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Articles 2611 - 2640 of 3460

Full-Text Articles in Ecology and Evolutionary Biology

How Sanak Aleut Fit Into The Intertidal Food Web, Jennifer A. Dunne, Spencer Wood, Roly Russell, Nancy Huntly, Matthew Betts, Herbert Maschner Aug 2012

How Sanak Aleut Fit Into The Intertidal Food Web, Jennifer A. Dunne, Spencer Wood, Roly Russell, Nancy Huntly, Matthew Betts, Herbert Maschner

Biology Faculty Publications

Almost everyone learns about the concept of ecological "food chains" in grade school. An example of a marine food chain is shown on the left-hand side of the chart to the left: phytoplankton are eaten by mussels, which are eaten by octopus, which are eaten by sea otters, which are eaten by humans. These feeding interactions can be abstractly represented as five "nodes" connected in a single chain by four links, as shown on the right-hand side of the diagram to the left. The food chain shown is one of the thousands of interlinked food chains that make up the …


The Ecological Interactions Of Sanak, Nancy Huntly, Spencer Wood, Roly Russell Aug 2012

The Ecological Interactions Of Sanak, Nancy Huntly, Spencer Wood, Roly Russell

Biology Faculty Publications

Ecology is not only about the interesting biology of organisms, as described for Sanak Island in the last chapter, but also about interactions: interactions between animals (including people), plants, microbes, the habitats that they live in, and the environments that surround them. The ecology of Sanak Island today reflects not only today's conditions, but also shows the imprint of the distant history of interactions of the island, especially, as we will see below, the imprint of the Aleut people that have made this island their home.


The Likelihood Of Extinction Of Iconic And Dominant Herbivores And Detritivores Of Coral Reefs: The Parrotfishes And Surgeonfishes, Mia T. Comeros-Raynal, John H. Choat, Beth A. Polidoro, Kendall D. Clements, Rene Abesamis, Matthew T. Craig, Muhammad E. Lazuardi, Jennifer Mcilwain, Andreas Muljadi, Robert F. Myers, Cleto L. Nanola Jr., Shinta Pardede, Luiz A. Rocha, Barry Russell, Jonnell C. Sanciangco, Brian Stockwell, Heather Harwell, Kent E. Carpenter Jul 2012

The Likelihood Of Extinction Of Iconic And Dominant Herbivores And Detritivores Of Coral Reefs: The Parrotfishes And Surgeonfishes, Mia T. Comeros-Raynal, John H. Choat, Beth A. Polidoro, Kendall D. Clements, Rene Abesamis, Matthew T. Craig, Muhammad E. Lazuardi, Jennifer Mcilwain, Andreas Muljadi, Robert F. Myers, Cleto L. Nanola Jr., Shinta Pardede, Luiz A. Rocha, Barry Russell, Jonnell C. Sanciangco, Brian Stockwell, Heather Harwell, Kent E. Carpenter

Biological Sciences Faculty Publications

Parrotfishes and surgeonfishes perform important functional roles in the dynamics of coral reef systems. This is a consequence of their varied feeding behaviors ranging from targeted consumption of living plant material (primarily surgeonfishes) to feeding on detrital aggregates that are either scraped from the reef surface or excavated from the deeper reef substratum (primarily parrotfishes). Increased fishing pressure and widespread habitat destruction have led to population declines for several species of these two groups. Species-specific data on global distribution, population status, life history characteristics, and major threats were compiled for each of the 179 known species of parrotfishes and surgeonfishes …


Quantitative Acoustic Analysis Of The Vocal Repertoire Of The Golden Rocket Frog (Anomaloglossus Beebei), Beth A. Pettitt, Godfrey R. Bourne, Mark A. Bee Jun 2012

Quantitative Acoustic Analysis Of The Vocal Repertoire Of The Golden Rocket Frog (Anomaloglossus Beebei), Beth A. Pettitt, Godfrey R. Bourne, Mark A. Bee

Biology Faculty Publications

This study describes the vocal repertoire of the Guyanan golden rocket frog, Anomaloglossus beebei, a bromeliad specialist with biparental care. Using multivariate analyses of nine call properties, as well as the occurrence of nonlinear phenomena, three signal types were distinguished—advertisement, courtship, and aggressive calls. Although all three call types were composed of a short series of rapidly repeated pulses, advertisement calls were produced at higher amplitudes and had longer pulse durations than both courtship calls and aggressive calls. Courtship calls exhibited lower dominant frequencies than both advertisement and aggressive calls, which had similar dominant frequencies. Aggressive calls had more …


The Spatial Games: Investigating The Spatial Distribution Of Anax And Select Prey As A Function Of Predator-Prey Interactions, Shane Elizabeth Johnson, Nene Kumashe Ugbah Jun 2012

The Spatial Games: Investigating The Spatial Distribution Of Anax And Select Prey As A Function Of Predator-Prey Interactions, Shane Elizabeth Johnson, Nene Kumashe Ugbah

Biological Sciences

Our study investigated the spatial distribution and movement behavior of predatory dragonfly larvae (Anax) and of two prey types: mosquito larvae and amphipods. Predator-prey interactions have important consequences for the population dynamics of both predator and prey groups and these interactions can shape community structure. We measured behavior of each prey type in the presence of the Anax predator and the behavior of the predator in the presence of these alternative prey types. Observations were made in five-gallon aquaria where a grid pattern allowed us to track the number of moves made by individuals. We compiled data from …


Seasonal Food Habits Of Burrowing Owls (Athene Cunicularia) In Human-Altered Landscapes, Carie Marie Wingert Jun 2012

Seasonal Food Habits Of Burrowing Owls (Athene Cunicularia) In Human-Altered Landscapes, Carie Marie Wingert

Master's Theses

In 2004, I initiated a year-long study to investigate the food habits of burrowing owls (Athene cunicularia). Burrowing owls have been found in a variety of human-altered landscapes; however, little is known about burrowing owl food habits in urban landscapes. Burrowing owl food habits during the non-breeding season are also largely undocumented, despite increasing concern over the survival of overwintering burrowing owls. Differences in prey consumption between reproductive and non-reproductive owls during the breeding season have not yet been examined. I collected pellets over a 12 month period at four study sites affected by different levels of human …


Macrophyte Communities Of Lake Winnebago: Baseline Study Of Species Composition With Abundances And Water Quality Conditions, Mackenzie Kessenich May 2012

Macrophyte Communities Of Lake Winnebago: Baseline Study Of Species Composition With Abundances And Water Quality Conditions, Mackenzie Kessenich

Lawrence University Honors Projects

Historical records from Lake Winnebago show minimal macrophyte growth; however, reports from recent years claim that macrophyte growth in some areas of the lake has reached nuisance levels. This study aimed to investigate the species of macrophytes present and their abundances in four near-shore locations, as well as measurements of multiple water quality conditions. Rake sampling was used to identify species and quantify their abundances and distributions. In addition, data were collected on light penetration, Secchi depths, and suspended algae chlorophyll concentrations at each site. These data from shallow near-shore sites reveal trends in changing water clarity and light penetration …


Will Extreme Climatic Events Facilitate Biological Invasions?, Jeffrey M. Diez, Carla M. D'Antonio, Jeffrey S. Dukes, Edwin D. Grosholz, Julian D. Olden, Cascade J.B. Sorte, Dana M. Blumenthal, Bethany A. Bradley, Regan I. Early, Inés Ibáñez, Sierra J. Jones, Joshua J. Lawler, Luke P. Miller May 2012

Will Extreme Climatic Events Facilitate Biological Invasions?, Jeffrey M. Diez, Carla M. D'Antonio, Jeffrey S. Dukes, Edwin D. Grosholz, Julian D. Olden, Cascade J.B. Sorte, Dana M. Blumenthal, Bethany A. Bradley, Regan I. Early, Inés Ibáñez, Sierra J. Jones, Joshua J. Lawler, Luke P. Miller

Faculty Publications, Biological Sciences

Extreme climatic events (ECEs) – such as unusual heat waves, hurricanes, floods, and droughts – can dramatically affect ecological and evolutionary processes, and these events are projected to become more frequent and more intense with ongoing climate change. However, the implications of ECEs for biological invasions remain poorly understood. Using concepts and empirical evidence from invasion ecology, we identify mechanisms by which ECEs may influence the invasion process, from initial introduction through establishment and spread. We summarize how ECEs can enhance invasions by promoting the transport of propagules into new regions, by decreasing the resistance of native communities to establishment, …


Devil In A Hula Skirt: Comparative Conservation Efforts Between The United States And Australia Using Two Endemic Species As Case Studies, Sarah Catherine Kuchinsky May 2012

Devil In A Hula Skirt: Comparative Conservation Efforts Between The United States And Australia Using Two Endemic Species As Case Studies, Sarah Catherine Kuchinsky

Undergraduate Honors Thesis Collection

There is a biodiversity crisis in our midst. An abundance of technology and innovation in the last 200 years has revamped our world but at the cost of pushing many species to extinction. Since the passing of the Endangered Species Act by Congress in 1973, more than 100 species in the United States have disappeared (Wilson 2006). The International Union for the Conservation of Nature (IUCN) stats that "19,265 species out of the 59,507 so far assessed as threatened with extinction," (IUCN 2011). Extinction is the result of a combination of factors caused by human environmental disturbance. Disturbances in the …


Evolution Of The Bacteriorhodopsin Gene Bop In Haloarchaea, Maulik Jitesh Jani May 2012

Evolution Of The Bacteriorhodopsin Gene Bop In Haloarchaea, Maulik Jitesh Jani

Honors Scholar Theses

Rhodopsins are among the most studied protein families. They all incorporate a light based chromophore and an opsin. They are found in all three domains of life and can be found in diverse environments, such as the membrane of the haloarchaea, Haloarcula (Haa) marismortui, growing on salt flats in Death Valley, to within human tissues and deep sea water (Briggs and Spudich 2005). Rhodopsins can serve multifarious purposes, from phototaxis away from harmful light to efficient energy generation. Bacteriorhodopsin is the best studied rhodopsin in haloarchaea. The evolutionary pattern of bacteriorhodopsin through its gene, bop, is the main focus of …


Genetic Variation In Potentially Virulent Vibrio Parahaemolyticus From The Northern Gulf Of Mexico, Nicholas Felix Noriea Iii May 2012

Genetic Variation In Potentially Virulent Vibrio Parahaemolyticus From The Northern Gulf Of Mexico, Nicholas Felix Noriea Iii

Dissertations

Vibrio parahaemolyticus (Vp) is a gram-negative bacterium found naturally in marine and estuarine environments. Vp is found in oysters including those which are later consumed by the public. Sub-populations of potentially virulent Vp contain specific virulence factors and are relevant human pathogens capable of causing gastroenteritis, wound infection, and death. The tdh and trh genes, both encoding hemolysins, have been correlated with the majority of clinical Vp isolates but have not been shown to be the definitive virulence factors.

A total of 146 Vp isolates from the northern Gulf of Mexico were collected and probed …


Comparison Of Creel Survey Data To Traditional Sampling Techniques In Pit-Lake Fisheries Of Muhlenberg County, Kentucky, Derek L. Rupert May 2012

Comparison Of Creel Survey Data To Traditional Sampling Techniques In Pit-Lake Fisheries Of Muhlenberg County, Kentucky, Derek L. Rupert

Masters Theses & Specialist Projects

Populations of largemouth bass, Micropterus salmoides, and bluegill, Lepomis macrochirus, were evaluated from five pit-lakes in Muhlenberg County, Kentucky, to determine if accurate proportional stock density (PSD) data can be obtained from a mandatory creel survey. It was hypothesized that the proportion of stock-to-quality (300-400mm) and quality (+400mm) largemouth bass from four years (2007-2010) of creel survey data would be statistically similar to those generated through on-site sampling in 2011. Fish were collected via a combination of gill netting, seining, hook-and-line fishing, and boat-mounted electro-fishing. In two of the pit-lakes, the sampling-generated length frequency data was not significantly …


The Ecological Roles Of Podostemum Ceratophyllum And Cladophora In The Habitat And Dietary Preferences Of The Riverine Caddisfly Hydropsyche Simulans, Brenna Tinsley May 2012

The Ecological Roles Of Podostemum Ceratophyllum And Cladophora In The Habitat And Dietary Preferences Of The Riverine Caddisfly Hydropsyche Simulans, Brenna Tinsley

Mahurin Honors College Capstone Experience/Thesis Projects

The net-spinning caddisfly Hydropsyche simulans can be a common inhabitant of shallow reaches in riverine systems, and is easily the most common hydropsychid in the upper Green River, Kentucky. This study was performed in summer 2011 and focused on two main questions: 1. Do the larvae of the riverine caddisflies H. simulans and Cheumatopsyche preferentially inhabit dense patches of P. ceratophyllum compared to bare substrates in the upper Green River?, and 2. Do larvae of H. simulans and Cheumatopsyche consume the filamentous alga Cladophora during the annual late summer algal bloom in the upper Green River? Densities of both hydropsychid …


The Effects Of Invasive Grasses On The Survival And Germination Of Native Forbs, Rachel A. King Apr 2012

The Effects Of Invasive Grasses On The Survival And Germination Of Native Forbs, Rachel A. King

Scripps Senior Theses

Invasive species are an increasingly severe conservation problem that can dramatically alter native plant communities. Although ecologists have proposed many mechanisms for the dominance of invasive species, superior competitive ability is one of the longest standing hypotheses. In winter annual plant communities, germination timing affects competition among plants. I hypothesized that grass invasion has changed the costs and benefits of early and late germination, and that species with flexible germination timing would compete better with invasives. Experiments at the Bernard Field Station in Claremont, CA and germinator trials in a controlled environment were used to examine the survivorship and germination …


Nutrient Limitation Of Algal Biomass In Streams Of The Sacramento Mountains, New Mexico, Bianca C. Luong^, Vanessa Lougheed* Apr 2012

Nutrient Limitation Of Algal Biomass In Streams Of The Sacramento Mountains, New Mexico, Bianca C. Luong^, Vanessa Lougheed*

COURI Symposium Abstracts, Spring 2012

Anthropogenic activities often alter the chemistry and overall quality of freshwater ecosystems. Algae are good indicators of ecosystem health because they respond quickly to influxes of nutrients. In this study, we used algal biomass and nutrient limitation experiments to assess the water quality in streams of the Sacramento Mountains of New Mexico. We sampled 7 sites monthly from May to September 2011, with the exception of June (n = 27) for both periphyton (attached algae) and phytoplankton. Phytoplankton algal biomass was significantly related to phosphorus in the water column; while there were no relationships between periphyton biomass and nutrients. The …


The Causes And Consequences Of Increased Active Layer Depth In Arctic Tundra Ponds Over The Past 40 Years, Gabriela Contreras ^, Vanessa L. Lougheed * Apr 2012

The Causes And Consequences Of Increased Active Layer Depth In Arctic Tundra Ponds Over The Past 40 Years, Gabriela Contreras ^, Vanessa L. Lougheed *

COURI Symposium Abstracts, Spring 2012

Active layer is used to describe the upper most layer of soil that thaws each summer in the Arctic. With a warming Arctic, permafrost is expected to thaw and active layer depth to increase, thus releasing organic material and nutrients into aquatic environments; however, there are few long-term datasets available to test these predictions. The ponds at the International Biological Program (IBP) site in Barrow, Alaska represent one of the only locations in the Arctic with long-term aquatic ecosystem data. The objective of this study was to determine whether the temperature of tundra ponds has increased over the past 40 …


Human-Wildlife Interaction In Serengeti And Ngorongoro Districts Of Tanzania: A Case Study On Small Mammals, Fj Magige Mar 2012

Human-Wildlife Interaction In Serengeti And Ngorongoro Districts Of Tanzania: A Case Study On Small Mammals, Fj Magige

Tanzania Journal of Science

In the Serengeti and Ngorongoro Districts, small mammals are said to provide protein and income to the local people. However, they are simultaneously reported to conflict with farming activities. These conflicting aspects have not been investigated there. The present article considers humansmall mammal interactions in six villages adjacent to the protected areas. Data were obtained through questionnaire administration to the local people in the districts. Small mammals were hunted in the two districts for food purposes and some villagers declared that they were earning income from selling small mammals products. Seventy percent of the respondents in Serengeti District claimed that …


Plant Species Richness And Species Area Relationships In A Florida Sandhill, Monica Ruth Downer Mar 2012

Plant Species Richness And Species Area Relationships In A Florida Sandhill, Monica Ruth Downer

USF Tampa Graduate Theses and Dissertations

Pine sandhill are integral pyrogenic communities in the southeastern United States. Though once widespread, habitat destruction, fire suppression and fragmentation have reduced the population to nearly 3%. It is important to learn as much as possible about these unique areas in order to implement best management practices to conserve and restore the existing populations of these communities.

Fire is central to the maintenance of pine sandhill communities and two conceptual hypothesis regarding burn frequency have come to light in maintaining the unique species composition and richness of these areas. The first is the Intermediate Disturbance Hypothesis which suggests that intermediate …


Assessing The Link Between Coastal Development And The Quality Of Fish Habitat In Mangrove Tidal Tributaries, Justin Micheal Krebs Mar 2012

Assessing The Link Between Coastal Development And The Quality Of Fish Habitat In Mangrove Tidal Tributaries, Justin Micheal Krebs

USF Tampa Graduate Theses and Dissertations

To assess the potential influence of coastal development on the quality of estuarine habitat for nekton, we characterized land use and the intensity of land development surrounding small tidal tributaries of Tampa Bay. Based on this characterization, we classified tributaries as undeveloped, industrial, urban or man-made (i.e., mosquito-control ditches). Over one-third (37%) of tributaries were determined to be heavily developed, while fewer than one-third (28%) remain relatively undeveloped. We then examined the nekton community from eleven tributaries in watersheds representing the defined land-use classes. Whereas mean nekton density and species richness were both independent of land use, nekton-community structure differed …


Herbivory Affects Patterns Of Plant Reproductive Effort And Seed Production, Natalie M. West Feb 2012

Herbivory Affects Patterns Of Plant Reproductive Effort And Seed Production, Natalie M. West

School of Biological Sciences: Dissertations, Theses, and Student Research

Insect herbivory can have a major influence on plant reproduction, and potentially drive selection for strategies that reduce or resist herbivore effects. I used a combination of field experiments and ecological modeling to examine how modifications in the patterns and timing of reproductive investment might ameliorate the consequences of herbivore damage for plant reproduction. I performed experiments to examine how changes in reproductive effort after apical damage and reduction of insect herbivory affected seed production in two thistles native to Nebraska. I then used field data to parameterize a life history model predicting the resource allocation among buds and size …


Two New Species Of Trigonotylus (Hemiptera: Heteroptera: Miridae: Stenodemini) From Western Canada And Northwestern United States, G G.E. Scudder, Michael D. Schwartz Jan 2012

Two New Species Of Trigonotylus (Hemiptera: Heteroptera: Miridae: Stenodemini) From Western Canada And Northwestern United States, G G.E. Scudder, Michael D. Schwartz

Green Canyon Environmental Research Area, Logan Utah

No abstract provided.


The Role Of Fine-Scale Habitat Associations In Structuring Spider Assemblages: Determinants Of Spatial Patterns In Community Compostion, Stephanie M. Cobbold Jan 2012

The Role Of Fine-Scale Habitat Associations In Structuring Spider Assemblages: Determinants Of Spatial Patterns In Community Compostion, Stephanie M. Cobbold

Green Canyon Environmental Research Area, Logan Utah

No abstract provided.


Livestock Foraging Behavior In Response To Sequence And Interactions Among Alkaloids, Tannins, And Saponins, Tiffany L. Jensen Jan 2012

Livestock Foraging Behavior In Response To Sequence And Interactions Among Alkaloids, Tannins, And Saponins, Tiffany L. Jensen

Green Canyon Environmental Research Area, Logan Utah

No abstract provided.


Soil C And N Changes With Afforestation Of Grasslands Across Gradients Of Precipitation And Plantation Age, Sean T. Berthrong, Gervasio Pinero, Esteban G. Jobbagy, Robert B. Jackson Jan 2012

Soil C And N Changes With Afforestation Of Grasslands Across Gradients Of Precipitation And Plantation Age, Sean T. Berthrong, Gervasio Pinero, Esteban G. Jobbagy, Robert B. Jackson

Scholarship and Professional Work - LAS

Afforestation, the conversion of unforested lands to forests, is a tool for sequestering anthropogenic carbon dioxide into plant biomass. However, in addition to altering biomass, afforestation can have substantial effects on soil organic carbon (SOC) pools, some of which have much longer turnover times than plant biomass. An increasing body of evidence suggests that the effect of afforestation on SOC may depend on mean annual precipitation (MAP). The goal of this study was to test how labile and bulk pools of SOC and total soil nitrogen (TN) change with afforestation across a rainfall gradient of 600–1500 mm in the Rio …


Salinity Preference Of Alaskan Threespine Stickleback: Test For Divergence In Halotaxis Between Ancestral And Landlocked Populations, David Fryxell, Eric T. Schultz Jan 2012

Salinity Preference Of Alaskan Threespine Stickleback: Test For Divergence In Halotaxis Between Ancestral And Landlocked Populations, David Fryxell, Eric T. Schultz

EEB Articles

Glacial retreat during the Pleistocene caused landlocking of anadromous Alaskan threespine stickleback, Gasterosteus aculeatus, furnishing a natural ‘experiment’ in osmoregulatory divergence. The objective of this study was to assess the effects of individual acclimation and population divergence on salinity preference. Full-sibling families of marine, anadromous, and freshwater-landlocked populations of stickleback were reared in common environments until 3 weeks post-hatch, then were split and acclimated to low or high salinity. At 6 to 8 weeks of age the six experimental groups were tested for salinity preference in a tank that offers fish a choice of compartments with different salinities arranged …


Ramphotyphlops Braminus (Brahminy Blindsnake): Predation, Louis A. Somma Jan 2012

Ramphotyphlops Braminus (Brahminy Blindsnake): Predation, Louis A. Somma

Papers in Herpetology

Ramphotyphlops braminus currently has the most widespread, near worldwide, nonindigenous distribution of any snake. In Florida, USA, R. braminus is rapidly expanding its distribution.

The stomach contents of a necropsied Dasypus novemcinctus (nine-banded armadillo) found on the premises of the Florida Department of Agriculture and Consumer Services in Gaisesville in March 2011 included an intact adult R. braminus. Dasypus novemcinctus is nonindigenous in Florida. It has a primarily insectivorous diet but occasionally preys upon small vertebrates, including reptiles. This is the first record of R. braminus in the diet of D. novemcinctus.


Leptodactylus Pentadactylus, Miriam M. Heyer, W. R. Heyer, Rafael O. De Sá Jan 2012

Leptodactylus Pentadactylus, Miriam M. Heyer, W. R. Heyer, Rafael O. De Sá

Biology Faculty Publications

Adult Leptodactylus pentadactylus (Figure 1) are large, the head is about as wide as long, and the hind limbs are moderately long (Table 1; Heyer and Thompson [2000] provided definitions of adult size and leg length categories for Leptodactylus). Male vocal sacs are not visible externally or are moderately expanded as a single sac. Sexually active males usually do not have hypertrophied forearms (the largest male examined, 195 mm SVL, has very weakly hypertrophied forearms), only the largest males have a single small to moderate size black spine on each thumb. No males have chest spines. Dorsolateral folds are …


Biology Department Newsletter, No.2, Sacred Heart University Jan 2012

Biology Department Newsletter, No.2, Sacred Heart University

Biology Newsletter

No abstract provided.


Experimental And Molecular Studies Of Bryophyte Dispersal On Alpine Summits, Sean Charles Robinson Jan 2012

Experimental And Molecular Studies Of Bryophyte Dispersal On Alpine Summits, Sean Charles Robinson

Legacy Theses & Dissertations (2009 - 2024)

A review of bryophyte dispersal studies over the last sixty years revealed that the role of gametophytic fragments in bryophyte population dynamics has not been adequately addressed. In this dissertation, I investigate the dispersal potential of wind blown bryophyte fragments on Adirondack alpine summits using a combination of field and laboratory methods.


Experimental Studies Of The Multitrophic Effects Of Anti-Herbivore Defense In Three Pine Barrens Shrub Species, Elizabeth R. Coffey Jan 2012

Experimental Studies Of The Multitrophic Effects Of Anti-Herbivore Defense In Three Pine Barrens Shrub Species, Elizabeth R. Coffey

Legacy Theses & Dissertations (2009 - 2024)

Plants produce numerous chemical defense compounds in response to herbivory. Induced chemical defenses potentially influence soil dynamics by changing the nutrient ratio of the leaves, which are seasonally shed and contribute to the soil, or by affecting the detritus community through leaching of chemicals in the root zone. Manual clipping was utilized in this study to induce chemical compounds with the goal of examining the secondary effects of induced plant defenses. I removed 25% of the mass of fifteen plants of three different plant species in the Albany Pine Bush annually for two consecutive years. I then compared the soil …