspecific venom variation between male and female garter snakes collected from the University of South Alabama of different sizes.