Zoology Commons^™

Scorpio Basraensis Sp. N. From Southern Iraq (Scorpiones: Scorpionidae), Ersen A. Yağmur, Jafar A. Makki, Shurooq A. Najim

Euscorpius

A new scorpion species, Scorpio basraensis sp. n., is described and illustrated from Basra Province, southern Iraq. Scorpio basraensis sp. n. is compared with all known species of the genus Scorpio Linnaeus, 1758 occurring in countries surrounding Iraq. The new species can be distinguished by the circular genital operculum in males and the cordiform genital operculum in females (not posteriorly elongated), a densely hirsute body, appendages and metasoma, a distinctly elongated telson vesicle, discrete granules on the external surface of the chela manus that are not flattened, and pectinal tooth counts of 11–13 in males and 9–11 in females. …

Historical Chaos Among Tetrabothriideans (Eucestoda) Of Odontocetes (Artiodactyla)—Revising Generic Taxonomy And Species Diversity For Cestodes Among Oceanic Dolphins, Eric P. Hoberg, Kaylen Marie Soudachanh, Robert L. Rausch

MANTER: Journal of Parasite Biodiversity

Explorations over the past 150 years that have defined the diversity and structure of the tetrabothriidean cestode fauna among Odontocete cetaceans (Artiodactyla) are summarized and reviewed. Persistent errors in interpretation of comparative morphological characters, inconsistent taxonomic decisions, and an understanding of generic-level and species diversity are addressed. We outline a revision of the Tetrabothriidea, which now encompasses 8 genera. Prosthecocotyle is resurrected from synonymy with Tetrabothrius, and Cetaceobothridius n. gen is proposed. A clear basis is outlined for differentiation of generic and species diversity among Cetaceobothridius, Priapocephalus, Prosthecocotyle, Strobilocephalus, Tetrabothrius, and Trigonocotyle that constitute the tetrabothriidean fauna of odontocetes; species …

A New Species Of Liocheles Sundevall, 1833 (Scorpiones: Hormuridae) From The Western Himalayas, Virender К. Bhardwaj, Sunil Katal, H. T. Lalremsanga, Zeeshan A. Mirza

Euscorpius

Sampling in the Western Himalayas of India led to the discovery of a population of the genus Liocheles Sundevall, 1833, that differes from all Indian congeners. Molecular and morphological data confirmed the distinctiveness of the population, which is here described as a new species. The new species is morphologically similar to L. longimanus (Werner, 1939), from which it differs in the arrangement of trichobothria on external patella and structure of the hemispermatophore. A preliminary molecular phylogeny of the family Hormuridae based on the mitochondrial cytochrome oxidase I gene is presented.

New Locality Records And Future Habitat Suitability Of Sassanidotus Gracilis (Birula, 1900) (Scorpiones: Buthidae) In The Iran–Afghanistan–Pakistan Arid Belt, Ersen A. Yağmur, Kerim Çiçek, Homa Barati, Mohammad Moradi

Euscorpius

This study reports new locality records from Iran and a climate-based habitat suitability assessment for Sassanidotus gracilis (Birula, 1900) (Scorpiones: Buthidae) in the Iran–Afghanistan–Pakistan arid belt. We document new specimens from Sistan and Baluchestan Province, Iran, with an updated distribution map and illustrations, and we model the species' potential distribution across south-eastern Iran and adjacent parts of Afghanistan and Pakistan.

Habitat suitability was associated mainly with precipitation seasonality, precipitation of the wettest month, precipitation of the driest month, and slope. In an arid setting these rainfall variables are unlikely to indicate a need for moist habitat; they more plausibly track …

First Insular Record And Dna Barcoding Of Euscorpius Vailatii Tropea & Fet, 2015 From Trizonia Island (Fokida, Greece), Konstantinos Kalaentzis, Francesco Frigioni, Alexios Panagiotopoulos, Elias Tzoras, Styliani Minoudi, Alexandros Triantafyllidis

Euscorpius

Euscorpius vailatii Tropea & Fet, 2015 (Scorpiones: Euscorpiidae), originally described from Aitoloakarnania and Fokida (Greece), is herein reported from Trizonia Island in the Gulf of Corinth, representing the first insular record for the species. The examined specimens are consistent with the diagnostic morphology of E. vailatii. The first molecular data for the species are provided, based on both mainland and insular populations. Comparative analyses further highlight the distinctiveness of E. vailatii from geographically close congeners, while phylogenetic results indicate a sister-taxon relationship with E. erymanthius, supporting a Quaternary biogeographic history shaped by the evolution of the Gulf of …

Snails In The Diet Of The Cutlip Minnow, Exoglossum Maxillingua, Tess A. Taloumis, Steven L. Powers

Southeastern Fishes Council Proceedings

The Cutlip Minnow, Exoglossum maxillingua, is an important stream fish native to the eastern United States. It has an unusual lower jaw morphology. Early hypotheses suggested the jaw morphology is an adaptation for feeding on snails. Despite the publication of numerous diet studies on this species, there is no documentation of snails as prey items. During recent snorkel observations in the Roanoke River, E. maxillingua was spotted carrying snails within their mouths, prompting further investigation. An in-depth temporal examination of the diet from 145 specimens of E. maxillingua across multiple sites was conducted to further investigate the diet of …

Formalization Of Nomenclatural Acts Proposed In Tang (2026) (Scorpiones: Buthidae, Chaerilidae, Scorpiopidae), Victoria Tang

Euscorpius

Several nomenclatural, taxonomic, and faunistic decisions put forth in Tang (2026) are formalized herein: (1) four nomina dubia, Chaerilus tessellatus Qi et al., 2005, Scorpiops langxian Qi et al., 2005, S. pococki Qi et al., 2005, and Isometrus (Reddyanus) wenjieae Huang et al., 2025; (2) two junior subjective synonyms, Scorpiops langxian = S. pococki = S. tibetanus Hirst, 1911 syn. n.; (3) one new “aspidonym”, Scorpiops obluda Tang, 2026 sp. indet. nomen nudum; and (4) removal of Scorpiops hardwickii (Gervais, 1843) and S. petersii Pocock, 1893 from the Chinese scorpiofauna. Reddyanus tibetanus (Lourenço & Zhu, 2008) is maintained …

Scorpiops Walteri Sp. N. (Scorpiones: Scorpiopidae) From Myanmar, František Kovařík

Euscorpius

A new species Scorpiops walteri sp. n. (Scorpiopidae) is described from Myanmar, fully illustrated with color photos showing its morphology and habitus. Scorpiops walteri sp. n. is the only species of the genus with a combination of the following five characters: chelal trichobothrium Eb₃ located in middle of manus between trichobothria Dt and Est; patella of pedipalp with 11–12 ventral and 19 external trichobothria; fingers of pedipalps strongly undulate in male; pedipalp movable finger with ca. 47 IAD; and chela length to width ratio 2.8 in male.

Behavioral Analysis Of Cryptoprocta Ferox At The Central Florida Zoo And Botanical Gardens, Rita R. Albert

The Pegasus Review: UCF Undergraduate Research Journal

The fossa, Cryptoprocta ferox, is the largest carnivorous mammal endemic to the island of Madagascar. C. ferox is currently listed as a vulnerable species by the IUCN Red List (2015) due to deforestation. Most fossas are now located in zoos for breeding and repopulation purposes, but research on captive mating has been limited due to the low numbers of successful breeding attempts in captivity. The Central Florida Zoo and Botanical Gardens is home to two fossas, one male and one newly introduced female. This study notes behavioral changes in captive mating fossas, with a focus on the male fossa …

Euscorpius Emae Sp. N. (Scorpiones: Euscorpiidae), A New Scorpion Species From Montenegro, František Kovařík

Euscorpius

A new species Euscorpius emae sp. n. (Scorpiones: Euscorpiidae) from Montenegro is described, fully complemented with color photographs. E. emae sp. n. is morphologically unique by yellow to orange color, very narrow metasoma and enlarged chela. Pedipalp patella external trichobothria numbers are 4eb, 4eb_a, 2esb, 4em, 4est, 5et, and ventral aspect of patella with 9 trichobothria.

Tails From The Isle Of Devils: Two New Records Of Non-Native Invasive Scorpions (Arachnida: Scorpiones) Successfully Intercepted In Bermuda, Including The First Record Of Caraboctonidae, Danniella Sherwood, Mark Outerbridge, Struan R. Smith, Jenny Gosling, Myles Darrell

Euscorpius

Hadruroides sp. (Scorpiones: Caraboctonidae) is newly recorded as a successfully intercepted non-native invasive species in Bermuda, based on examination of a desiccated and damaged immature female specimen. A second record of an intercepted Centruroides gracilis (Latreille, 1804) (Scorpiones: Buthidae) is reported based on examination of an adult male.

Metabolic Markers And Gut Microbiota In Big Cats: A Relationship Between Diet, Health And Environment, Javeria Saman, Saima Sharif, Bushra Nisar, Shagufta Naz

Journal of Bioresource Management

The wild animals that occupy large territories were found to adapt less easily to captive environments compared to those with small home ranges. Among big cats, captivity posed a range of challenges, with stress emerging as central factor influencing overall welfare. Transition into captive environment was shown to alter both behavior and physiological processes, largely due to changes in their natural habitat, environmental complexity and diet. Several health issues reported in big cats including alopecia, gastrointestinal issues, neuromuscular disorders such as hindquarter paralysis were associated with underlying metabolic dysregulation. Metabolic health was recognized as critical component of overall well-being, closely …

Scorpions Of Angola (Arachnida, Scorpiones). Part I. Family Buthidae, With Descriptions Of Two New Species., František Kovařík

Euscorpius

All scorpion species of the family Buthidae known from Angola are listed, with color photographs and maps of their distribution. Diagnosis of Babycurus ansorgei Hirst, 1911 is revised; its male is described for the first time. Babycurus crassicaudatus Roewer, 1952, is revalidated (it was erroneously synonymized with B. ansorgei, since the females of both species are morphologically almost identical). Uroplectes angolensis sp. n. and U. xavieri sp. n. are described from Angola. Validity of Uroplectes ngangelarum Monard, 1930 is confirmed as different from U. planimanus (Karsch, 1879); its lectotype is designated. Lectotype is also designated for Uroplectes planimanus …

History And Distribution Of Great-Tailed Grackles (Quiscalus Mexicanus) In Arkansas, R. Tumlison, V. Andersen

Journal of the Arkansas Academy of Science

Great-tailed Grackles (Quiscalus mexicanus) were reported first in southwestern Arkansas in 1969, were nesting there in 1977, and established a resident population by 1980. Nesting was reported in far northwestern Arkansas in 1986. Presently, the species is known from 44 of Arkansas’s 75 counties, those mostly in the western and central counties. Most individuals appear to be migratory, with most reports occurring in April and May. However, breeding and now resident populations have been reported in the southwestern, northwestern, and central counties, particularly in urban areas. Birds are very commonly reported to forage at fast-food establishments and truck …

Morphometric And Genetic Analyses Of The Sheat Catfish Genus Ompok And Kryptopterus (Siluriidae) From The Mahakam River, East Kalimantan, Indone-Sia, Jusmaldi Jusmaldi, Rudhy Gustiano, Haryono Haryono, Rusdianto Rusdianto, Gema Wahyudewantoro, Syahroma Husni Nasution, Yulianto Yulianto, Muhammad Hunaina Fariduddin Aththar, Safar Dody, Amran Ronny Syam, Irin Iriana Kusmini, Yosmaniar Yosmaniar

Makara Journal of Science

Anthropogenic impacts within the watershed threaten the survival of sheat catfish. Therefore, species of the genera Ompok and Kryptopterus should be conserved and domesticated to ensure their long-term persistence. Moreover, the accurate documentation of the existence and taxonomic validity of Ompok and Kryptopterus species is crucial. Accordingly, this study aims to assess the species diversity of the genera Ompok and Kryptopterus in the Mahakam River, East Kalimantan, Indonesia. The study was conducted between June and December 2014, with an update in August 2024. Fish samples were collected from the upstream and middle reaches of the Mahakam River. A total of …

Effect Of Temperature On The Defensive And Predatory Behavior Of The Scorpion Hadrurus Obscurus (Scorpiones: Hadruridae), Zia Nisani, Megan Bennett, Audrey Enriquez, Mel Iniquez, Emilia Martinez Rios, Brooklyn Stone

Euscorpius

Scorpions employ their stinging mechanism both for defense and predation, and being ectothermic, temperature influences their physiology and behavior. As such, we hypothesized that temperature both affects defensive stinging and prey-capture behavior in scorpions. We tested the effects of body temperature on these behaviors in the desert hairy scorpion, Hadrurus obscurus Williams, 1970. In the first experiment, scorpions were placed in temperature-controlled chambers with either high (26-28°C) or low (21- 22°C) temperatures and probed to elicit a defensive sting. We recorded the reaction time and the number of probes needed to elicit a sting. In the second experiment, we observed …

Compsobuthus Hadeki Sp. N. From Iraq (Scorpiones: Buthidae), František Kovařík, František Šťáhlavský

Euscorpius

A new species, Compsobuthus hadeki sp. n. from Iraq is described and fully complemented with color photographs and compared with morphologically the most similar species, C. matthiesseni (Birula, 1905). The two species can be unequivocally separated by the following morphological features: in C. hadeki sp. n., movable finger of pedipalp bears 9 rows of granules; telson with aculeus shorter than vesicle, and metasoma III with 10 carinae. Cytogenetic analysis was performed, revealing a diploid chromosome count of 22 in C. hadeki sp. n.

A Reassessment Of “Tityus” Tenuimanus Banks, 1910, As A Misidentified Leiurus Sp. (Scorpiones: Buthidae), Mohamed Mousaid, Ersen A. Yağmur, Zoe Flores, Alana Rivera, Aziza Lansari, Abdellah Bouazza

Euscorpius

This study re-examines the holotype of “Tityus” tenuimanus Banks, 1910, previously synonymized with Buthus atlantis Pocock, 1889 by Lourenço & Francke (1984). Our results indicate that the specimen is a misidentified representative of Leiurus Ehrenberg, 1828, as it exhibits the diagnostic characters of this genus: the central lateral and posterior median carinae of the carapace fused into a lyre-shaped configuration, and tergites I–II bearing five granular carinae (a character absent in the genus Buthus Leach, 1815). Moreover, the holotype lacks one of the most distinctive characters of the genus Tityus Koch, 1836, namely the presence of a subaculear tubercle. The …

Scorpions In Brazilian Caves And Other Subterranean Habitats, Jonas E. Gallão, Bruno D, Lenhare, Misael Oliveira-Neto, Maria E. Bichuette

Euscorpius

Arachnids have successfully colonized subterranean environments. Compared to spiders, harvestmen, and pseudoscorpions, scorpions have a moderate number of subterranean species, including troglophiles and troglobites. Among the four families and 184 species and three subspecies of scorpions found in Brazil, three families and at least 33 species and one subspecies have been documented in subterranean habitats. This study presents a faunistic list of scorpion occurrences in caves and other types of subterranean habitats in Brazil, along with distribution maps and an attempt to classify species based on their use of and dependence on subterranean environments.

The Genus Nebo Simon, 1878 (Scorpiones: Diplocentridae) In Oman, Graeme Lowe

Euscorpius

No abstract provided.

Mammalian Biodiversity Of The Easley Conservation Stewardship Area, Grayson C. Belt

Honors College Theses

Monitoring wildlife is an important part of conservation and is particularly vital for newly acquired conservation areas to provide a baseline of what is present. In 2025, the 22 ha Murray State University Easley Conservation Stewardship Area (MSU ECSA) was established following a donation to the university by Melissa Easley. I conducted a survey using trail cameras at the ECSA in fall 2025 and winter 2025/26 to record mammalian diversity. The survey consisted of two subprojects: a mesomammal subproject of 18 knee height trail cams and a small mammal subproject using 15 custom small mammal bucket cameras. I uploaded all …

The Hitchhiker's Guide To The Host: Fleas And The History Of Pikas In North America, Josh Pletcher

All NMU Master's Theses

Parasite phylogenetics can offer windows into the cryptic evolutionary history of hosts, often highlighting events that are not recorded by the phylogenetics of the host. Pikas (small lagomorphs) host a diverse parasite community comprising endoparasites (e.g., coccidians, nematodes, and cestodes) and ectoparasites (e.g., fleas and mites). Phylogenetic analysis of pika endoparasites has revealed pikas evolved in Asia and colonized North America in two waves, and that contact between these two pika lineages allowed parasites to be transferred from one lineage to another and to survive the extinction of their original host. To uncover these cryptic histories, understanding the biogeography of …

An Integrative Description Of Euscorpius Diagorasi Sp. N. From Rhodes, Greece (Scorpiones: Euscorpiidae), Konstantinos Kalaentzis, Francesco Frigioni, Eliza Kaitetzidou, Alessio Iannucci, Alexandros Triantafyllidis

Euscorpius

The genus Euscorpius Thorell, 1876 comprises a diverse and taxonomically challenging group of scorpions in the Mediterranean, with Greece representing one of its principal centers of diversity. In this study, we provide an integrative description of Euscorpius diagorasi sp. n., a new species from Rhodes Island, Greece. The new species is described on the basis of adult male and female morphology and mitochondrial COI sequence data. It is a small oligotrichous species characterized by a total length of approximately 21–25 mm, pale yellow to light brown coloration with darker reddish-brown pedipalps, pectinal tooth count of 8 in the male …

Fish Community Assessment Of Goose Pond Natural Area, Macy Sparks

ATU Scholars Symposium

Natural areas play an integral role in the preservation and conservation of unique ecosystems (i.e., natural wetlands). However, many natural areas have never had a biological inventory of species utilizing these areas. Goose Pond Natural Area (Conway County and Pope County, Arkansas) is one of those natural areas lacking species occurrence data. We addressed this data gap by sampling fish species across multiple habitat types (shallow woody wetland, open grassy wetland, and stream). We employed multiple gear types (mesh umbrella minnow traps, seines, gill nets, mini-fyke nets, and backpack electrofishing) in hopes of increasing species detection. Sites within the natural …

A Revision Of The Psammophilic Genus Liobuthus Birula, 1898 (Scorpiones: Buthidae), With A Description Of Four New Species From Central Asia, Victor Fet, František Kovařík, Graeme Lowe, Matthew R. Graham

Euscorpius

The rare Central Asian psammophilic scorpion genus, Liobuthus Birula, 1898 (Scorpiones: Buthidae), is revised. Five species are delineated according to results from three species delimitation approaches using mitochondrial DNA. Results are supported by diagnostic morphology, including an assessment of the distinctive neobothriotaxy. The type species, L. kessleri Birula, 1898, is confirmed for Iran (northeast), Turkmenistan and Uzbekistan (south). Four new species are described: L. atamuradovi sp. n. (Turkmenistan),  L. danovi sp. n. (Kazakhstan, northern Turkmenistan, Uzbekistan), L. shukurovi sp. n. (southwestern Tajikistan), and L. vachoni sp. n. (southern Kazakhstan), The unique additive neobothriotaxy (on both pedipalp femur and patella) …

Western Massasauga (Sistrurus Tergeminus): A Species Conservation Assessment For The Nebraska Natural Legacy Project, Colleen Rothe-Groleau, Melissa J. Panella

Nebraska Game and Parks Commission: Publications

The Nebraska Natural Legacy Project recognizes the western massasauga (Sistrurus tergeminus) as a Tier I at-risk species. Provided are some general management recommendations regarding western massasaugas. Conservation practitioners will need to use professional judgment to make specific management decisions based on objectives, location, and a multitude of variables. This resource was designed to share available knowledge of this at-risk species that will aid in the decision-making process or in identifying research needs to benefit the species.

Criteria for selection as Tier I: State listed, G3

Estimated Population in NE: 1,000–2,500

Estimate based on: Field surveys

Trends since 2005 …

Systematics Of Disk-Winged Bats (Thyroptera), Courtney L. Klumpp

Capstone Showcase

This thesis investigates the evolutionary relationships among disk-winged bats in the genus Thyroptera, a small group of Neotropical bats known for the adhesive suction disks on their wings and feet that allow them to roost inside tightly rolled leaves. The distinctive morphology and behavior of these bats have not been extensively documented, and their phylogenetic relationships remain only partially resolved. The goal of this research is to better understand how these species are related to one another and to determine whether the five recognized species of Thyroptera form a single monophyletic group. To address this question, morphological and genetic …

Indirect Effects Of Coral Bleaching On Tursiops Truncatus Diet And Foraging Behavior In Bocas Del Toro, Panama, Annabel Fiero

Scripps Senior Theses

Indirect effects, where another species or factor mediates the effects of a given species, are critical to ecosystem structure and resilience. Shifts in fish community structure, as a result of rapidly declining coral reefs in the Caribbean due to warming-induced bleaching, pose a threat to apex predators like Tursiops truncatus (bottlenose dolphins). Branching corals create habitat niches for numerous fish species. These fish rely on coral for food and habitat, and in turn provide food for upper trophic levels. Prior research has highlighted how coral bleaching will disrupt Acroporid reef building corals and fish community structure, but the effects of …

The Effect Of Noise Pollution On The Diversity And Abundance Of Bird Species At Different Locations At Warner Parks, Nashville, Tn, Jeremy T. Little, Wyatt L. Fooks, Darlene Panvini, Jackson Whiteside

Science University Research Symposium (SURS)

Noise pollution is a significant abiotic factor that can alter bird biodiversity and abundance of birds in urbanized areas. A new road is proposed through a relatively untouched area of Warner Parks in Nashville, TN, which may affect the bird community. Acoustic Song Meters were used to monitor bird sounds and soundscapes in the proposed road site and two areas of the park with different levels of noise pollution (roadside=high noise; isolated=low noise) for three weeks. Recordings were analyzed to determine bird species and abundance at each site to predict how increasing noise pollution may affect bird diversity in the …

First Report Of A Bat Guano Fire From A Cave In Southeast Asia, Sopark Jantarit, Martin Ellis, Pruet Kalasuwan, Neeranuch Phusunti, Pimonsri Mittraparp-Arthorn, Jiranan Pattano, Arnon Chukamnerd, Turdtun Tong-In, Natthawut Khrueaaunruean

International Journal of Speleology

Fires occurring in cave guano are a rare and unusual phenomenon and the occurrence of such events in Southeast Asia is particularly novel. In this study, we document the first recorded instance of a bat guano fire in a tropical cave at Tham Chao Ram in Sukhothai province, northern Thailand. A bacteriological analysis was made of the guano at the fire site and from a control site within the cave. We give a comprehensive review of global reports on bat guano fires in caves, conduct an initial investigation of the physical factors at the fire site and discuss the potential …