Zoology Commons^™

Fire Scarring Does Not Significantly Impact Conifer Growth Response To Forest Fires In Utah, Connor B. Biser, Russell Torgersen, Otto Degroff, Steve Petersen

Student Works

BYU Wildfire Symposium 2026 - First Place Research Application, People's Choice Award

Wildfires are an increasingly prevalent disturbance in forests in the western United States, and play an essential role in ecosystem function. Micro-variations in fire dynamics and intensity can leave individual trees with varying levels of damage. We hypothesized that trees with fire-scars would suffer a growth depression directly following the disturbance. Core samples were taken for dendrochronological analysis from 185 conifers at 29 points, representing 11 fires from 6 years. Our findings showed a growth depression in the first year following the fire, with no significant difference between …

Maturation Of An Off-Channel Habitat Concept To Conserve Native Fishes In The Lower Colorado River, Paul C. Marsh, Thomas E. Dowling, Thomas F. Turner, Megan J. Osborne, Brian R. Kesner

Monographs of the Western North American Naturalist

Two endemic, “large river” fishes of the Colorado River basin of western North America, bonytail Gila elegans and razorback sucker Xyrauchen texanus, are among several critically endangered species in the system. Wild populations of bonytail are gone, and there are no self-sustaining populations of razorback sucker anywhere; reproduction occurs but recruitment does not. Both species have been under intensive management in the Lower Colorado River since the 1980s. Today, with the single exception of Lake Mead, remaining populations are composed entirely of repatriated individuals that depend on stocking for their continued existence. In 2003, a conceptual off-channel habitat (OCH) …

Reproductively Viable Population Of American Black Bears (Ursus Americanus) In Lowland Chihuahuan Desert Habitat Of Trans-Pecos Texas, Franklin D. Yancey Ii, Stephen Kasper

Western North American Naturalist

During a 6.5-year camera-trap study in Big Bend Ranch State Park (situated in the Trans-Pecos region of West Texas), we documented a persistent breeding population of American black bears (Ursus americanus) in the Solitario region of the park. The habitat in the Solitario, which is an isolated, eroded remnant of volcanic activity, is low-elevation Chihuahuan Desert scrub but includes significant and complex localized topography and prolonged water sources in the form of tinajas. From January 2016 to May 2022, camera traps captured 367,301 photographs. Of these, 868 contained one or more black bears, resulting in 1010 black bear …

First Documentation Of Least Chub In The Snake River Drainage In Idaho, Eric J. Billman, Dallin Stokes, Isaac Calderwood, Cody Diehl

Western North American Naturalist

The Least Chub (Iotichthys phlegethontis) is a small minnow endemic to the Bonneville Basin in Utah, where the current distribution of extant populations is limited to a few small spring complexes. We report on the first documentation of Least Chub in the upper Snake River drainage in Idaho. In June 2021, October 2021, and June 2022, we surveyed for small-bodied fishes in off-channel habitat of lower Henrys Fork in Cartier Slough Wildlife Management Area west of Rexburg, Idaho. Small-bodied fishes were sampled at 17 sites in off-channel habitat; 5 baited minnow traps (galvanized steel) were set for approximately …

Western Pearlshell (Margaritifera Falcata) Extirpation In The Smith River, Montana, With A Possible Link To Warming Water Temperatures, David M. Stagliano

Western North American Naturalist

Western pearlshell (Margaritifera falcata) populations are in decline across the species’ range in western North America. In 2008, Montana categorized the western pearlshell as an imperiled species of concern (S2) based on reduced detections at historically occupied sites and few viable populations. Between 2004 and 2014, we surveyed 24 Smith River reaches (61 total sites in the watershed) for the presence of western pearlshell populations. Despite historical and anecdotal descriptions of abundant western pearlshell in the Smith River, intensive structured surveys between 2004 and 2007 documented only 4 sections with low-density, nonviable western pearlshell populations (<10 individuals, large size classes [average total length = 70 mm], with no signs of recruitment). In 2014, we revisited these occupied sites to repeat surveys (expending significantly more search time) and observed no live individuals. This was surprising given that, although those populations would certainly die out without recruitment, older individuals can oftentimes persist for decades. Therefore, we consider the western pearlshell mussel extirpated from the Smith River watershed. We reviewed long-term USGS gage discharge and temperature data near the western pearlshell populations to evaluate the potential causal factors for, and timing of, the extirpation. From 3 July to 3 August 2007, water temperatures were at or above the thermal stress levels of the western pearlshell (>25 °C, …

Mexican Gartersnakes (Thamnophis Eques) Along The Gila River In Southwestern New Mexico, Keith Geluso

Western North American Naturalist

The Mexican gartersnake (Thamnophis eques) is a federally threatened species in the United States, with only a few isolated populations remaining in Arizona and New Mexico. For some populations, only limited knowledge exists regarding their status and natural history; thus, all observations are noteworthy. Along the Gila River in New Mexico, for example, only 2 observations of this species have been published, one in 1883 and one in 1973. Herein, I report on 6 additional observations of Mexican gartersnakes from along the Gila River in southwestern New Mexico. Five individuals were documented in riparian habitats close to or …

Puma (Puma Concolor) Sex Influences Diet In Southwest New Mexico, Kelly M.T. Bernard, Travis W. Perry, Nokubonga Mgqatsa

Western North American Naturalist

Puma (Puma concolor) is a wide-ranging, large felid species, and site-specific research on its diet is important for local management. Like the diets of other large felids, puma diets may differ between sex due to size dimorphism and between seasons due to changes in prey vulnerability and availability. We investigated the influence of sex and season on puma diet in southwest New Mexico in terms of prey species and size categories. Pumas (10 males, 6 females) were tracked with GPS collars for an average of one year per individual between February 2008 and July 2020. Puma location was …

Are Shrubs A Necessary Component Of The Habitat For Ammospermophilus Nelsoni In The San Joaquin Desert?, David J. Germano, Larry R. Saslaw

Western North American Naturalist

The San Joaquin antelope squirrel (Ammospermophilus nelsoni; SJAS) is a state-listed Threatened species in California that evolved in the shrublands of the San Joaquin Desert. Due to livestock overgrazing and fires, much of the desert has lost its shrub cover, which would be a conservation concern if shrubs are a necessary component of the habitat for squirrels. We established two 64-trap plots on the Lokern area of Kern County, California: one covered by saltbush scrub (Atriplex spp.) and another nearby without shrubs. We trapped quarterly starting in January 2019 and ending in November 2021 (12 trap sessions). …

First Documentation Of Ergasilus Centrarchidarum On Smallmouth Bass In Idaho, Danielle Perkins, John W. Heckel Iv, Brett High, Eric J. Billman

Western North American Naturalist

Ergasilus centrarchidarum is a parasitic copepod that infects the gills of fishes in the family Centrarchi­dae. This parasite has been introduced in many regions of North America through introduction of infected Smallmouth Bass (Micropterus dolomieu); however, effects of E. centrarchidarum on its hosts have been poorly studied. Ergasilus centrarchidarum was first discovered infecting Smallmouth Bass in Ririe Reservoir in 2020, although timing of introduction remains uncertain. Our objective was to determine prevalence and intensity of infection of E. centrarchidarum on Smallmouth Bass in Ririe Reservoir. We used angling to collect Smallmouth Bass in the upstream portion of Ririe …

First Shasta Ground Sloth (Nothrotheriops; Xenarthra) From The Eastern Great Basin, Nevada, Jim I. Mead, Sandra L. Swift, H. Gregory Mcdonald, Steven D. Emslie

Western North American Naturalist

We present the first record of the Shasta ground sloth (Nothrotheriops shastensis) for the eastern Great Basin. The single right upper second molariform was recovered from the back dirt pile of archaeological test pits excavated during the 1960s and 1970s in Smith Creek Cave, northern Snake Range, White Pine County, Nevada. The precise age of the specimen is indeterminate, although all sediment layers in the area produced radiocarbon dates older than 11,000 cal BP; thus, the specimen is considered of the Rancholabrean Land Mammal Age. All previous accounts of this sloth in the Great Basin are from its …

Conspecific Tolerance In A Solitary Small Carnivore, The Pacific Marten, Matthew S. Delheimer, Marie E. Martin, Katie M. Moriarty

Western North American Naturalist

Holarctic martens (genus Martes) are small carnivores that are considered functionally solitary, with both resource scarcity and martens’ inability to store energetic reserves posited as precluding social behavior. Interactions between martens are expected to be largely agonistic with the exception of mating and rearing of kits by adult females. Nonetheless, we documented multiple instances of conspecific tolerance by Pacific martens (M. caurina) during long-term monitoring of radio-marked animals in 2 study populations. Observations from radio-tracking and remotely triggered cam­era trap photographs suggested that pairs of martens traveled, foraged, and rested together. Presumed marten associations occurred both inter- …

Nest Success By A Female Black-Chinned Hummingbird (Archilochus Alexandri) With Male-Like Plumage, Cooper M. Farr, John Middleton

Western North American Naturalist

Female-limited polymorphism, a phenomenon in which female animals can exhibit male-like coloration, is known to occur in approximately 25% of hummingbird species, but it has not previously been recognized in Black-chinned Hummingbirds (Archilochus alexandri). Between May and July 2021, we documented a Black-chinned Humming­bird with male plumage features that built a nest, incubated eggs, and successfully fledged 2 chicks in Salt Lake City, Utah. The bird’s gorget feathers, including the chin and upper throat, were predominantly dark with a small number of white feathers interspersed. Because the bird participated in nest building, presumed egg production, and incubation—behaviors not …

Use Of Highway Infrastructure By Bats: A Literature Review And Survey Of The Trans-Pecos Region Of Texas, Holly G. Wilson, Stirling J. Robertson, Richard D. Stevens

Western North American Naturalist

Bats are important components of nature and provide both monetary and nonmonetary benefits to mankind. While anthropogenic effects on wildlife and their habitats are typically viewed as negative, highway infrastructure might benefit numerous species of plants and animals, bats in particular. Cracks, crevices, and grooves within structures provide places for bats to roost. Here we review the literature on this topic as well as describe results of our own survey of bridges and culverts in the Trans-Pecos of Texas. We surveyed 204 bridges and 768 culverts between 2018 and 2019. We observed 7 species of bats using bridges or culverts …

Channel Island Foxes (Urocyon Littoralis) Are Viable Seed Dispersal Agents Of Toyon, Savannah L. Bartel, Juliann T. Schamel, Bridget A. Parrino, Melissa K. Marshall, John L. Orrock

Western North American Naturalist

Mammals that consume fleshy fruit and deposit viable seeds in their scat can be important agents of seed dispersal for plants. Despite the knowledge that the island fox (Urocyon littoralis) consumes large quantities of fleshy fruits, it is unknown whether seeds survive digestion by island foxes and are able to germinate. In order to evaluate whether island foxes are agents of viable seed dispersal on the Channel Islands, we collected island fox scat on Santa Cruz Island in February 2021 and conducted germination assays on digested seeds found in scat. We compared the germinability and germination rates of …

Using Detection Dogs And Rspf Models To Assess Habitat Suitability For Bears In Greater Yellowstone, Jon P. Beckmann, Lisette P. Waits, Aimee Hurt, Alice Whitelaw, Scott Bergen

Western North American Naturalist

In the northern U.S. Rockies, including the Greater Yellowstone Ecosystem (GYE), connectivity is a concern because large carnivores have difficulties dispersing successfully between protected areas. One area of high conservation value because of its importance for connecting the GYE to wilderness areas of central Idaho is the Centennial Mountains and surrounding valleys (2500 km2) along the Idaho–Montana border just west of Yellowstone National Park. The current expansion of grizzly bears (Ursus arctos) and other large carnivore populations outside protected areas of Yellowstone National Park and Grand Teton National Park has placed a greater emphasis on potential linkage zones …

Bird Use Of Riparian Vegetation Along The Truckee River, Califonia And Nevada, Suellen Lynn, Michael L. Morrison, Amy J. Kuenzi, Jennifer C. C. Neale, Benjamin N. Sacks, Robin Hamlin, Linnea S. Hall

Great Basin Naturalist

The Truckee River in California and Nevada is subject to diverse water regimes and a corresponding variety of flow rates. Original riparian vegetation has been altered by these variable flow rates and by a variety of human uses resulting in loss of native riparian vegetation from its historic extent. We conducted bird surveys along the Truckee River during spring 1993 to (1) determine relationships between birds and the present vegetation; (2) determine the importance of different vegetation types to sensitive bird species that have declined recently in the western United States due to competition from exotic plant species, cowbird ( …

Gap Analysis Of The Vegetation Of The Intermountain Semi-Desert Ecoregion, David M. Stoms, Frank W. Davis, Kenneth L. Driese, Kelly M. Cassidy, Michael P. Murray

Great Basin Naturalist

A conservation gap analysis was conducted for the Intermountain Semi-Desert ecoregion to assess the representation of land-cover types within areas managed primarily for biodiversity objectives. Mapped distributions of plant communities were summarized by land-management status categories. The total amount of land permanently protected in the ecoregion is

Winter Macroinvertebrate Communities In Two Montane Wyoming Streams, Christopher M. Pennuto, Frank Denoyelles Jr., Mark A. Conrad, Frank A. Vertucci, Sharon L. Dewey

Great Basin Naturalist

Macroinvertebrate communities were examined on 4 winter dates over a 4-yr period in 2 high-altitude Rocky Mountain streams to document overwintering assemblages potentially experiencing spring acid pulses. Taxa richness values were comparable to other published lists for alpine and montane stream systems despite the fact that most literature reflected summer collections. Mean benthic density ranged from 1406 to 19,734 organisms/m², and drift rates ranged from 0 to 1740 organisms/100 m³. Benthic collections showed higher taxa richness than drift collections while the Ephemeroptera and Plecoptera occurred in greater proportions in drift than in benthos. The Nemouridae (Plecoptera), …

Randomly Amplified Polymorphic Dna Analysis (Rapd) Of Artemisia Subgenus Tridentatae Species And Hybrids, E. Durant Mcarthur, Joann Mudge, Renée Van Buren, W. Ralph Andersen, Stewart C. Sanderson, David G. Babbel

Great Basin Naturalist

Species of Artemisia (subgenus Tridentatae) dominate much of western North America. The genetic variation that allows this broad ecological adaptation is facilitated by hybridization and polyploidization. Three separate studies were performed in this group using randomly amplified polymorphic DNA (RAPD). Fifty-seven 10-mer primers generated nearly 400 markers from genomic DNA obtained from leaf tissue. These studies were (1) a measure of the variability of plants within and between populations and between subspecies using 5 A. tridentata ssp. wyomingensis populations, 2 A. cana ssp. cana populations, and 1 A. cana ssp. viscidula population; (2) an examination of the hypothesis that …

Density, Distribution, And Habitat Of Flammulated Owls In Idaho, Craig Groves, Terry Frederick, Glenn Frederick, Eric Atkinson, Melonie Atkinson, Jay Shepherd, Gregg Servheen

Great Basin Naturalist

From 1990 to 1992 we surveyed for Flammulated Owls (Otus flammeolus) in 3 areas in Idaho: Salmon National Forest (SNF), Payette National Forest and adjacent Hells Canyon National Recreation Area (PNF-HCNRA), and Nez Perce National Forest (NPNF). We also collected and summarized information on all historic and modern records of Flammulated Owls in Idaho. Flammulated Owls were detected on 65% of 68 routes (2–16 km in length) surveyed at densities ranging from 0.04 to 1.25 singing males/40 ha. Owls were detected on survey routes as early as 10 May and as late as 23 July. Mean percent canopy …

Erratum

Great Basin Naturalist

No abstract provided.

Diplostomiasis In Native And Introduced Fishes From Yellowstone Lake, Wyoming, Victor H. Inchausty, Michael Foutz, Richard A. Heckmann, Claudete Ruas, Paulo Ruas

Great Basin Naturalist

Totals of 101 native Yellowstone cutthroat (Oncorhynchus clarki bouvieri), 27 introduced lake trout (Salvelinus namaycush), and 40 introduced longnose sucker (Catostomus catostomus) from Yellowstone Lake, Wyoming, USA, were examined for eye flukes. Metacercariae of the trematode fluke Diplostomum were in vitreous humor and/or lens of 94% of Yellowstone cutthroat trout, 92% of lake trout, and 78% of longnose sucker. Longnose sucker had 7% prevalence of infection in both lens and vitreous humor of metacercariae, while Yellowstone cutthroat trout had 3% and lake trout 8%. Diplostomum spathaceum was in lens tissue of 5% of infected …

Natural Variability Of Vegetation, Soils, And Physiography In The Bristlecone Pine Forest Of The Rocky Mountains, Brigitte M. Ranne, William L. Baker, Tom Andrews, Michael G. Ryan

Great Basin Naturalist

Pinus aristata Engelm. forest vegetation in Colorado was studied to determine vegetation composition and the relationship between vegetation and environment. Species percent cover, tree size class, and environmental variables were recorded for 49 plots. Previously collected data for 4 plots from New Mexico were included. Environmental variables included latitude, longitude, elevation, slope, aspect, topographic position, parent material, percent rock cover, mean rock size, litter depth, estimated plot age class, and evidence of anthropogenic disturbance. Soils were analyzed for texture, depth, and percent carbon and nitrogen. Pinus aristata foliage was analyzed for percent nitrogen and phosphorus. Direct and indirect gradient analyses …

Helminths Of The Southwestern Toad, Bufo Microscaphus, Woodhouse's Toad, Bufo Woodhousii (Bufonidae), And Their Hybrids From Central Arizona, Stephen R. Goldberg, Charles R. Bursey, Keith B. Malmos, Brian K. Sullivan, Hay Cheam

Great Basin Naturalist

The gastrointestinal tracts, lungs, and urinary bladders from 77 Bufo microscaphus, 61 Bufo woodhousii, and 8 of their hybrids were examined for helminths. One species of trematode (Glypthelmins quieta), 1 species of cestode (Distoichometra bufonis), and 5 species of nematodes (Aplectana incerta, A. itzocanensis, Rhabdias americanus, Physaloptera sp., and Physocephalus sp.) were found. The greatest prevalence (41%) and mean intensity (231.7) were recorded for Aplectana incerta in Bufo woodhousii. It appears hybrids harbor fewer parasites than either parent species.

Oochoristica Scelopori (Cestoda: Linstowiidae) In A Grassland Population Of The Bunch Grass Lizard, Sceloporus Scalaris (Phrynosomatidae), From Arizona, Stephen R. Goldberg, Charles R. Bursey, Chris T. Mcallister, Hobart M. Smith, Quynh A. Truong

Great Basin Naturalist

No abstract provided.

Effects Of Douglas-Fir Foliage Age Class On Western Spruce Budworm Oviposition Choice And Larval Performance, Kimberly A. Dodds, Karen M. Clancy, Kathryn J. Leyva, David Greenberg, Peter W. Price

Great Basin Naturalist

The western spruce budworm (Choristoneura occidentalis Freeman) prefers to feed on flushing buds and current-year needles of Douglas-fir (Pseudotsuga menziesii [Mirb.] Franco). Budworm larvae will not typically consume older age classes of needles unless all current-year foliage is depleted. We tested the following null hypotheses: (1) budworm larvae can feed on foliage with a wide range of qualities (i.e., current 1-, 2-, or 3-year-old needles) without measurable effects on fitness; and (2) budworm adults do not show any oviposition preference linked to the age of the foliage they fed on as larvae. We used both laboratory and field …

Selecting Wilderness Areas To Conserve Utah's Biological Diversity, Diane W. Davidson, William D. Newmark, Jack W. Sites Jr., Dennis K. Shiozawa, Eric A. Rickart, Kimball T. Harper, Robert B. Keiter

Great Basin Naturalist

Congress is currently evaluating the wilderness status of Bureau of Land Management (BLM) public lands in Utah. Wilderness areas play many important roles, and one critical role is the conservation of biological diversity. We propose that objectives for conserving biodiversity on BLM lands in Utah be to (1) ensure the long-term population viability of native animal and plant species, (2) maintain the critical ecological and evolutionary processes upon which these species depend, and (3) preserve the full range of communities, successional stages, and environmental gradients. To achieve these objectives, wilderness areas should be selected so as to protect large, contiguous …

Snakes Of Utah, Douglas C. Cox, Wilmer W. Tanner

Books by Faculty of the Monte L. Bean Life Science Museum

Snakes of Utah is a complete, up-to-date, authoritative description of the snakes that occur in the state of Utah. It includes 33 species in 4 families: Leptotyphlopidae, Erycidae, Colubridae, and Crotalidae. Each description includes field marks, photographs, habitats, and distribution maps.

Table of Contents

Introduction

Classification

Leptotyphlopidae

Erycidae

Colubridae

Crotalidae

Glossary

Index

Acknowledgments

On The Relative Importance Of Floral Color, Shape, And Nectar Rewards In Attracting Pollinators To Mimulus, Steven D. Sutherland, Robert K. Vickery Jr.

Great Basin Naturalist

Pollinator preferences were observed for the six species of section Erythranthe of the genus Mimulus using greenhouse-grown plants placed in a meadow in the Red Butte Canyon Natural Area, Salt Lake County, Utah. The principal pollinators were hummingbirds and bumble bees. Hummingbirds preferred the species with the most reflexed tubular flowers regardless of color, whereas bumblebees preferred pink. lavender, or yellow flowers to red flowers regardless of shape. Results for the six species were confirmed by observations of F₂ hybrid recombinant plants selected such that flower color could be held constant and flower shape varied and vice versa.

Red Butte Canyon Research Natural Area: History, Flora, Geology, Climate, And Ecology, James R. Ehleringer, Lois A. Arnow, Ted Arnow, Irving B. Mcnulty, Norman C. Negus

Great Basin Naturalist

Red Butte Canyon is a protected, near pristine canyon entering Salt Lake Valley, Utah. It contains a well-developed riparian zone and a perennial stream; hillside vegetation ranges from grasslands on the lower limits to Douglas-fir and aspen stands at the upper elevations. In this paper we describe the history of human impact, natural history aspects of climate, geology, and ecology, and faunal and floral information for key species in the canyon. The role and importance of Research Natural Areas is discussed, particularly with respect to the need to protect Red Butte Canyon—one of the few remaining undisturbed riparian ecosystems in …