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Articles 1 - 30 of 37
Full-Text Articles in Population Biology
Coho Salmon (Oncorhynchus Kisutch) Response To Habitat Restoration Through Space And Time: Monitoring Growth, Survival, And Movement In A Restored Habitat Network, Carter Perez Adamson
Coho Salmon (Oncorhynchus Kisutch) Response To Habitat Restoration Through Space And Time: Monitoring Growth, Survival, And Movement In A Restored Habitat Network, Carter Perez Adamson
Cal Poly Humboldt theses and projects
Despite the decline of coho salmon (Oncorhynchus kisutch), Prairie Creek, a tributary of Redwood Creek in northern California, remains a coho salmon stronghold. The restoration of juvenile coho rearing habitat was identified as a recovery objective in this watershed by the National Marine Fisheries Service. Among the restoration projects addressing this objective are an ongoing project on Prairie Creek and its tributary Skunk Cabbage Creek, and a prior effort on Strawberry Creek (a tributary of Redwood Creek’s estuary). Standard monitoring methods for assessing project success have limitations and may be improved by measuring salmon use of restored habitats …
Geographic Variation In Proteomic Responses To Ocean Acidification In A Cold-Water Coral (Balanophyllia Elegans), Keana A. Richmond
Geographic Variation In Proteomic Responses To Ocean Acidification In A Cold-Water Coral (Balanophyllia Elegans), Keana A. Richmond
Cal Poly Humboldt theses and projects
In the face of a rapidly changing climate, assessing organismal responses to future stressors in the context of current, natural exposure to stress could provide key insights to understanding marine ecosystem resilience. I used Balanophyllia elegans, a cold-water, solitary, azooxanthellate coral as a model to better understand how varying oceanographic conditions across its geographic range have shaped its ability to tolerate and potentially adapt to current and future ocean acidification conditions. I collected B. elegans individuals from four sites across 2,500km of their range and subjected them to two pH treatments to investigate site-specific protein expression in response to …
Jaguar (Panthera Onca) Population Status And Conservation Needs In Panama Revealed Using Spatial Density And Occupancy Models., Rebeca E. Becdach
Jaguar (Panthera Onca) Population Status And Conservation Needs In Panama Revealed Using Spatial Density And Occupancy Models., Rebeca E. Becdach
Cal Poly Humboldt theses and projects
Jaguars (Panthera onca) are key predators and the largest felid in the Americas. The species is listed as near threatened globally due to habitat loss, fragmentation, and human persecution. Their population decline can negatively affect the structure and functioning of the ecosystems they inhabit. Conservation efforts for jaguars and their habitat require understanding jaguar populations and how they are impacted by human presence across their range. This information is lacking in Panama, a critical portion of the Jaguar range. Panama forms a narrow land bridge connecting Central and South America, integral to wildlife movement, including jaguars, across both …
Bat Species Diversity In The Klamath Basin Prior To Dam Removals, Ryan Page Matilton
Bat Species Diversity In The Klamath Basin Prior To Dam Removals, Ryan Page Matilton
Cal Poly Humboldt theses and projects
River ecosystems make up a small percentage of the earth’s surface, but they support high levels of biodiversity and attract high levels of human use. Dams, for example, disrupt a river’s natural course and flow, alter water temperatures in the stream, redirect river channels, transform floodplains, and fragment river continuity, which affects ecological balance and food webs. The Klamath River in northern California, which has been dammed for the last century, experienced dramatic disruptions including a large fish die-off in 2002. With the negative impacts of the Klamath dams apparent, multiple entities negotiated to remove the dams and restore the …
Just Keep Swimming: Outmigration Survival Of Upper Klamath Basin Juvenile Chinook Salmon, James M. Whelan
Just Keep Swimming: Outmigration Survival Of Upper Klamath Basin Juvenile Chinook Salmon, James M. Whelan
Cal Poly Humboldt theses and projects
The removal of four impassable hydroelectric dams on the Klamath River provides salmonids with the opportunity to recolonize their historic range in the upper Klamath River watershed. However, successful reestablishment of salmon populations requires juvenile salmon born in Oregon's upper tributaries to safely migrate downstream through the river's upper reaches. This stretch of the Klamath River, immediately downstream of Upper Klamath Lake, presents multiple migration challenges, including two flow-control dams not scheduled for removal (Link River Dam and Keno Dam), significant water diversions, and water quality issues. To evaluate these challenges, I conducted experimental releases of radio-tagged juvenile Chinook salmon …
Characterizing The Habitat Use Of Pacific Coast Feeding Group Gray Whales (Eschrichtius Robustus) And The Spatial And Temporal Variability Of Their Benthic And Planktonic Invertebrate Prey In Northern California, Robyn E. Norman
Cal Poly Humboldt theses and projects
As opportunistic foragers, the Eastern North Pacific population (~20,000) of gray whales (Eschrichtius robustus) feed on diverse benthic and planktonic invertebrates in northern Alaska foraging grounds before they undertake one of the largest yearly migrations of any mammal to breed in Baja California, Mexico. While most of the population travels to the summer foraging grounds in Alaska, a sub-group of whales (~230) called the Pacific Coast Feeding Group (PCFG) summer between British Columbia, Canada, and northern California. The diet of PCFG whales typically includes high-density and/or high-caloric prey items like mysids and diverse species of amphipods, yet a …
Post-Fire Vegetation And Pyrodiversity Influence Breeding Abundances Of Lewis's Woodpeckers In The Eastern Cascades, Oregon, Brittany Welch
Post-Fire Vegetation And Pyrodiversity Influence Breeding Abundances Of Lewis's Woodpeckers In The Eastern Cascades, Oregon, Brittany Welch
Cal Poly Humboldt theses and projects
Lewis’s woodpeckers (Melanerpes lewis) are described as "burn specialists" due to their preference for breeding in recently burned pine forests in the western US. However, despite increasing fire activity, this species experienced a 48% range-wide decline between 1968 and 2019, which raises questions about their adaptability to altered fire regimes in the region. We partnered with the Oregon Department of Fish & Wildlife to investigate how Lewis’s woodpeckers in the eastern Cascades, Oregon were influenced by post-fire habitat characteristics such as snag size, snag density, and burn severity across five wildfires that varied in fire age, size, and …
Evaluating The Trojan Y Chromosome Strategy For The Removal Of Invasive Sacramento Pikeminnow From The Eel River, Ca, Alexander W. Juan
Evaluating The Trojan Y Chromosome Strategy For The Removal Of Invasive Sacramento Pikeminnow From The Eel River, Ca, Alexander W. Juan
Cal Poly Humboldt theses and projects
The recent introduction and spread of Sacramento Pikeminnow (Ptychocheilus grandis) in Northern California’s Eel River Basin represents a significant threat and impediment to the recovery of several threatened native fish species. This study was undertaken to evaluate the Trojan Y Chromosome Strategy (TYC) for the extirpation of pikeminnow from the basin. TYC is a genetic biocontrol method that relies on the production and stocking of fish with YY sex chromosomes, which may be phenotypically male (YY male) or female (YY female). These YY fish produce all-male offspring when mating with their wild conspecifics and TYC can lead to …
Using Novel And Traditional Survey Techniques To Monitor Small Mammal Species In Northwestern California, Sydney Mccluskey
Using Novel And Traditional Survey Techniques To Monitor Small Mammal Species In Northwestern California, Sydney Mccluskey
Cal Poly Humboldt theses and projects
Small mammal species play a critical role in forest ecosystems contributing significantly to overall forest biodiversity and ecological dynamics. Forest-dwelling species are among the most vulnerable to habitat loss and fragmentation. It is essential to develop efficient techniques for monitoring small mammal populations in forested ecosystems to inform conservation efforts and management plans; yet, monitoring small mammals in these habitats can be particularly challenging due to their size, activity patterns, and frequently nocturnal nature. In this study, I aimed to address two primary research objectives: 1) compare the effectiveness of 3 distinct camera-trap survey techniques (i.e., ground, tree, and tube) …
Bird Predation On Coho Salmon (Oncorhynchus Kisutch) In Humboldt Bay, Colton J. Dixon
Bird Predation On Coho Salmon (Oncorhynchus Kisutch) In Humboldt Bay, Colton J. Dixon
Cal Poly Humboldt theses and projects
Coho Salmon (Oncorhynchus kisutch) populations along the west coast of the United States are declining. In Humboldt County, located in northwestern California, Coho Salmon have been in decline since their federal listing as threatened in 1997, primarily due to habitat loss from logging, agriculture, and urban development. Although these anthropogenic factors caused salmon populations to decline, bird predation may also be hindering population recovery, as evidenced by the recovery of Passive Integrated Transponder (PIT) tags, originally implanted in Coho Salmon, from bird colonies in Humboldt Bay. This study examines predation by Double-crested Cormorants (Phalacrocorax auritus), Caspian …
Pond Characteristics That Influence Oviposition Site Selection By Two Pond-Breeding Amphibians, Northern Red-Legged Frog (Rana Aurora) And Northwestern Salamander (Ambystoma Gracile), Jessica Jones
Cal Poly Humboldt theses and projects
Pond-breeding amphibians connect aquatic and terrestrial habitats through their biphasic life cycle, and understanding pond characteristics that support oviposition sites for multiple species is important for amphibian conservation. Two common amphibians in the Pacific Northwest, Northern red-legged frog (Rana aurora) and Northwestern salamander (Ambystoma gracile) often oviposit in the same pond; however, prior studies have focused on pond use by one species or the other, but not both together. I surveyed pond and oviposition site characteristics of 26 ponds in Humboldt County, CA, 10 of which were used only by R. aurora and 16 of which …
Scale And Habitat Effects On Measurement Of Streptomyces Biogeography And Biodiversity, Terilyn A. Stoflet
Scale And Habitat Effects On Measurement Of Streptomyces Biogeography And Biodiversity, Terilyn A. Stoflet
Cal Poly Humboldt theses and projects
Bacteria in the genus Streptomyces are detected in most soils around the world, as well as on and in seeds, plants, animals, insects and the wind. Currently, more than 700 valid species of Streptomyces have been identified worldwide. Streptomyces produce the majority of antibiotics, antivirals and antifungals, as well as important agricultural bioactive molecules due to their unique genetic capabilities. Their distinct genetic abilities allow them to produce an unlimited amount of biosynthetic gene clusters, or BGC’s. Although many compounds come from Streptomyces, scientists have not discovered the biogeographic patterns of this genus. The biogeography of Streptomyces is a …
Interactions Between New Zealand Mud Snails (Potamopyrgus Antipodarum) And Toxic Cyanoabacteria (Anabaena) In Baduwa’T (Mad River), Humboldt County, California, Cecilia F. Kennelly
Interactions Between New Zealand Mud Snails (Potamopyrgus Antipodarum) And Toxic Cyanoabacteria (Anabaena) In Baduwa’T (Mad River), Humboldt County, California, Cecilia F. Kennelly
Cal Poly Humboldt theses and projects
Invasive species in freshwater systems are circulated around the globe, wreaking havoc on natural communities. The New Zealand mud snail (Potamopyrgus antipodarum) is a very successful aquatic invader that has established populations in freshwater streams, lakes, and rivers around the globe. The goal of this project was to evaluate the interactions of invasive New Zealand mud snails (NZMS) and toxic cyanobacteria Anabaena in the Mad River in Humboldt County, California. These two species of concern are found in similar habitats along the river, but their whether NZMS consume Anabaena or affect toxin production is unknown. First, I conducted …
Insights Into The Conservation Of The Rare Perennial Herb, Astragalus Applegatei, Sean Stewart
Insights Into The Conservation Of The Rare Perennial Herb, Astragalus Applegatei, Sean Stewart
Cal Poly Humboldt theses and projects
As anthropogenic activities expand, conservation of rare and endangered species is critical to reduce extinction risks. Collecting and analyzing demographic data can assist in the development of conservation strategies and can help evaluate the success of existing strategies. In this study, I built integral projection models from five years of demographic data and conducted a population viability analysis on four populations of the rare perennial herb, Astragalus applegatei, to inform conservation decisions. I also assessed how planting A. applegatei seedlings might contribute to the growth of an extant population. I found that one population was already below the extinction …
Western Gulls’ (Larus Occidentalis) Reproductive And Behavioral Responses To Human Disturbance, Lilamarie 'Asta Bowen
Western Gulls’ (Larus Occidentalis) Reproductive And Behavioral Responses To Human Disturbance, Lilamarie 'Asta Bowen
Cal Poly Humboldt theses and projects
Human disturbances can negatively affect wildlife by causing stress, altering behavior, or even impacting populations through changes in survival or productivity. Colonial-nesting seabirds are of particular concern due to population declines and their gregarious and conspicuous nature, which may attract human visitors. However, the effects of nearby human activities, though frequently negative, could be neutral or even positive through phenomena like habituation to human activities or subsidization by human-supplied food sources. In this observational study, I evaluated relative support for three hypotheses which may explain how Western Gulls are impacted by human activity: disturbance, habituation, and subsidization. I investigated this …
Carnivore And Ungulate Occurrence In A Fire-Prone Region, Sara J. Moriarty-Graves
Carnivore And Ungulate Occurrence In A Fire-Prone Region, Sara J. Moriarty-Graves
Cal Poly Humboldt theses and projects
Increasing fire size and severity in the western United States causes changes to ecosystems, species’ habitat use, and interspecific interactions. Wide-ranging carnivore and ungulate mammalian species and their interactions may be influenced by an increase in fire activity in northern California. Depending on the fire characteristics, ungulates may benefit from burned habitat due to an increase in forage availability, while carnivore species may be differentially impacted, but ultimately driven by bottom-up processes from a shift in prey availability. I used a three-step approach to estimate the single-species occupancy of four large mammal species: mountain lion (Puma concolor), coyote …
Sympatric Carnivores And Vegetation Structure Influence The Distribution And Abundance Of Humboldt Martens In Northern California, Erika L. Anderson
Sympatric Carnivores And Vegetation Structure Influence The Distribution And Abundance Of Humboldt Martens In Northern California, Erika L. Anderson
Cal Poly Humboldt theses and projects
The Humboldt marten (Martes caurina humboldtensis) is a species facing conservation concern in Oregon and California. Historical populations were subjected to unregulated fur trapping and timber harvest throughout the late 19th and early 20th centuries leading to a 90% decline in their distribution. As such, the Humboldt marten is listed as a state endangered species in California, and the analogous ‘coastal marten’ is listed as federally threatened under the Endangered Species Act and classified as a Species of Greatest Conservation Need in Oregon and California. Contemporary populations are small, and isolated, and threatened by habitat loss …
Identifying Habitat Suitability And Connectivity For Roosevelt Elk (Cervus Canadensis Roosevelti) On The North Coast Of California, Ashley Ann Harper
Identifying Habitat Suitability And Connectivity For Roosevelt Elk (Cervus Canadensis Roosevelti) On The North Coast Of California, Ashley Ann Harper
Cal Poly Humboldt theses and projects
As human development continues to expand, wildlife managers are faced with problems arising from human-wildlife conflict. To address conflict, it is vital to identify how animals view the landscape and move across areas of differing land use. On the North Coast of California, Roosevelt elk (Cervus canadensis roosevelti) are widely considered to be a valuable resource but are viewed by some as a nuisance due to economic losses caused by property damage. The aim of this study was to identify suitable habitat and movement corridors for Roosevelt elk across Humboldt and Del Norte counties in northern California. Suitable …
Forecasting The Outcomes Of Managing A Rare Endemic Plant At Its Elevational Limit Under Climate Change Scenarios, Ethan Reibsome
Forecasting The Outcomes Of Managing A Rare Endemic Plant At Its Elevational Limit Under Climate Change Scenarios, Ethan Reibsome
Cal Poly Humboldt theses and projects
Lupinus constancei (Lassics lupine) is an exceptionally rare forb with a population that in 2020 consisted of 1,300 individuals located on two adjacent mountaintops in northwestern California, USA. Past work has demonstrated the species is threatened by high levels of seed predation by small mammals, which is likely the result of fire exclusion and the subsequent vegetation encroachment that both increases small mammal habitat and directly reduces suitable habitat for the species. Because of this, active management through caging of reproductive individuals to reduce seed consumption has been conducted every year since 2003. In addition, growing season temperatures have increased …
Evaluating Relative Abundance, Fish Length, And Marine Protected Area Effectiveness For Four Key Rocky Reef Species Along The Northern Californian Coast, Leon Davis
Cal Poly Humboldt theses and projects
Marine protected areas (MPAs) were created with the purpose of helping conserve and restore diminished populations of marine organisms. Measuring the effectiveness of MPAs requires long-term monitoring, investigating the abundance and size distributions of the species that utilize the conservation areas, and comparing the results to neighboring reference sites that are not currently protected. In this study, observations from long-term MPA monitoring in northern California (2010-2019) were modeled with substrate, oceanographic, spatial, temporal, and body size variables to describe the variability in abundance and size of three fish groups: Black rockfish (Sebastes melanops), the Blue rockfish group (comprised …
Degradation And Nitrogen Cycling In The Context Of Biodiversity-Ecosystem Function Relationships In The Inquiline Bacterial Community Of Darlingtonia Californica, Megan Teigen
Cal Poly Humboldt theses and projects
Biodiversity-ecosystem function (BEF) research aims to explain how species and their environments interact with each other. Microbial communities engage in vital biogeochemical pathways in a variety of natural ecosystems, and yet there are large knowledge gaps about the specific metabolic pathways in which they are involved. Degradation specifically contributes to nitrogen cycling globally through the breakdown of large organic nitrogen compounds into small inorganic nitrogen that is necessary for the survival of many other organisms. In this study, I focused on the degradative function of the inquiline microbial communities found within the carnivorous pitcher plant, Darlingtonia californica. Darlingtonia grows in …
Demographic Modeling Of Conservation Strategies For The Yosemite Toad (Anaxyrus Canorus), Corrina Rebecca Kamoroff
Demographic Modeling Of Conservation Strategies For The Yosemite Toad (Anaxyrus Canorus), Corrina Rebecca Kamoroff
Cal Poly Humboldt theses and projects
The Yosemite toad (Anaxyrus canorus) is an anuran species endemic to the Sierra Nevada in California that, like many amphibians globally, has suffered population declines. The documented decline in A. canorus populations across their historic range highlights the need for an effective management strategy to protect the species from future extirpation. For this study, I estimated survival rates of A. canorus using a Cormack-Jolly-Seber model populated with data from a demographic study. I then used a female-only post-birth pulse stochastic Lefkovitch matrix model using vital rates I estimated and from the literature to simulate the effect of different …
Noninvasive Genetic Sampling With A Bayesian Spatial Capture-Recapture Analysis To Estimate Abundance Of Roosevelt Elk (Cervus Canadensis Roosevelti), Makenzie Henk
Cal Poly Humboldt theses and projects
Determining abundance of Roosevelt elk (Cervus canadensis roosevelti) in central Humboldt County, California has presented a unique challenge to wildlife managers due to the dense forest habitat and the animals’ elusive behavior. As the elk population has increased, so has human-wildlife conflict, and wildlife agencies need efficient and repeatable methods for determining abundance to inform management decisions. Traditional monitoring methods such as helicopter surveys are ineffective due to low sighting probability and strong behavioral responses to the aircraft. They also often lead to biased sex ratios when the distribution of males and females varies across the landscape. Non-invasive …
Comparison Of Standard And Environmental Dna Methods For Estimating Chinook Salmon Smolt Abundance In The Klamath River, California, Doyle Joseph Coyne
Comparison Of Standard And Environmental Dna Methods For Estimating Chinook Salmon Smolt Abundance In The Klamath River, California, Doyle Joseph Coyne
Cal Poly Humboldt theses and projects
Evaluating abundance of juvenile salmonids is critical to conservation and management. Current abundance estimation involves use of rotary screw traps and mark-recapture studies. Use of environmental DNA (eDNA) in water samples offers a noninvasive and less expensive approach that may potentially improve or eventually replace traditional monitoring. Here I evaluate the utility of eDNA to predict weekly abundance estimates of outmigrating Chinook salmon (Oncorhynchus tshawytscha) smolts in the Klamath River, California. A total of 15 water samples were collected per week over the 17-week smolt outmigration in both 2019 and 2020. Chinook salmon eDNA concentration in each water …
Phylogenetic And Population Genetic Analysis Of The Humboldt's Flying Squirrel Using High-Throughput Sequencing Data, Stella Chi Yuan
Phylogenetic And Population Genetic Analysis Of The Humboldt's Flying Squirrel Using High-Throughput Sequencing Data, Stella Chi Yuan
Cal Poly Humboldt theses and projects
The intraspecific genetic variation and diversity within the Humboldt’s flying squirrel (Glaucomys oregonensis) has not yet been characterized despite its elevation to full species in 2017. The San Bernardino flying squirrel (G. o. californicus) is thought to be the southernmost population of G. oregonensis and is restricted to the San Bernardino and San Jacinto Mountains in California, but recent surveys indicate they have been extirpated from the latter locality. In order to provide baseline genetic data across the geographic range of G. oregonensis, I had the following objectives: 1) investigate the intraspecific molecular variation in …
The Effects Of Woody Vegetation Encroachment And Removal Within A Coastal Fen, Joseph Saler
The Effects Of Woody Vegetation Encroachment And Removal Within A Coastal Fen, Joseph Saler
Cal Poly Humboldt theses and projects
Early successional wetland habitat is being lost in temperate regions worldwide as a result of changes in disturbance regimes that allow for the establishment and dominance of woody species. In particular, this phenomenon is pronounced in fens, which harbor high numbers of rare herbaceous species that require early successional habitat. I investigated the relationship between woody vegetation encroachment and herbaceous species diversity within a Northern California coastal fen that has been undergoing encroachment by woody vegetation for ca. 80 years by recording species richness and cover data from 338 permanent plots throughout the fen. I also investigated the effect of …
Spatial And Temporal Genetic Structure Of Winter-Run Steelhead (Oncorhynchus Mykiss) Returning To The Mad River, California, Steven R. Fong
Spatial And Temporal Genetic Structure Of Winter-Run Steelhead (Oncorhynchus Mykiss) Returning To The Mad River, California, Steven R. Fong
Cal Poly Humboldt theses and projects
Distinct populations of steelhead in the wild are in decline. The propagation of steelhead in hatcheries has been used to boost population numbers for recreational fisheries and for use in conservation. However, hatchery breeding practices of steelhead can result in changes in genetic structure. I investigated the genetic structure of winter-run steelhead (Oncorhynchus mykiss) returning to the Mad River, California, where a hatchery has been used enhance production for recreational fisheries since 1971. Genetic variability in Mad River steelhead was evaluated using 96 single nucleotide polymorphisms (SNPs) among 4203 individuals, including the Mad River and nearby locations, and …
Multivariate Habitat-Based Predictive Modeling Of Three Demersal Rockfish Species In Central California, Georgia R. Martel
Multivariate Habitat-Based Predictive Modeling Of Three Demersal Rockfish Species In Central California, Georgia R. Martel
Cal Poly Humboldt theses and projects
Accurate, spatially explicit models of rockfish abundance are critical in implementing ecosystem-based management strategies and designating essential fish habitats and marine protected areas. Multibeam bathymetry methods and visual, non-extractive submersible transect surveys were combined to collect environmental variables and fish abundance data at three distinct sites within the study region. Zero-adjusted models were developed using habitat classification analyses of high-resolution (5 m) digital elevation models. Model accuracies were assessed by using a reserved subset of the original datasets. To demonstrate that a model’s predictive power was linked to its spatial origins, Mean Absolute Error and coefficient of determination values were …
A Tale Of Two Cervids: Habitat Selection And Spatial Ecology Of Roosevelt And Tule Elk In California, Adam S. Mohr
A Tale Of Two Cervids: Habitat Selection And Spatial Ecology Of Roosevelt And Tule Elk In California, Adam S. Mohr
Cal Poly Humboldt theses and projects
California’s Roosevelt (Cervus canadensis rooseveltii) and tule elk (C. c. nannodes) populations have experienced a remarkable recovery after over-hunting and habitat loss nearly extirpated them from the state. Greater clarity of Roosevelt and tule elk habitat selection patterns would help managers continue to effectively support the recovery of these two iconic subspecies. As the manner in which elk populations balance their competing resource needs is unique to each population (Skovlin et al. 2002), I have examined Roosevelt and tule elk habitat selection patterns in separate analyses and present these results in independent chapters. In both cases, …
Estimates Of Calf Survival And Factors Influencing Roosevelt Elk Mortality In Northwestern California, Erin Nigon
Estimates Of Calf Survival And Factors Influencing Roosevelt Elk Mortality In Northwestern California, Erin Nigon
Cal Poly Humboldt theses and projects
Survival of juvenile ungulates is known to be highly variable, yet it is fundamental to understanding the dynamics and trends of wildlife populations. Factors influencing calf survival are poorly understood in northwestern California where Roosevelt elk (Cervus canadensis roosevelti) are known to exist. The objectives of this study were 1) to estimate summer and annual survival rates and determine recruitment rates for Roosevelt elk calves in the area, 2) to evaluate differences in calf survival by examining the effects of individual and population level covariates on summer and annual calf survival, 3) to identify factors influencing the timing …