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Articles 511 - 540 of 2030

Full-Text Articles in Other Animal Sciences

Goats Distinguish Between Positive And Negative Emotion-Linked Vocalisations, Luigi Baciadonna, Elodie Briefer, Livio Favaro, A. G. Mcelligott Jul 2019

Goats Distinguish Between Positive And Negative Emotion-Linked Vocalisations, Luigi Baciadonna, Elodie Briefer, Livio Favaro, A. G. Mcelligott

Bioacoustics Collection

Background: Evidence from humans suggests that the expression of emotions can regulate social interactions and promote coordination within a group. Despite its evolutionary importance, social communication of emotions in non-human animals is still not well understood. Here, we combine behavioural and physiological measures, to determine if animals can distinguish between vocalisations linked to different emotional valences (positive and negative). Using a playback paradigm, goats were habituated to listen to a conspecific call associated with positive or negative valence (habituation phase) and were subsequently exposed to a variant of the same call type (contact call) associated with the opposite valence (dishabituation …


Sex-Specific Personalities In The Purple Marsh Crab, Jillian Sterman, Jessica Barton, Panagiota Delmedico, Samantha Sweeney Jun 2019

Sex-Specific Personalities In The Purple Marsh Crab, Jillian Sterman, Jessica Barton, Panagiota Delmedico, Samantha Sweeney

DePaul Discoveries

Animals are considered to possess personalities when individuals differ in behavior, and these differences are consistent between situations. Several studies have identified personalities in diverse groups but less is known about personality variation between the sexes. In this study, we examined variation in two key personality traits (boldness, activity) in female and male purple marsh crabs (Sesarma reticulatum) using a semi-field approach. Specifically, we measured boldness and activity on two consecutive days using the same behavioral assays during each time point. Consistency (personality) was determined using Kendall’s coefficient of concordance based on Spearman correlation coefficients for each behavior. …


Wyoming Wildlife: A Natural History, Paul Johnsgard, Thomas D. Mangelsen Jun 2019

Wyoming Wildlife: A Natural History, Paul Johnsgard, Thomas D. Mangelsen

Zea E-Books Collection

This book surveys Wyoming’s mammal, bird, reptile, and amphibian faunas. In addition to introducing the state’s geography, geology, climate, and major ecosystems, it provides 65 biological profiles of 72 mammal species, 195 profiles of 196 birds, 9 profiles of 12 reptiles, and 6 profiles of 9 amphibians. There are also species lists of Wyoming’s 117 mammals, 445 birds, 22 reptiles, and 12 amphibians. Also included are descriptions of nearly 50 national and state properties, including parks, forests, preserves, and other public-access natural areas in Wyoming. The book includes a text of more than 150,000 words, nearly 700 references, a glossary …


The Horse's Back As Influenced By Saddle Fit, Katie Millican Jun 2019

The Horse's Back As Influenced By Saddle Fit, Katie Millican

Agricultural Education and Communication

Therefore, this project will use devices called the Lameness Locator and a thermal camera along with facilities at the Equine Center in San Luis Obispo, California. The objective of the project is to utilize a population of sound horses and compare the heat detection images of their backs and saddles after being ridden to provide saddle fit data. This project will provide information contributing to closing the gap of equestrians uneducated about the importance of saddle fit.

The reason for researching and conducting a study to prove this point is simple: a horse owner needs to look at the bigger …


Sensory Perception, Adrian Rodriguez-Contreras May 2019

Sensory Perception, Adrian Rodriguez-Contreras

Open Educational Resources

Different types of sensory systems with their functional modalities will be presented. The biological bases for how these functions are generated and modified will then be described. As vision is the principal means of perception, we will focus in this course most on visual processing. Scientific data will be integrated into the lectures, such that students develop critical skills in analyzing data and proposing hypotheses.


Oxyrase And Pyruvate Inclusion In Extenders For Cold Stored Stallion Spermatozoa, Jordan T. Shore May 2019

Oxyrase And Pyruvate Inclusion In Extenders For Cold Stored Stallion Spermatozoa, Jordan T. Shore

Graduate Theses/Dissertations

A current method in addressing subfertility in stallions include manipulating the semen extender media by supplementing antioxidants and energy sources. Two experiments (EXP 1/EXP 2) were conducted to assess the effects of pyruvate and Oxyrase® suspended in commercial diluents for preservation of stallion spermatozoa. Assessment of total (TM) and progressive motility (PM), velocity and direction of movement (VAP, VSL, VCL, and elongation) were recorded by CASA. In EXP 1, 3 different ejaculates were collected from each of 4 stallions. Aliquots of each ejaculate were suspended into 4 treatments of INRA96 with or without Oxyrase® supplemented at 2.4 U/ml …


Considerations For Nisc’S Future Without Isac, Invasive Species Advisory Committee May 2019

Considerations For Nisc’S Future Without Isac, Invasive Species Advisory Committee

National Invasive Species Council

The Invasive Species Advisory Committee (ISAC) was established in 1999 by Executive Order 131121 and furthered by Executive Order 137512 to provide consultation, feedback, information on particular issues, and expert advice generally for consideration by the National Invasive Species Council (NISC). ISAC is composed of distinguished, experienced volunteers from various organizations that are appointed to serve on the Committee by the Secretary of the Interior. Their task is to develop assessments on many aspects of the invasive species issue in the United States and to provide recommendations for the management, control, and response to key invasive species as articulated in …


American White Pelicans, Tommy King May 2019

American White Pelicans, Tommy King

Wildlife Damage Management Technical Series

American white pelicans (Pelecanus erythrorhynchos, Figure 1) threaten aquaculture producers by direct predation and the spread of disease. They are also considered competition and a nuisance by some sports fishermen. Pelicans can also damage pond levees and crops, such as rice, by trampling the vegetation and depositing guano. A combination of wildlife damage management techniques is often necessary to reduce pelican damage to these resources. Aquaculture Prior to the winter of 1992, American white pelican depredations at catfish facilities in the Delta regions of Arkansas and Mississippi were limited, and birds were easily dispersed from the area. Since 1992, however, …


The Effects Of Pre And Post Exercise Low-Level Laser Therapy On Biochemical Markers Of Skeletal Muscle Fatigue In Equines, Carson Freeman May 2019

The Effects Of Pre And Post Exercise Low-Level Laser Therapy On Biochemical Markers Of Skeletal Muscle Fatigue In Equines, Carson Freeman

Animal Science Undergraduate Honors Theses

Our objective was to determine whether administering low-level laser therapy before or after exercise had the greatest effect on biochemical markers of skeletal muscle fatigue in equines such as cortisol and blood lactate. Twelve quarter horses were divided into three groups: Group A received no laser therapy, Group B received laser therapy before exercise, and Group C received laser therapy after exercise. A Class II ERCHONIA ® PL500 handheld low-level laser was utilized for treatment with a wavelength of 635nm. Exercise was utilized using a horse walker system for 30 minutes five days a week for three weeks. Blood was …


Bottlenose Dolphin (Tursiops Truncatus) Vocal Responses To Sonar And Spectrally Pink Background Noise, Maria Zapetis May 2019

Bottlenose Dolphin (Tursiops Truncatus) Vocal Responses To Sonar And Spectrally Pink Background Noise, Maria Zapetis

Dissertations

As human populations rise, the level of man-made noise increases globally. Naval sonar and boat traffic are underwater sound sources of particular concern to marine mammal welfare. To better understand the impact of these noise increases on cetaceans, studies can explore animals’ behavioral changes in response to noise. Studies have investigated the ‘dose-response’ relationship between the received sound pressure level of sonar signals and the behavior of cetaceans in the wild, but exposure studies in controlled environments are limited. The studies in this dissertation examined bottlenose dolphin vocal modifications during various experimental noise treatments. Acoustic recordings previously obtained for bottlenose …


The Diversity Of Terrestrial Mammals Surrounding Waterfall At Billy Barquedier National Park, Kelsey Johnson May 2019

The Diversity Of Terrestrial Mammals Surrounding Waterfall At Billy Barquedier National Park, Kelsey Johnson

Animal Science Undergraduate Honors Theses

Billy Barquedier is a National Park located in the Stann Creek district of Belize that contains Neotropical vegetation and wildlife. This study was performed in order to provide a baseline inventory and appearance frequency patterns of the terrestrial mammals located within Zone 1 of the park near a waterfall. The goal of the data and information collected during this study is to gain a greater understanding of the biodiversity and activity patterns of terrestrial mammals within the park and across Belize as a whole. The use of camera traps, small Sherman live traps, large live traps, and tracking methods were …


Initiation Of A Roundtable Meeting To Determine Safety Hazards And Provide Education To Range Bison Herd Workers, Lucia Finocchiaro May 2019

Initiation Of A Roundtable Meeting To Determine Safety Hazards And Provide Education To Range Bison Herd Workers, Lucia Finocchiaro

Capstone Experience: Master of Public Health

This capstone project serves as an important piece of greater project studying bison handling. Bison production is both a growing and dangerous industry. As a new enterprise in many parts of the country and on tribal lands, significant numbers of untrained employees may be hired. This project undertakes methods to better understand the risks associated with bison handling and to thereby improve handling procedures and increase safety. Herd managers (from tribal and non-tribal sites), researchers, safety experts, and stakeholders were brought together for a roundtable meeting in Omaha. This meeting was designed to provide stakeholders a forum to discuss common …


Animal Experimentation: Working Towards A Paradigm Change, Kathrin Herrmann (Ed.), Kimberley Jayne (Ed.) Apr 2019

Animal Experimentation: Working Towards A Paradigm Change, Kathrin Herrmann (Ed.), Kimberley Jayne (Ed.)

eBooks

Animal experimentation has been one of the most controversial areas of animal use, mainly due to the intentional harms inflicted upon animals for the sake of hoped-for benefits in humans. Despite this rationale for continued animal experimentation, shortcomings of this practice have become increasingly more apparent and well-documented. However, these limitations are not yet widely known or appreciated, and there is a danger that they may simply be ignored. The 51 experts who have contributed to Animal Experimentation: Working Towards a Paradigm Change critically review current animal use in science, present new and innovative non-animal approaches to address urgent scientific …


Morphological And Color Variation In Poicephalus Parrots, Eileen Connon Apr 2019

Morphological And Color Variation In Poicephalus Parrots, Eileen Connon

Undergraduate Honors Thesis Projects

Though describing and delimiting species is one of the principle aspects of many disciplines within biology, there is often debate about which methods and concepts should be used to make such decisions. The Tobias criteria for quantitative species delimitation represent one attempt to create a standard method of delimiting species based on the morphological species concept. However, previous examples of using these criteria have not always been completely quantitative. This study uses quantitative morphological and color data of three Poicephalusparrots as a case study for the effectiveness of the Tobias criteria. The results show varying levels of support for …


Assessment Of Wildlife Rehabilitation Trends: Domestic Dog And Cat Attacks, Rachel Wilson Mar 2019

Assessment Of Wildlife Rehabilitation Trends: Domestic Dog And Cat Attacks, Rachel Wilson

Undergraduate Honors Thesis Projects

Human-wildlife conflict is a growing concern, and domestic cat predation is estimated to be responsible for billions of wildlife deaths every year in the Unites States. Domestic dog predation has been less well studied, but as dogs are the second most numerous pet in the U.S. after cats, their impact on native wildlife should also be assessed. One method to assess domestic animal predation is to examine trends in wildlife rehabilitation records. Using the online database WILD-ONe, I analyzed over 31,000 wildlife hospital records in the United States from 2011-2016. During this time span, data showed over 14,000 domestic dog …


Crispr-Mediated Gene Editing: Scientific And Ethical Issues, Jarrod Bailey Mar 2019

Crispr-Mediated Gene Editing: Scientific And Ethical Issues, Jarrod Bailey

Genetics Collection

There remains substantial evidence to warrant great concern over the poor efficiency and specificity of clustered regularly interspaced short palindromic repeats (CRISPR)-mediated genetic modification (GM), despite relatively minor improvements compared to other GM methods. These issues cause persistent, adverse, ethical, and scientific consequences for GM animals, which may never be sufficiently resolvable.


Salmonid Species Diversity Predicts Salmon Consumption By Terrestrial Wildlife, Christina N. Service, Andrew N. Bateman, Megan S. Adams, Kyle A. Artelle, Thomas E. Reimchen, Paul C. Paquet, Chris T. Darimont Mar 2019

Salmonid Species Diversity Predicts Salmon Consumption By Terrestrial Wildlife, Christina N. Service, Andrew N. Bateman, Megan S. Adams, Kyle A. Artelle, Thomas E. Reimchen, Paul C. Paquet, Chris T. Darimont

Habitat and Trophic Ecology Collection

1. Resource waves—spatial variation in resource phenology that extends feeding opportunities for mobile consumers—can affect the behaviour and productivity of recipient populations. Interspecific diversity among Pacific salmon species (Oncorhynchus spp.) creates staggered spawning events across space and time, thereby prolonging availability to terrestrial wildlife.

2. We sought to understand how such variation might influence consumption by terrestrial predators compared with resource abundance and intra- and interspecific competition.

3. Using stable isotope analysis, we investigated how the proportion of salmon in the annual diet of male black bears (Ursus americanus; n = 405) varies with species diversity and density of spawning …


Farm Animal Cognition—Linking Behavior, Welfare And Ethics, Christian Nawroth, Jan Langbein, Marjorie Coulon, Vivian Gabor, Susan Oesterwind, Judith Benz-Schwarzburg, Eberhard Von Borell Feb 2019

Farm Animal Cognition—Linking Behavior, Welfare And Ethics, Christian Nawroth, Jan Langbein, Marjorie Coulon, Vivian Gabor, Susan Oesterwind, Judith Benz-Schwarzburg, Eberhard Von Borell

Social Cognition Collection

Farm animal welfare is a major concern for society and food production. To more accurately evaluate animal farming in general and to avoid exposing farm animals to poor welfare situations, it is necessary to understand not only their behavioral but also their cognitive needs and capacities. Thus, general knowledge of how farm animals perceive and interact with their environment is of major importance for a range of stakeholders, from citizens to politicians to cognitive ethologists to philosophers. This review aims to outline the current state of farm animal cognition research and focuses on ungulate livestock species, such as cattle, horses, …


Starvation Resistance Is Associated With Developmentally Specified Changes In Sleep, Feeding And Metabolic Rate, Elizabeth B. Brown, Melissa E. Slocumb, Milan Szuperak, Arianna Kerbs, Allen G. Gibbs, Matthew S. Kayser, Alex C. Keene Feb 2019

Starvation Resistance Is Associated With Developmentally Specified Changes In Sleep, Feeding And Metabolic Rate, Elizabeth B. Brown, Melissa E. Slocumb, Milan Szuperak, Arianna Kerbs, Allen G. Gibbs, Matthew S. Kayser, Alex C. Keene

Life Sciences Faculty Research

Food shortage represents a primary challenge to survival, and animals have adapted diverse developmental, physiological and behavioral strategies to survive when food becomes unavailable. Starvation resistance is strongly influenced by ecological and evolutionary history, yet the genetic basis for the evolution of starvation resistance remains poorly understood. The fruit fly Drosophila melanogaster provides a powerful model for leveraging experimental evolution to investigate traits associated with starvation resistance. While control populations only live a few days without food, selection for starvation resistance results in populations that can survive weeks. We have previously shown that selection for starvation resistance results in increased …


Ecological Response Of Phytoplankton To The Oil Spills In The Oceans, Danling Tang, Jing Sun, Li Zhou, Sufen Wang, Ramesh P. Singh, Gang Pan Feb 2019

Ecological Response Of Phytoplankton To The Oil Spills In The Oceans, Danling Tang, Jing Sun, Li Zhou, Sufen Wang, Ramesh P. Singh, Gang Pan

Mathematics, Physics, and Computer Science Faculty Articles and Research

Oil spills in oceans have substantial influence on marine ecosystems. This study investigates 21 oil spills in the world. Analyzing Chlorophyll-a (Chl-a) from Moderate Resolution Imaging Spectroradiomerer (MODIS) data after Penglai oil spills on 4 June 2011, found a bloom with peak value of Chl-a (13.66 mg m−3) spread over an area of 800 km2 during 18–25 June 2011, and a pronounced increase in the monthly Chl-a concentration (6.40 mg m−3) on June 2012 in the Bohai Sea. Out of the 21 oil spills, 14 blooms were observed, while 11 …


Perceptual Lateralization Of Vocal Stimuli In Goats, Luigi Baciadonna, Christian Nawroth, Elodie F. Briefer, Alan G. Mcelligott Feb 2019

Perceptual Lateralization Of Vocal Stimuli In Goats, Luigi Baciadonna, Christian Nawroth, Elodie F. Briefer, Alan G. Mcelligott

Communication Skills Collection

Functional asymmetries, for example, the preferential involvement of 1 brain hemisphere to process stimuli, may increase brain efficiency and the capacity to carry out tasks simultaneously. We investigated which hemisphere was primarily involved in processing acoustic stimuli in goats using a head-orienting paradigm. Three playbacks using goat vocalizations recorded in different contexts: food anticipation (positive), isolation (negative), food frustration (negative), as well as 1 playback involving dog barks (negative) were presented on the left and right sides of the test subjects simultaneously. The head-orienting response (left or right) and latency to resume feeding were recorded. The direction of the head-orienting …


Perceptual Lateralization Of Vocal Stimuli In Goats, Luigi Baciadonna, Christian Nawroth, Elodie F. Briefer, Alan G. Mcelligott Feb 2019

Perceptual Lateralization Of Vocal Stimuli In Goats, Luigi Baciadonna, Christian Nawroth, Elodie F. Briefer, Alan G. Mcelligott

Social Cognition Collection

Functional asymmetries, for example, the preferential involvement of 1 brain hemisphere to process stimuli, may increase brain efficiency and the capacity to carry out tasks simultaneously. We investigated which hemisphere was primarily involved in processing acoustic stimuli in goats using a head-orienting paradigm. Three playbacks using goat vocalizations recorded in different contexts: food anticipation (positive), isolation (negative), food frustration (negative), as well as 1 playback involving dog barks (negative) were presented on the left and right sides of the test subjects simultaneously. The head-orienting response (left or right) and latency to resume feeding were recorded. The direction of the head-orienting …


Behavioral Flexibility Of A Generalist Carnivore, Sarah E. Daniels, Rachel E. Fanelli, Amy Gilbert, Sarah Sarah Benson‑Amram Feb 2019

Behavioral Flexibility Of A Generalist Carnivore, Sarah E. Daniels, Rachel E. Fanelli, Amy Gilbert, Sarah Sarah Benson‑Amram

United States Department of Agriculture Wildlife Services: Staff Publications

Innovative problem solving, repeated innovation, learning, and inhibitory control are cognitive abilities commonly regarded as important components of behaviorally flexible species. Animals exhibiting these cognitive abilities may be more likely to adapt to the unique demands of living in novel and rapidly changing environments, such as urbanized landscapes. Raccoons

(Procyon lotor) are an abundant, generalist species frequently found in urban habitats, and are capable of innovative problem solving, which makes them an ideal species to assess their behavioral flexibility. We gave 20 captive raccoons a multi-access puzzle box to investigate which behavioral and cognitive mechanisms enable the generation …


A Global Reassessment Of Solitary-Sociable Dolphins, Laetitia Nunny, Mark P. Simmonds Jan 2019

A Global Reassessment Of Solitary-Sociable Dolphins, Laetitia Nunny, Mark P. Simmonds

Personality Collection

Dolphins are typically regarded as highly social animals. However, some individuals live apart from their own species and may come to socialize with people through a recognized series of stages which are presented and expanded on in this paper. The term “solitary-sociable dolphins” has been used to describe these animals and such individuals have been identified from several different species and reported in many parts of the world. In many instances, the interactions with people that may follow their original isolation, and which typically become more intense over time, have created situations where the welfare of the animal has been …


Tiger Sharks Eat Songbirds: Scavenging A Windfall Of Nutrients From The Sky, J. Marcus Drymon, K. Feldheim, A. M.V. Fournier, E. A. Seubert, A. E. Jefferson, A. M. Kroetz, Sean P. Powers Jan 2019

Tiger Sharks Eat Songbirds: Scavenging A Windfall Of Nutrients From The Sky, J. Marcus Drymon, K. Feldheim, A. M.V. Fournier, E. A. Seubert, A. E. Jefferson, A. M. Kroetz, Sean P. Powers

University Faculty and Staff Publications

No abstract provided.


Elevated Sensitivity To Tactile Stimuli In Stereotypic Horses, Sabrina Briefer Freymond, Déborah Bardou, Sandrine Beuret, Iris Bachmann, Klaus Zuberbühler, Elodie Briefer Jan 2019

Elevated Sensitivity To Tactile Stimuli In Stereotypic Horses, Sabrina Briefer Freymond, Déborah Bardou, Sandrine Beuret, Iris Bachmann, Klaus Zuberbühler, Elodie Briefer

Personality Collection

Although stereotypic behaviors are a common problem in captive animals, why certain individuals are more prone to develop them remains elusive. In horses, individuals show considerable differences in how they perceive and react to external events, suggesting that this may partially account for the emergence of stereotypies in this species. In this study, we focused on crib-biting, the most common stereotypy displayed by horses. We compared how established crib-biters (“CB” = 19) and normal controls (“C” = 18) differed in response to a standard “personality” assessment test battery, i.e., reactivity to humans, tactile sensitivity, social reactivity, locomotor activity, and curiosity …


Nebraska Grazing Conference Proceedings, 2019, Center For Grassland Studies, University Of Nebraska-Lincoln Jan 2019

Nebraska Grazing Conference Proceedings, 2019, Center For Grassland Studies, University Of Nebraska-Lincoln

Nebraska Grazing Conference Proceedings

Rangeland Health Session

Managing Sand Deposits after the Flood by Daren Redfearn

Rangeland Health: What Is It and Why Do I Care?” by Patrick Shaver

Managing a Sandhills Ranch with an Eye to Soil Health by Dana Larsen

Grazinglands Management Session

Soil Health: Expectations and Evidence by Virginia Jin

Mob Grazing on Nebraska Sandhills Meadow by Walt Schacht

Effect of Length of Grazing Period on Upland Sandhills Range by Jerry Volesky

Carbon Input and Loss in Semi-arid Sandy Rangeland by Martha Mamo

Graze 365 by Jacob Miller

Soil Health and Grazing Strategies: Opportunities for Increasing Soil Water by Andrea Basche …


Trade-Offs Shape Carotenoid-Based Color Variation In Redheaded Pine Sawfly (Neodiprion Lecontei) Larvae, Maranda Gaines Jan 2019

Trade-Offs Shape Carotenoid-Based Color Variation In Redheaded Pine Sawfly (Neodiprion Lecontei) Larvae, Maranda Gaines

Lewis Honors College Capstone Collection

Carotenoids serve various ecological roles in animals including coloration, immune responses, and vision. Carotenoid-derived coloration is greatly emphasized in the literature, particularly relating to mate choice and aposematic warning. However, the trade-offs between the color and non-color functions of carotenoids are not thoroughly explored. In the redheaded pine sawfly (Neodiprion lecontei), some larval populations have yellow pigmentation, using carotenoids derived from their diets for aposematic warning coloration. Other larval populations are white in color, having genetically lost the ability to produce the yellow pigment. Because carotenoids are essential to life functions in both the yellow and white populations, we aim …


Predicting Black Bear Activity At Backcountry Campsites In Bryce Canyon National Park, Utah, Wesley G. Larson, Tom Smith Jan 2019

Predicting Black Bear Activity At Backcountry Campsites In Bryce Canyon National Park, Utah, Wesley G. Larson, Tom Smith

Human–Wildlife Interactions

Developing the capacity to predict black bear (Ursus americanus; bear) activity in a diversity of habitats will help conserve bear populations and their habitats and minimize human–bear conflicts. This capacity will be particularly important in areas that provide bear habitat and offer backcountry hiking and camping experiences. Bryce Canyon National Park (BRCA), located on the edge of the Paunsaugunt Plateau in southern Utah, USA, provides important bear habitat and offers visitors 12 backcountry campsites. To effectively manage these areas to minimize human–bear conflicts, park managers will need better information about black bear use of these campsites and other …


Tree Squirrels And Fishers In Northern California: The Effects Of Masting Hardwoods On Stand Use, Andria M. Townsend Jan 2019

Tree Squirrels And Fishers In Northern California: The Effects Of Masting Hardwoods On Stand Use, Andria M. Townsend

Cal Poly Humboldt theses and projects

In western North America, tree squirrels such as western gray (Sciurus griseus) and Douglas squirrels (Tamiasciurus douglasii) are potentially important prey for fishers (Pekania pennanti). Western gray squirrels in particular may be highly ranked due to their large body size. Masting trees including black oak (Quercus kelloggii) and tanoak (Notholithocarpus densiflorus) produce an important food source for tree squirrels; therefore, forest stands containing these trees may be useful to foraging fishers. I hypothesized that; 1) the abundance of western gray and Douglas squirrels in a stand is influenced by the …