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Morphology, Reproduction And Mortality Of Equus Hemionus Hemionus In Mongolia, Annegret Stubbe, Michael Stubbe, Nyamsuren Batsaikhan

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

There are collected materials on morphology, reproduction rate, and mortality of Dschiggetajs Equus h. hemionus between 2001 and 2006 in the South Gobi of Mongolia. It were found 794 skeletons of wild asses, 95% of these were pouched, ⅔ of them in winter coat. Pouching was also located in the protected areas of this region. The main living resources of Equus hemionus are registered in the South eastern Gobi. For the first time statistical data on morphology measurements of body length, tail and tuft, ear length, and hoofs are given. There are no significant differences between the sexes. The reproduction …

Up-Date On The Behaviour And Status Of The Critically Endangered Onager, Equus Hemionus Onager, From Iran, Laurent Tatin, Bijan F. Darreh-Shoori, Christophe Tourenq, David Tatin, Bijan Azmayesh

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The onager, Equus hemionus onager, an Asiatic wild ass endemic to Iran, is categorized as Critically Endangered on the IUCN Red List. Its biology and conservation requirements are poorly documented. We report some observations made in 1997 and 2000 on the behavior and ecology of the two remaining populations, located in the Touran Protected Area and the Bahram-e-Goor Reserve. Recent population counts by the Department of Environment of Iran (471 in the Protected Area, and 96 in the Reserve) are markedly lower than the count of 600-770 made in the 1970s in the Touran Protected Area. We observed social …

Asiatic Wild Ass In Israel, David A. Blank

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The Syrian wild ass (Equus hemionus hemippus) lived in Israel in historic times until the 1930s, when it disappeared from the entire Middle East region. Khulans from other subspecies (E. h. onager - six individulas and E. h. kulan - five individuals) were brought to Israel from European zoos during 1968-69 (the authors of project considered that onager and kulan were the same subspecies) and these animals bred together in the semi-captivity conditions of the Haibar Yotvata Reserve. Later, during a 12-year period from 1982 through 1993, six groups totalling 38 khulans (17 males and 21 females) …

Surviving A Drought: Population Dynamics Of Ochotona Pallasi Pricei In A Dry Steppe, Gobi Altai, Mongolia, Karin Nadrowski, Georg Miehe

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Two distinct life history traits are described from the genus Ochotona, the group of “burrowing” species exhibiting high but fluctuating population densities and the “non-burrowing” species exhibiting relatively stable low population densities. The life history traits are linked to ecosystem dynamics with climatically variable steppe environments hosting “burrowing” species and relatively stable mountainous and rocky habitats hosting “non-burrowing” species. There are few intermediate species, living in both steppe and rocky environments. This study presents survival rates, reproductive rates and a tentative model of population dynamics for Ochotona pallasi pricei, an intermediate species with respect to habitat preference. We …

The Emerging Dziggetai (Equidae: Equus Hemionus Pallas): An Illustrated History Of Taxonomic Concepts For The Identification, Classification, And Distribution Of Hemiones From Central Asia, Arnd Schreiber

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The history of the discovery and the exploration of the hemione populations (Equus hemionus PALLAS) from Central Asia (Mongolia, northwest China, northeast Tibet, Kazakhstan, and Russian Siberia) from the 13th century to approximately 1950 are compiled with the aim to provide materials for a taxonomic and population genetic revision of the dziggetais. Data on their phenotypic differentiation, historical distribution status, and their husbandry history in European zoological gardens, are reviewed, as is the gradual emergence of concepts of how to identify and to classify these populations. Hemiones were an important historical case in the discussion of how to represent …

Quo Vadis Equus Hemionus Hemionus In Mongolia?, Annegret Stubbe, Michael Stubbe, O. Shagdarsuren, Nyamsuren Batsaikhan, Ravčigijn Samjaa

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Inspired by studies on biodiversity in the South Gobi we were confronted with the human impact on animal communities and endangered animal species, especially the Asiatic wild ass. So the idea was born to organize an international conference with the aim to actualize the status of Equus hemionus in Central Asia and to find ways for better conservation of this species and its habitats. A short review is given on historical exploration, decreasing area and population number of the Dschiggetai. Today the range of the nominate form of Equus hemionus is located in the Gobi region of southern Mongolia/northern China …

Remarks On The Social System Of The Mongolian Wild Ass (Equus Hemionus Hemionus), Gertrud Neumann-Denzau, Helmut Denzau

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

So far the social system of Mongolian wild asses is non-uniformly interpreted. The authors describe their own observations from different parts of Mongolia and review data from available publications. The arguments support a territorial social system, which agrees with the social system of all other wild asses in Asia and Africa.

Asiatic Wild Asses In The Literature: What Do We Need To Know Now?, Sarah R.B. King

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Asiatic wild asses were once found across central Asia, but over time their range has been reduced until today all subspecies are under threat. This paper examined the literature about wild asses with an aim to illustrating gaps in our knowledge as a target for future research.

A search was made on BIOSIS ISI for the following strings: Equus hemionus, wild ass, khulan, kulan, kiang, khur, and onager. In total 61 relevant references were found, published between 1900 and 2005. Most of these references were published in the 1990s, with only one published between 1900 and 1970. Khur and …

Comparative Demography And Dietary Resource Partitioning Of Two Wild Ranging Asiatic Equid Populations, Ellen Schulz, Thomas Kaiser, Annegret Stubbe, Michael Stubbe, Ravčigijn Samjaa, Nyamsuren Batsaikhan, Joachim Wussow

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Tooth wear signatures allow inference on the dietary traits of herbivorous ungulates. Comparing dietary regimes of taxonomically closely related populations further allows inference on habitat structure and food availability. The mesowear method of tooth wear evaluation has opened up a pathway to reconstruct subtle differences in dietary behavior and resource partitioning based on skeletal material as the only source of information. Eighty cheek dentitions of Asian wild asses (Equus hemionus) from the Southern Gobi (Mongolia) and 61 dentitions of African free ranging donkeys (Equus asinus) from the Emirate Sharjah (United Arab Emirates) were investigated for their …

Population Assessment Of Khulan (Equus Hemionus) In Mongolia, Badamjav Lkhagvasuren

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The data presented here suggest that 18,411 (± 224) khulan inhabit southern Mongolia with a density of 1.4 individuals per 1,000 km² within the 157,525 km² territory of its actual distribution. The Dornogobi province contains the highest number of khulan with a mere 67%, while 20% are found in the Ömnögobi province, 12% are in Djungarian Gobi and only 1% in the Gobi-Altay and Bayankhongor provinces. According to the proportion of foals and yearlings within each aimag, the Dornogobi, Ömnögobi east and Khovd populations have an average reproduction rate.

Kulan (Equus Hemionus Pallas 1775) In Turkmenistan, Victor S. Lukarevskiy, Yu. K. Gorelov

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The kulan (Equus hemionus kulan) is the only odd-toed ungulate left in the wild in northern Eurasia. In the 1930s, the kulan survived in southern Turkmenistan only. In the 1940s, kulans were found only in an area named Badkhyz in Turkmenistan. To conserve the last natural population of this subspecies of kulan the Badkhyz Natural Reserve was established in 1941. The historical and contemporary distribution, fluctuation in numbers, conservation and sci research of kulan populations in the Badkhyz Natural reserve are reviewed in this paper.

First Records And First Proven Breeding Of Lesser Grey Shrike, Lanius Minor, In Mongolia, Markus Deutsch, Axel Bräunlich

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

During a trip to the Dzungarian Gobi in southwestern Mongolia, a group of German birdwatchers found several territorial lesser grey shrikes, Lanius minor, and documented an active nest. The species had not previously been recorded from Mongolia. Observations were made near Bulgan gol (Bulgan river; nest coordinates: 46°6’N 91°32’E, 1190 m a.s.l.) in Khovd aimag (Khovd province) on June 24, 2006. Besides two territorial pairs, of which a female could be photographically documented on the nest, at least four more territorial adults were found within an area of about 1.5 km². Based on these records, an abundance …

Numbers, Distribution And Social Structure Of Kiang (Equus Kiang Moorcroft 1841) Population In The Southwestern Part Of Tibet, China, Natalia V. Paklina, Chris Van Orden

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In September - October 1998 we made an extensive survey in southwest Tibet to study numbers, distribution and social structure of the kiang population in this remote part of its range. Kiangs were sighted between 29º 40’ and 32º 50’ N, and 81º- 86º E. They preferred broad flat valleys with lakes at an altitude from 4,000 up to 5,000 m. Small kiang groups and solitary animals were sighted in Transhimalaya. There were no kiangs in Tsangpo (Brahmaputra) River valley, in the sand desert near Ali and in the Clay Mountains near Tsada. Seven old kiang corpses were found along …

Some Information On The Ecology Of Khulan (Equus Hemionus Pallas, 1775) In The Western Part Of The South Gobi Province, Mongolia, D. Tsendjav, S. Purevsuren

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

This short study provides information on the density and distribution of khulans in September 2003 and speculates about factors that negatively influence the khulan population in the sums of Khurmen, Bayandalai, Noyon, Sevrei, and Gurvantes in the South Gobi province.

Turkmenian Kulan (Equus Onager Kulan) Captive Population, Anna Mekarska

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The Turkmenian kulan (Equus onager kulan) is critically endangered, fewer then 1,500 animals are left in the wild. In 2004, there were additional 570 Turkmenian kulans living in captivity: 156 specimens in Euro-Asian zoos, 83 in Askania Nova Reserve and 331 in Canyon Colorado Equid Sanctuary. Population size in EAZA region is decreasing both in terms of numbers and genetic parameters. Those facts underline the urgent need for coordinating development of Eurasian kulan’s captive population. From 2004 onwards, large efforts to create healthy breeding groups in zoos raised bigger interest in this species in European zoos.

Identification Of Herder-Wild Equid Conflicts In The Great Gobi B Strictly Protected Area In Sw Mongolia, Petra Kaczensky, N. Enkhsaihan, O. Ganbaatar, Christian Walzer

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In protected areas of Mongolian overgrazing, competition with wild ungulates and poaching are important management concerns. The Great Gobi B Strictly Protected Area (SPA) in SW Mongolia is a re-introduction site for the Przewalski’s horse (Equus ferus przewalskii), a stronghold of the Asiatic wild ass (Equus hemionus), and remains an important grazing area for seminomadic herders. We show the power of simple inventory and monitoring methods to assess herder-wildlife conflicts, by combining data of: (1) human and livestock demographic data, (2) migration patterns, of semi-nomadic herders, (3) monthly surveys of wild- and domestic ungulates, and (4) …

Kiangs (Equus Kiang, Moorcroft 1841) In Sikkim, India, Natalia V. Paklina, Chris Van Orden

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Between the 5^th and 10^th of March 2001 we completed a 450 km route by car in Northern Sikkim. During the trip along the southern slopes of the Himalayas and Sikkim Plateau we didn’t see a single kiang. According to the local people and military personnel in Sikkim, kiangs are known only in the outer limits of the Sikkim Plateau, where wild animals have a possibility to migrate free over the Chinese-Indian border. In winter only small groups of kiang incidentally come to the Sikkim Plateau from South Tibet. But in May more of them come there and …

Current Status Of The Khulan (Equus Hemionus) In The Trans-Altai Gobi, Enkhbileg Dulamtseren, Adiya Yadamjav, Dovchindorj Ganbold, Mijidorj Batmunkh

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Since 1975, 25 wildlife surveys have been conducted in the Trans-Altai Gobi. In this paper we summarize results from the last 11 surveys, conducted between October 2000 and April 2005, and discuss our findings together with results from previous surveys.

Assessment Of Non-Metric Skull Characters And Age Determination In The Asiatic Wild Ass Equus Hemionus: A Methodological Approach, Hermann Ansorge, Annegret Stubbe, Nyamsuren Batsaikhan, Ravčigijn Samjaa, Michael Stubbe

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The report presents the first information on the use of non-metric skull characters and age determination by tooth section for Asiatic wild asses. It was initiated by a collection of more than 400 skulls of the Asiatic wild ass from the Bordzongijn-Gobi 2002-2004. The methodological overview gives a first research prospect in Asiatic Wild Ass concerning the use of non-metric skull characters as a morphological tool for assessing genetic variability, determining epigenetic divergences and evaluating fluctuating asymmetry. The time course of tooth eruption and replacement in the Asiatic wild ass is described with regard to its use for age determination. …

Semi-Wild Population Of Kulans In The Bukhara Breeding Centre And Their Co-Habitation With Przewalski’S Horses, Karim Bahloul, Olga B. Pereladova, Natalia V. Soldatova, Ekaterina V. Sidorenko, Antoine J. Sempere

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Asiatic wild asses and Przewalski's horses initially inhabited steppe, semi-desert and desert areas, but Przewalski's horses became extinct in the wild, and kulans are under threat of disappearance. The Bukhara Breeding Centre (Uzbekistan) was created in 1976 for conservation and reintroduction of wild ungulate species. In 1977-1978, five kulans (two males and three females) from Barsa-Kelmes Island on the Aral lake were introduced to the reserve. The group increased to 25-30 animals in 1995-1998, when five Przewalski's horses from Moscow and St. Petersburg zoos were introduced to the same territory. We analyzed the home ranges, preferred habitats and social interactions …

Social Networks In Wild Asses: Comparing Patterns And Processes Among Populations, Daniel I. Rubenstein, Siva Sundaresan, Ilya Fischhoff, David Saltz

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Asiatic wild asses inhabit some of the most arid environments in the world. All live in fissionfusion societies, but demography varies and the deserts in which they live often differ in subtle ways. Characterizing details of social structure of wild ass populations has been a challenge and has made it difficult to determine causes and consequences of any differences that might exist. We use network theory to compare the social structures of two populations of Asiatic asses/ onagers inhabiting the Negev desert, Israel and khur of the Little Rann of Kuch, India and show that populations differ in important structural …

Chlorophacinone Baiting For Belding’S Ground Squirrels, Craig A. Ramey, George H. Matschke, Richard M. Engeman

United States Department of Agriculture Wildlife Services: Staff Publications

The efficacy of using 0.01% chlorophacinone on steam-rolled oat (SRO) groats applied in CA alfalfa by spot-baiting/hand baiting around burrow entrances (~11.5 g) to control free-ranging Belding's ground squirrels (Spermophilus beldingi) were compared in 6 randomly assigned square treatment units (TUs). Four TUs were given the rodenticide and 2 treated with placebo bait. Each TU was a 0.4 ha square surrounded by a similarly treated 5.5 ha square buffer zone. Baits were applied on May 13 and re-applied, on May 20 and May 22, after 7 days of un-forecasted cool wet weather greatly reduced their above ground activity. Pesticide (EPA …

Estimates Of Breeding Bird Populations In The Sheyenne National Grassland, North Dakota, Mary Ann Cunningham, Douglas Johnson, Daniel N. Svingen

The Prairie Naturalist

We conducted a two-year survey of breeding birds in the Sheyenne National Grassland, North Dakota, to estimate total populations of breeding birds. The Grassland is of interest because it provides wildlife habitat and recreational birding opportunities. Indicated breeding pairs were counted on 100-m wide belt transects during morning surveys from late May to early July in 2002 and 2003. We surveyed approximately 6 to 7% of the Grassland each year. The most abundant species in both years were grasshopper sparrow (Ammodramus savannarum), red-winged blackbird (Agelaius phoeniceus), claycolored sparrow (Spizella pallida), Savannah sparrow (Passerculus …

Ecology Of The Missouri River. Progress Report, Dingell-Johnson Project F-75-R-24, Supplement I - Missouri River Creel Survey, Camp Creek To Kansas State Line, 1 April Through 13 October 2006., Gerald Mestl

Nebraska Game and Parks Commission: White Papers, Conference Presentations, and Manuscripts

This report describes Missouri River activities and results related to a channelized Missouri River creel survey conducted from 29 March through 10 October 2003. This is the fourth of a planned annual creel survey to be conducted on alternating sections of the channelized Missouri River to measure changes in recreational fishing activity, especially those changes due to large scale habitat restoration efforts. Future reports will contain additional analyses of these data. Anglers spent over 30,000 hours fishing the Missouri River from Camp Creek (rk 883.5) to the Kansas state line (rk 790.2) during the survey period. Effort peaked during the …

Celastrina Serotina (Lycaenidae: Polyommatinae): A New Butterfly Species From The Northeastern United States And Eastern Canada, Harry Pavulaan, David M. Wright

The Taxonomic Report of the International Lepidoptera Survey

A new light-venter Azure species, Celastrina serotina, is described from the northeastern United States and eastern Canada. The flight period of this univoltine spring species is interpolated (allochrony) between its sympatric congeners, C. lucia Auctorum (not Kirby) and C. neglecta (Edwards). In this regard it is similar to C. neglectamajor Opler & Krizek in the southern Appalachians and C. idella Wright & Pavulaan on the New Jersey coastal plain. In a significant portion of its range, C. serotina larvae feed on eriophyid mite-formed galls on the upper surface of Black Cherry Prunus serotina Ehrh. …

Biodemography Of A Long-Lived Tephritid: Reproduction And Longevity In A Large Cohort Of Female Mexican Fruit Flies, Anastrepha Ludens, James R. Carey, Pablo Liedo, Hans-Georg Müller, Jane-Ling Wang, Damla Senturke, Lawrence G. Harshman

Lawrence G. Harshman Publications

Age of sexual maturity, daily and lifetime reproductive rates, and life span were recorded in a laboratory cohort of Mexican fruit flies consisting of over 1100 females maintained individually. The results revealed that, relative to the medfly, the Mexfly is slower maturing (14 vs 17 days), more fecund (1400 vs 650–1100 eggs/female), and longer lived (50 vs 35 days). The results reinforced the generality of several earlier findings on the medfly including the deceleration of mortality at older ages and the weakness of the correlation between the rate of egg laying at early ages and both subsequent reproduction and remaining …

The Symmetry Of Longevity, Lawrence G. Harshman, Hans-Georg Müller, Xueli Liu, Yue Wang, James R. Carey

Lawrence G. Harshman Publications

We tested the hypothesis that relatively symmetrical flies live longer. Vein measurements on the left and right wings from the same individual were used to characterize bilateral symmetry in relationship to age-specific survival in defined cohorts. A longitudinal cohort study supported the hypothesis of a correlation between bilateral symmetry and longevity. For another type of experiment, wings were removed from females and males at approximately the beginning of adult life. Thus, there would be no effect of wings per se on adult survival. These wings were measured to characterize bilateral asymmetry, and the day of death of each dewinged individual …

Ecology Of The Missouri River: Missouri River Creel Survey, Bellevue Bridge To Camp Creek, 3 April Through 29 May 2004, Gerald Mestl

Nebraska Game and Parks Commission: White Papers, Conference Presentations, and Manuscripts

The Nebraska Game and Parks Commission's strategic plan has stated the following management goal for the Missouri River: Restore, protect, and maintain the diversity of historic Missouri River habitats, resources, and ecosystem functions in order that present and future generations may enjoy consumptive and non-consumptive outdoor recreational opportunities (NGPC 1996). To accomplish this goal the Nebraska Game and Parks Commission identified the following five objectives: • To restore terrestrial and aquatic floodplain habitat types by 2008. This would include old oxbows, chutes, side channels, sand bars, backwaters, wetlands, and other shallow water habitats. To restore ftows that reflect the natural …

Taxonomic Analysis Of The Genus Megisto (Satyridae) In The Eastern United States, Part I: Types, Type Localities, And Taxonomic Relationships Of The Available Names, Ronald R. Gatrelle

The Taxonomic Report of the International Lepidoptera Survey

The genus Megisto Hübner, 1819 in the eastern United States and southeastern Canada is currently comprised of the original epithets Papilio eurytus Fabricius, 1775, Papilio cymela Cramer, 1777, Papilio eurytris, Fabricius, 1793, and Neonympha eurytris viola Maynard, 1891. Because the eastern Megisto is considered by some, including the present author, to contain two or more sibling species, the types and type localities of each of these names were studied. Where no type was found and no type locality fixed, typification was established by lectotypification or neotypification and a type locality was fixed. The taxonomic relationship of these taxa were …

2002 Licensed Angler Survey: Summarized Results, Keith L. Hurley, Kristin L. Duppong-Hurley

Nebraska Game and Parks Commission: White Papers, Conference Presentations, and Manuscripts

On October 22, 2002, 5000 six-page surveys (Appendix 3) were sent to a stratified-random sample of Nebraska anglers drawn from the 2001 licensed anglers database. The database included all anglers who purchased a fishing license in 2001 as well as all lifetime fishing permit holders who purchased their license before or during 2001. The database did not include those possessing a free permit, such as veteran’s or senior citizen permits. The survey subsample (2.6% of the 2001-licensed angler database) was stratified such that the proportions of surveys in the subsample (4366 residents and 634 non-residents) matched the proportions of resident/non-resident …