Zoology Commons^™

Zinc Content In The Organs And Tissues Of Freshwater Fish From The Kardjali And Studen Kladenets Dam Lakes In Bulgaria, Iliana G. Velcheva

Turkish Journal of Zoology

A study was conducted to estimate zinc loads in the organs and tissues of Alburnus alburnus, Cyprinus carpio and Perca fluviatilis inhabiting 2 dam lakes Kardjali and Studen Kladenets, located along the Arda river (South Bulgaria). The zinc content was determined by atomic absorption spectrophotometry using a Perkin Elmer 380 B apparatus .The effect of the temporal factor was proved over a 3-year study. The highest zinc concentrations in fish organs and tissues were detected in summer and autumn. The highest zinc loads were found in the kidney and liver, and the lowest in the muscles of the 3 fish …

Long-Distance Dispersal And Population Trends Of Moose In The Central United States, Justin D. Hoffman, Hugh H. Genoways, Jerry R. Choate

University of Nebraska State Museum: Mammalogy Papers

Dispersal is a basic feature of the natural history of moose. Most information about moose dispersal pertains to short-distance movements because long-distance movements are uncommon and difficult to observe. Since the 1950s, moose populations have been increasing in Minnesota and North Dakota. This may have contributed to several long-distance dispersal events for moose that recently were reported in the central United States. These dispersal events provide an opportunity to investigate both the causes and the biological implications of this rare phenomenon. Herein, we review long-distance dispersal events based on information obtained from a variety of sources. Dispersal routes that could …

Hyla Squirella (Squirrel Treefrog): Refugia., Louis A. Somma, David Serrano

Papers in Herpetology

HYLA SQUiREUA (Squirrel Treefrog). REFUGIA. Entomologists commonly use trap-nests to monitor cavity-nesting Hymenoptera. These trap-nests consist of 5 x 10 cm pine timber cut into 12-cm lengths. Five lengths are then stacked and strapped together. Each pIece of timber is predrilled in each side to have either a 3.2, 4.8, 6.4, 7.9, or 12.7 Iiundiameterhole, 8 cmindepth. Thus, each trap-nest consisted of five pieces of wood containing 2 holes of each diameter. They are suspended with wire from tree trunks and branches. We set these wooden traps (N = 10) at Devil's Millhopper Geological State Park, Gainesville, Alachua Co., Florida, …

Parelaphostrongylus Odocoilei In Columbia Black-Tailed Deer From Oregon, Jack A. Mortenson, Arthur Abrams, Benjamin M. Rosenthal, Detiger Dunams, Eric P. Hoberg, Robert J. Bildfell, Richard L. Green

Harold W. Manter Laboratory of Parasitology: Faculty Publications

Documenting the occurrence of Parelaphostrongylus odocoilei has historically relied on the morphological examination of adult worms collected from the skeletal muscle of definitive hosts, including deer. Recent advances in the knowledge of protostrongylid genetic sequences now permit larvae to be identified. Dorsal-spined larvae (DSLs) collected in 2003–2004 from the lung and feces of six Columbian black-tailed deer (Odocoileus hemionus columbianus) from Oregon were characterized genetically. The sequences from unknown DSLs were compared to those from morphologically validated adults and larvae of P. odocoilei at both the second internal transcribed spacer (ITS-2) of ribosomal DNA and the mitochondrial cytochrome …

Leptodactylus Fragilis (Brocchi): White-Lipped Thin-Toed Frog, Miriam M. Heyer, W. Ronald Heyer, Rafael O. De Sá

Biology Faculty Publications

Adult Leptodactylus fragilis are small. The head is longer than wide, and the hind limbs are moderately long (Table 1; Heyer and Thompson 2000 provided definitions of adult size and leg length categories for Leptodactylus). Male vocal sacs are laterally expanded and range from lightly gray speckled to darkly pigmented. Male snout is spatulate; female snout is rounded. Male forearms are not hypertrophied and males lack asperities on the thumbs and chest. The dorsum is spotted or blotched with dark markings, which are often chevron-shaped and sometimes confluent. The supratympanic fold is dark brown. A pair of dorsolateral folds …

An Examination Of Seasonal Growth And Survivorship Of Sigmodon Hispidus In Southeastern Virginia, Heather Alyssa Green

Biological Sciences Theses & Dissertations

A reexamination of specific population dynamic aspects of Sigmodon hispidus, the hispid cotton rat, is necessary in order to gain additional knowledge and perspective on this species in its northernmost distribution on the east coast of the United States. Previous studies of Virginia cotton rats were based on data from necropsies, which do not allow for the determination of certain population characteristics, such as density, growth, and survival. General population trends and rates of growth and survival in males and females throughout the seasons in Virginia will also provide a basis for comparison with populations of cotton rats in …

Movement Patterns And Habitat Usage Of Shark Bay Dugongs, David K. Holley

Theses: Doctorates and Masters

In order to define small and large scale spatial and temporal individual movement patterns of dugongs (Dugong dugon) within the Shark Bay World Heritage Property (SBWHP) a total of 19 dugongs were fitted with remote location recording and transmitting devices. Combined locations from all units totalled over 10,000 locations. This spatial and temporal data was used to define movement patterns of dugongs within Shark Bay as well as areas of high use deemed to be indicative of foraging activity. Platform Transmitting Terminals (PTT?s) using the ARGOS location collection system tracked animals over large temporal scales with 4 animals tracked up …

The Dynamics Of Bay Anchovy In The Hudson River Estuary: Process-Oriented Studies And Long-Term Changes, Eric T. Schultz, Kamazima M. M. Lwiza, John R. Young, Kyle J. Hartman, R. C. Tipton

EEB Articles

We review three areas of recent research on Hudson River bay anchovy. One focus has been the along-estuary movement of early life stages. A cohort analysis of samples collected in a spatiotemporally extensive monitoring program has confirmed that early-stage anchovy migrate up-estuary, at an estimated rate of 0.6 km/d. Complementary fine-scale field sampling was designed to clarify behaviors that effect the migration. This work found that early-stage anchovy can show preferences for depth and can conduct periodic vertical migration. To determine whether these behaviors were sufficient to produce up-estuary migration, larval flux and velocity were estimated. These estimates were consistent …

Results Of The Alcoa Foundation-Suriname Expeditions. Xiv. Mammals Of Brownsberg Nature Park, Suriname, Burton K. Lim, Mark D. Engstrom, Hugh H. Genoways, François M. Catzeflis, Kelly A. Fitzgerald, Sandra L. Peters, Marijem Djosetro, Sandra Brandon, Sutrisno Mitro

University of Nebraska State Museum: Mammalogy Papers

An inventory of mammals in the vicinity of Brownsberg Nature Park, Suriname, incorporated a number of different sampling methods including examining museum voucher specimens, an animal-rescue operation, transect surveys, camera photo-traps, and interviews with local inhabitants. We document a total of 125 mammal species present in the Park. These include ten opossums, five pilosans, four armadillos, 58 bats, eight monkeys, 13 carnivores, one tapir, four artiodactyls, and 22 rodents. Nine of these species are reported for the first time from Suriname: one mouse opossum (Marmosops pinheiroi); one naked-backed moustached bat (Pteronotus gymnonotus); four fruit-eating bats ( …

Index To Volume 73

Nebraska Bird Review

Adams, Chris 32

Alexander,

George 32

Irene 32

Amiotte, Sue 97, 136

Anderson,

Kari 31

Riley 28

Anhinga 46, 51

Armknecht, Henry 4, 32, 96

Audubon Society 22

Aughey, Samuel 125

Avocet, American 46, 54, 87, 102, 140

Fall Field Report, August–November 2005, W. Ross Silcock

Nebraska Bird Review

This fall was notable for its lack of notable events; numbers of migrants reported in all species groups, especially waterfowl, shorebirds, and warblers, were low. Lingering birds were both insectivores and seed-eaters, and, coupled with a rather stable fall weather-wise, it seems that there were no disturbances to concentrate birds or otherwise impede a steady flow of migrants through the state. Without concentrations, migrants are often not noticed or, more accurately here, not reported by observers.

The best Western Grebe count was 2500, very low relative to expected high fall counts of at least 20,000. Mergansers were the only bright …

Fall Field Days At Harlan County Reservoir

Nebraska Bird Review

The annual Fall Field Days were held on September 9–11, 2005, at the 4-H Camp near Alma. Field trip destinations included Harlan County Reservoir and various other points of interest in Furnas, Harlan, Franklin, Phelps, Kearney, and Adams Counties. The total tally was 119 species, including Snowy Egret, Yellow-crowned Night-Heron, Mississippi Kite, Prairie Falcon, and Olive-sided Flycatcher.

Fifty-four members and guests attended. On Friday the group heard from Dr. Josef Kren on the topic of obligate brood parasites, species that have lost the ability to construct nests and incubate eggs and instead lay eggs in the nests of other species, …

Suggested Areas For Study, William F. Rapp Jr.

Nebraska Bird Review

After 100 years of organized ornithology we have a good picture of the bird fauna of Nebraska. Yes, we will still be able to add new species to the state list, but this will become more and more difficult. What problems are left for the amateur ornithologist to solve? Nebraska is a big state with many different ecological areas, few of which have been studied in detail. Only a few areas of the state have been well studied. Not only do we need data on what species occur, but how many. This type of study can be done by amateur …

Snowy Plover Nesting At Lake Mcconaughy In 2004 And 2005, Mark M. Peyton, Gabriel Wilson

Nebraska Bird Review

The Snowy Plover (Charadrius alexandrinus) is a small shore bird that upon casual observation resembles the Piping Plover (Charadrius melodus). The most obvious differences are in the dark beak and legs of the Snowy Plover in contrast to the orange bill with the black tip and orange legs of the Piping Plover. The two also have strikingly different calls (pers. obs.). In Nebraska the Piping Plover can be found along the Missouri, Platte, Niobrara, Loup, and Elkhorn Rivers and the sandpits associated with those rivers (Haig and Plissner 1993). Piping Plovers are also common along the …

Nebraska Bird Review (December 2005) 73(4), Whole Issue

Nebraska Bird Review

Fall Field Report, August–November 2005 ... 134

Fall Field Days at Harlan County Reservoir ... 150

Snowy Plover Nesting at Lake McConaughy in 2004 and 2005 ... 154

Suggested Areas for Study ... 157

Index to Volume 73 ... 160

Subscription and Organization Information ... 171

Subscription And Organization Information [December 2005]

Nebraska Bird Review

The Nebraska Bird Review is published quarterly by the Nebraska Ornithologists' Union, Inc., as its official journal, and is sent to members not in arrears of dues. Annual subscription rates (on a calendar-year basis only): $15 in the United States, $18 in Canada, and $30 in all other countries, payable in advance. Single copies are $4 each, postpaid, in the United States, $5 in Canada, and $8 elsewhere. Send orders for back issues to Mary Lou Pritchard, NOU Librarian, c/o University of Nebraska State Museum, W-436 Nebraska Hall, Lincoln, NE 68588-0514.

Memberships in the NOU (on a calendar-year basis only): …

Comparing Immune Responses Of Juvenile And Adult Northern Fence Lizards, Sceloporus Undulatus, Zaïna Sandrine Kabange

All-Inclusive List of Electronic Theses and Dissertations

Sex steroid hormones affect behavior and morphological traits important in sexual selection. Steroid hormones also play a role in the development and functioning of immune system components. The immunocompetence handicap hypothesis (Folstad and Karter, 1992) proposes that tradeoffs, mediated by plasma levels of testosterone, exist between the immune system and elaboration of secondary sexual traits. Endocrine-immune interactions are well documented in birds and mammals, and steroid hormones can be immunoenhancing or immunosuppressive. Relatively little work has examined the ontogeny of immune function, and the interactions between steroid hormones and the immune system in reptiles. The first experiment addressed the question …

Hybridization Of Lythrurus Fasciolaris (Cope) And Lythrurus Umbratilis (Girard) (Cypriniformes: Cyprinidae) In The Ohio River Basin, Robert L. Hopkins Ii

Morehead State Theses and Dissertations

A thesis presented to the faculty of the College of Science and Technology at Morehead State University in partial fulfillment of the requirements for the Degree of Master of Science by Robert L. Hopkins, II on November 14, 2005.

A Phylogenetic Analysis Of Vanzolinius Heyer, 1974 (Amphibia, Anura, Leptodactylidae): Taxonomic And Life History Implications, Rafael O. De Sá, W. Ronald Heyer, Arley Camargo

Biology Faculty Publications

The validity of the monotypic leptodactylid frog genus Vanzolinius Heyer, 1974 has been questioned recently. We explore the relationship of Vanzolinius discodactylus within the cluster of closely related genera Adenomera, Leptodacylus, and Lithodytes with both morphological and molecular data sets. Morphological and combined morphological and molecular data were analyzed using maximum parsimony; molecular data sets were analyzed with maximum likelihood methods. The resultant relationships are unambiguous in Vanzolinius being imbedded within Leptodactylus. In order to maintain Leptodactylus as a monophyletic genus, Vanzolinius is placed in the synonymy of Leptodactylus Fitzinger, 1826. The implications of relationships analyzed in …

Redescripción De La Morfología Larval Externa De Dos Especies Del Grupo De Leptodactylus Fuscus (Anura, Leptodactylidae), José A. Langone, Rafael O. De Sá

Biology Faculty Publications

Re-description of the larval external morphology of two species of the Leptodactylus fuscus group (Anura, Leptodactylidae). The larvae of Leptodactylus gracilis (D’Orbigny y Bibron, 1840) and L. mystacinus Bumeister, 1861 are re-described and compared with previous descriptions noting intraspecific variation in oral disc characteristics (arrangement of papillae and rows of cornified teeth). The external morphological analysis of larvae of the Leptodactylus fuscus group suggest that the lack of comparable descriptions among species, as well as the lack of analyses of their intraspecific variation, limits the use of larval characteristics for diagnostic purposes.

Mammalogy At Texas Tech University: A Historical Perspective, Lisa C. Bradley, John R. Suchecki, Brian R. Amman, Joel G. Brant, Hugh H. Genoways, L. Rex Mcaliley, Robert J. Baker, Francisca Mendez-Harclerode, Robert D. Bradley

University of Nebraska State Museum: Mammalogy Papers

The mammalogy program at Texas Tech University officially was established in 1962, when Robert L. Packard joined the faculty of the Department of Biological Sciences. As the institution's first mammalogist, Packard took the initiative to develop a strong program of mammalian research and education. Influenced by the successful program built by his mentor, E. Raymond Hall, at the University of Kansas, Packard modeled similar goals for Texas Tech University. Those goals included a strong emphasis on both undergraduate and graduate education and research, with several mammalogy faculty members, and the establishment and growth of a large and active mammal collection.

Online Dictionary Of Invertebrate Zoology: F, Mary Ann Basinger Maggenti, Armand R. Maggenti, Scott L. Gardner

Armand R. Maggenti Online Dictionary of Invertebrate Zoology

face facet facette facetted facial facialium facies facio-orbital factor facultative faeces Fahrenholz's falcate falces falciform falciger falcis falculate falsadentes false falx famulus fan fang farctate farinaceous fascia fasciate fascicle fasciculus fasciole fascio-maculata fascio-punctate fastening fastigiate fastigium fat fate fauces fauna faunal faveolate feces fecula fecundate fecundity feedback feeder feeding feeler feet felt female femoro-alary femorogenu femorotibial femur fenestra fenestrate fenestration fenestrella fenestrule fente feral ferment fermentation ferreous ferruginous ferrule fertile fertility fertilization fertilizin festivus festoon festoons fetid fiber fibril fibrillar fibrin fibrinogen fibrocyte fibroin fibroplasm fibrous fibula fide filament filamentary filaria filariform filate filator file filial filiation filibranch filicornia filiform …

Online Dictionary Of Invertebrate Zoology: B, Mary Ann Basinger Maggenti, Armand R. Maggenti, Scott L. Gardner

Armand R. Maggenti Online Dictionary of Invertebrate Zoology

baccate bacillary bacilliform back-cross back-mutation bacteremia bacteriophagous baculiform baculite baenomere baenopoda baenosome Baer's bailer balanced balancers balanoid balantin balantine Balbiani Balbiani's bald Baldwin ballonets ballooning band barb barbate, barbatus barbula bark baroceptor, baroreceptor barotropism barrier basad basal basalar basalia basalis base basement bases basicarnal basicerite basiconic basicosta basicostal basicoxite basifemoral basifemur basilaire basilar basilateral basilemma basimandibula basimaxilla basimetrical basinym basioccludent basiophthalmite basiperiphallus basipharynx basipod basipodite basiproboscis basipulvilli basirostral basis basiscopic basiscutal basisternum basistylus basitarsal basitarsus basitergal basitibial basitrich basitrichous basivalvula basket basopinacocytes batatiform Batesian bathmis bathyal bathylimnetic bathymetric bathypelagic battery batumen bave B-chromosome, bdelloid bead beaded beak beaked bean beard …

Online Dictionary Of Invertebrate Zoology: A, Mary Ann Basinger Maggenti, Armand R. Maggenti, Scott L. Gardner

Armand R. Maggenti Online Dictionary of Invertebrate Zoology

abactinal abambulacral A-band abapertural abapical abaxial abbreviate abcauline abdomen abduct aberrant abient abiocoen abiogenesis abiology abiotic abjugal ablation aboral aboriginal abortion abranchiate abreptor abscission absorption abterminal abullate abyssal abyssobenthos abyssopelagic acanthaceous acanthella acanthocephalans acanthocyst acanthodion acanthodrilin acanthoparia acanthophore acanthophorites acanthopod acanthosphenote acanthostegous acanthostyle acanthozooid acariasis Acarina acarinarium acarocecidium acarology acarophily acaryote acaudal acclimation acclivous accretion acelomate acentric Acephala acephalocyst acephalous acerata acervate acescence acetabulum achaetous achlamydate achroacyte achroglobin achroic achromasia achromatic A-chromosome achroous acia acicle aciculiform acidic acidobiontic acidophilic acidophobic acidotheca aciduric acies aciform acinacicate acinaciform acinarious aciniform acinose aclid acline aclitellate acme Acoelomata acoelous acone acontioids acraein Acraspeda acrembolic …

Online Dictionary Of Invertebrate Zoology: C, Mary Ann Basinger Maggenti, Armand R. Maggenti, Scott L. Gardner

Armand R. Maggenti Online Dictionary of Invertebrate Zoology

cadavericole caddis cadre caducous caecum Caenogaea caespitose Cainogea caisson calabar calamistrum calamus calathiform Calcarea calces calciferous calcipala calcospherites calice calicle calicoblastic caliology callous callow callus calobiosis calomus caloric calorigenic calorimetry calotte caltrop calva calx calyciform calycine calycle calyoptis calypto calyptra Calyptratae calyptron calyx camarodont camera camerostome campaniform campestral campi campodeiform campus Canadian canal canalaria canali cancellate cancrisocial cancroid cane canella canines canities cannibalism cannula canthariasis canthus capillary capillitium capita caprification capsule capuliform caput capylus caraboid carapace carbohydrates carcinoid carcinology cardate cardiac cardines cardinosternal cardinostipital cardioblasts cardiocoelomic cardo cardosubmental caria caridean caridoid carina carious carmine carnivore carnose Carolinian carolites carotene carotin …

Online Dictionary Of Invertebrate Zoology: D, Mary Ann Basinger Maggenti, Armand R. Maggenti, Scott L. Gardner

Armand R. Maggenti Online Dictionary of Invertebrate Zoology

dacryoid dactyl dactylethra dactylognathite dactyloid dactylopodite dactylopore dactylozooid dactylus dance daphnid dart dart sac Darwinism dauer larvae dauermodification daughter daughter cells daughter cyst day-eye De Vriesianism dealate dealation death deaurate decacanth decalcification decamerous decapodid larvae decathecal decephalic deciduous deck declinate declivitous declivous deconjugation decorticate decumbent decurved decussated dedetermination dedifferentiation defaunate defecate deferent deficiency definition definitive host deflected deflected front deflexed defoliator deformed degenerate degenerate code degeneration dehiscence dehiscent deirids delamination delimitation, diagnosis, differential diagnosis delthyrial delthyrium deltidial deltidium deltoid demanian demarcation deme demersal demibranchs demiplate demiprovinculum Demospongiae denatant dendriform dendrite dendritic thickening dendrobranch dendrogram dendroid dendron dendrophagous dendrophilous denematize denizen …

Online Dictionary Of Invertebrate Zoology: E, Mary Ann Basinger Maggenti, Armand R. Maggenti, Scott L. Gardner

Armand R. Maggenti Online Dictionary of Invertebrate Zoology

ear eave eaves eburnean ecalcarate ecarinate ecaudate ecbolic eccentric ecdemic ecdysial ecdysis ecdysone ecdysotrophic ecesis echinate echinating Echinodermata echinoderms echinopaedium echinostome echiopluteus Echiura echiurans ecitophile ecium eclectic eclipsed eclosion ecoclimate ecocline ecogeographical ecological ecology ecomorph ecomorphic economic ecoparasite ecophene ecophenotype eco¬phenotypic ecosite ecospecies ecosystem ecotone ecotype ecsoma ectad ectadenia ectal ectally ectoblast ectochone ectocochleate ectocommensal ectocrine ectocyst ectoderm ectodermal ectognathous ectohormones ectolecithal ectomere ectomesenchyme ectomesoderm ectoneural ectooecium ectoparasite ectopara¬sitic ectophagous ectophallus ectopic ectoplasm ectoproct Ectoprocta ectoprocts ectoptygma ectosomal ectosome ectospermalege ectostracum ectosymbion ectosymbi¬ont ectothermal ectotrophous ectozoon ecumenical edaphic edaphic factors edaphon edeagus edentate edentulous edge editum edoeagus effect effector efferent …

Online Dictionary Of Invertebrate Zoology: G, Mary Ann Basinger Maggenti, Armand R. Maggenti, Scott L. Gardner

Armand R. Maggenti Online Dictionary of Invertebrate Zoology

galea gallery gallicola gallicolae gallicolous galliphagous galvanotaxis galvanotropism gametangiogamy gametangium gamete gametocyte gametogamy gametogenesis gametogenetic gametogeny gametogonium gametogony gametotoky gamma gamobium gamodeme gamogenesis gamogonic gamogony gamones gamophase ganglia gangliate ganglion ganglionic gap gaper gaping gas gaseous gaster gasterostome gasterotheca gastraea gastraeum gastral gastralia gastric gastrilegous gastrocoele gastrocoelus gastrodermis gastro-ileal gastrolith gastroorbital Gastropoda gastropods gastropores Gastrotricha gastrotrichs gastrovascular gastrozooid gastrula gastrular gathering Gause's gelatigenous gelatin gelatine gelatinous geminate gemma gemmate gemmation gemmipara gemmipares gemmule gemmulostasin gena genacerore genae genal genaponta genatasinus gene genera generalized generation generative generitype generotype Gene's genesiology genetic genetics genic genicular geniculate genital genitalia genitointestinal genocline genocopy genodeme …

Online Dictionary Of Invertebrate Zoology: N, Mary Ann Basinger Maggenti, Armand R. Maggenti, Scott L. Gardner

Armand R. Maggenti Online Dictionary of Invertebrate Zoology

nacre nacreous naiad nail naked nanism nanoid nanitic worker nanometer nanoplankton nanozooid narcosis nasale nascent naso Nassanoff's gland Nassanov's gland Nassanov's gland nasus nasute natal natality rate natant natatory natatory lamellae nates naticid naticiform native natural classification artificial classification natural decrease natural increase natural group natural increase natural requeening supersedure natural selection naupliar eye nauplius eye naupliiform nauplius eye nauplius larva nautilicone nautiliform nautiloid nautiloid navicular neala neallotype neanic neap Nearctic neascus larva nebulous neck neck organ nuchal organ necrobiosis necrocytosis necrophagous necrophoresis necrophoric behavior necrosis necrotize nectar nectobenthic nectocalyx nectophore nectochaeta larva necton nekton nectophore pneumatophore nectopod nectosome nectozooid …

Online Dictionary Of Invertebrate Zoology: L, Mary Ann Basinger Maggenti, Armand R. Maggenti, Scott L. Gardner

Armand R. Maggenti Online Dictionary of Invertebrate Zoology

labella labellum labellar abductor apodeme labellar basal sclerites labellar mesial sclerite labellar sclerite labellum labial labial area labial disc labial glands labial gutter premental gutter labial kidneys labial nephridia labial lumen premental gutter labial mask labial nephridia labial palp/palpus palpi maxillary palp labial plate labial stipes labiostipes labial suture labial veil oral lappets labiate labidophorous labiella labiellum labile labiostipes labis socii labium bia inner lip labral labropalatum clypeopala¬tum labrum labra labrum-epipharynx lac lacerated lacinia lacinea laciniae lacinia mandibulae lacinia mobilis lacinia mobilis laciniate lacteous lactescent lactic acid lacuna ae lacunar system lacunose lacustrine laeotorma ae laeotropic lagena utriculus Lamarckism lamella …