Animal Sciences Commons^™

Revision And Taxonomic Position Of Genera Afghanorthochirus Lourenço & Vachon, Baloorthochirus Kovařík, Butheolus Simon, Nanobuthus Pocock, Orthochiroides Kovařík, Pakistanorthochirus Lourenço, And Asian Orthochirus Karsch, With Descriptions Of Twelve New Species (Scorpiones, Buthidae), František Kovařík

Euscorpius

The genera Baloorthochirus, Butheolus, Orthochiroides, Pakistanorthochirus, and Asian species of the genus Orthochirus are revised and keys are presented to all species of Butheolus and Orthochiroides, Asian species of Orthochirus, and to species of all related genera. Afghanorthochirus Lourenço & Vachon, 1997 is synonymized with Orthochirus Karsch, 1892; Nanobuthus Pocock, 1895 is synonymized with Butheolus Simon, 1882; and Pakistanorthochirus Lourenço, 1997 is synonymized with Baloorthochirus Kovařík, 1996. Orthochiroides socotrensis sp. n. from Socotra Island and eleven new species of Orthochirus are described: O. afghanus sp. n., O. heratensis sp. n., O. jalalabadensis …

Antillean Scorpions Deposited At The Montana State University (Arachnida: Scorpiones), Luis F. De Armas

Euscorpius

A total of 71 specimens of Antillean scorpions belonging to three families [Liochelidae, Scorpionidae (Diplocentrinae), and Buthidae] and seven genera are deposited at the Montana Entomology Collection (MTEC) of the Montana State University. The most represented genus is Tityus C. L. Koch, 1836, which contains seven species from Dominican Republic. Those materials are from the following countries: Cuba, Dominican Republic, U.S. Virgin Islands, British Virgin Islands, Montserrat, and St. Kitts. The male of Tityus septentrionalis Armas & Abud Antun, 2004 is described for the first time.

A New Vaejovis From The Mogollon Highlands Of Northern Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey

Euscorpius

A new scorpion species, Vaejovis halli sp. nov., is described. This relatively small, brown new species is found on Mount Ord in the Mazatzal Mountains along the Mogollon Rim of northern Arizona. The new species appears most similar to V. vorhiesi Stahnke and V. deboerae Ayrey. The most distinguishing characteristic of this new species is the number of inner denticles (ID) found on the pedipalp fingers with six on the movable finger and usually five on the fixed finger, which more closely correlates with Vaejovis species from the mountains of southern Arizona rather than those geographically closer …

A Review Of The Genus Heterometrus Ehrenberg, 1828, With Descriptions Of Seven New Species (Scorpiones, Scorpionidae), František Kovařík

Euscorpius

The genus Heterometrus is revised and a key is presented to all species except H. tristis. Subgenera Chersonesometrus Couzijn, 1978, Gigantometrus Couzijn, 1978, Javanimetrus Couzijn, 1981 and Srilankametrus Couzijn, 1981 are synonymized with the subgenus Heterometrus Ehrenberg, 1828. Seven new species of Heterometrus are described: H. beccaloniae sp. n., H. mysorensis sp. n., H. rolciki sp. n., and H. ubicki sp. n. from India, H. nepalensis sp. n., from Nepal, H. sejnai sp. n. from Thailand, and H. cimrmani sp. n. from Thailand and Vietnam. The following species and subspecies are synonymized: H. ( …

A Note On The Scorpions From The Pirin Mountains, Southwestern Bulgaria (Scorpiones: Buthidae, Euscorpiidae), Rolando Teruel, Victor Fet, Luis F. De Armas

Euscorpius

In the present note, four scorpion species are recorded from the Pirin Mountains in southwestern Bulgaria, including the first records for Bulgaria of both Mesobuthus gibbosus (Brullé, 1832) (Buthidae) and the “mingrelicus complex” of the genus Euscorpius Thorell, 1876 (Euscorpiidae) (the latter based upon an undetermined species). Also, E. hadzii Caporiacco, 1950 and another undetermined species of Euscorpius belonging to the “carpathicus complex” are recorded from Pirin. The taxonomic affinities of the taxa are discussed, as well as some biogeographical considerations about the scorpions of this poorly studied mountainous area.

A Scorpion From A Peatbog: The First Arthropod Fossil From The Late Viséan Of The Moscow Coal Basin, Victor Fet, Michael E. Soleglad, Yulia V. Mosseichik, Dmitry E. Shcherbakov

Euscorpius

This note describes a fossil fragment from the Upper Viséan (ca. 330 Ma), of the Moscow Coal Basin (Novgorod Region, Russia). The fossil is identified as the cuticle of a single leg segment (left femur) of a scorpion (Arachnida: Scorpiones), without any further taxonomic placement. The anatomical details of the fragment are given, along with ecological considerations. It is the first fossil scorpion record from Russia.

Alloscorpiops Wongpromi Sp. N. From Thailand And Laos (Scorpiones: Euscorpiidae: Scorpiopinae), František Kovařík, Michael E. Soleglad, Ondřej Košulič

Euscorpius

Alloscorpiops wongpromi sp. n. from Thailand and Laos is described and compared with all other species of the genus Alloscorpiops Vachon, 1980. A key is presented to all species of the genus. A. wongpromi sp. n. is characterized mainly by a higher number of trichobothria on the patella, which has 33–37 external and 21–22 ventral trichobothria, and a dark-colored telson. The subgenus Laoscorpiops Lourenço, 2013 is synonymized with the nominotypical subgenus, because it was based on misinterpretation of the trichobothrial pattern, with two ventral accessory trichobothria on the chela regarded as part of the external Eb series.

Response Of Male Centruroides Vittatus (Scorpiones: Buthidae) To Aerial And Substrate-Borne Chemical Signals, Steffany B. Steinmetz, Karen C. Bost, Douglas D. Gaffin

Euscorpius

Chemical signaling promotes mate location throughout numerous animal taxa. In this study we investigated the possibility that striped scorpions, Centruroides vittatus, use chemical signaling in the form of odor plumes or substrate-borne deposits as communication channels. A Y-shaped arena was constructed to test scorpions’ use of air-borne chemical cues in the detection of potential mates. A second, circular behavioral choice chamber was used to test male scorpions’ responses to female deposits by direct substrate contact. Male scorpions showed no tendency to move toward the female in tests of air-borne chemical transmission but they did demonstrate behavior associated with chemical …

Scorpiofauna Of Kashan (Esfahan Province, Iran) (Arachnida: Scorpiones), Valerio Vignoli, František Kovařík, Pierangelo Crucitti

Euscorpius

This work contains the results of the zoological expedition of the Società Romana di Scienze Naturali to Kashan, in Iran. Four days of intensive field research provided interesting data on the community structure of scorpions of Kashan. Thirty-two scorpion specimens, all belonging to the family Buthidae, are listed and ecological notes are included. Iranobuthus krali Kovařík, 1997, Kraepelinia palpator (Birula, 1903) and Polisius persicus Fet, Capes & Sissom, 2001 were found for the first time in the examined localities. The species collected belong to different ecomorpho-types, characteristic for the particular geographic position of Kashan. We report presence of two species …

A Powerful New Light Source For Ultraviolet Detection Of Scorpions In The Field, Graeme Lowe, Steven R. Kutcher, Doug Edwards

Euscorpius

We describe a new portable solid-state ultraviolet (UV) light that should greatly improve the ease and effectiveness of fluorescent detection of scorpions in the field. The design is based on an array of InGaN light emitting diodes (LEDs). It is in-expensive, lightweight, simple to implement, energetically efficient and mechanically rugged. Scorpion cuticular fluorescence excited by an LED array is 13-fold brighter than excitation by previously available fluorescent tube lamps drawing the same current from the battery.

Mesobuthus Eupeus (C.L. Koch, 1839) (Scorpiones: Buthidae) In Turkey, Ayşegül Karataş, Ahmet Karataş

Euscorpius

Distribution of Mesobuthus eupeus (C.L. Koch, 1839) (Scorpiones: Buthidae) in Turkey is investigated and new distributional records are given. In addition, data on the variation of the number of pectinal teeth, and some ecological features of M. eupeus are provided.

Proceedings Of The 3rd Scorpiology Symposium (American Arachnological Society, 28th Annual Meeting), Victor Fet, Michael E. Soleglad

Euscorpius

Table of Contents

Victor Fet and Douglas D. Gaffin. Preface p. i

Karen C. Bost and Douglas D. Gaffin. Sand scorpion home burrow navigation in the laboratory p. 1

Douglas D. Gaffin and Mark E. Walvoord. Scorpion peg sensilla: are they the same or are they different? p. 7

J. Zachary Porterfield, Douglas D. Gaffin, Caitlin Porterfield and Curtis Johnston. Screening for scorpions: A non-invasive approach to tracking the movements of arachnids in sand p. 17

Tsunemi Yamashita. Surface activity, biomass, and phenology of the striped scorpion, Centruroides vittatus (Scorpiones: Buthidae) in Arkansas, USA p. 25

C. Neal McReynolds. Temporal …

Prey Capture Behavior In The East African Scorpions Parabuthus Leiosoma (Ehrenberg, 1828) And P. Pallidus Pocock, 1895 (Scorpiones: Buthidae), Jan O. Rein

Euscorpius

Prey capture behavior in Parabuthus leiosoma (Ehrenberg, 1828) and P. pallidus Pocock, 1895 was studied in the laboratory. The behavioral components involved in prey capture were identified and an ethogram is presented. The occurrence of the different prey capture components are analyzed and discussed.

A Review Of The Genus Isometrus Ehrenberg, 1828 (Scorpiones: Buthidae) With Descriptions Of Four New Species From Asia And Australia, František Kovařík

Euscorpius

Four new species of the genus Isometrus are described: Isometrus (Reddyanus) bilyi from Australia, I. (R.) petrzelkai from Vietnam and Thailand, I. (R.) problematicus from India and I. (R.) khammamensis from India. Keys are presented to all species of both subgenera of Isometrus. Lectotypes are designated for I. (I.) thurstoni Pocock, 1893 and I. (R.) assamensis Oates, 1888. I. (I.) sankariensis Tikader & Bastawade, 1983 is synonymized with I. (I.) thurstoni Pocock, 1893. First time records include I. (I.) thurstoni Pocock, …

High-Level Systematics And Phylogeny Of The Extant Scorpions (Scorpiones: Orthosterni), Michael E. Soleglad, Victor Fet

Euscorpius

A number of authors (e. g. Birula, 1917a, 1917b; Mello-Leitão, 1945; Stockwell, 1989) addressed above-level systematics of extant scorpions, and accepted the grouping of scorpion families in several superfamilies. At the same time, Kjellesvig-Waering (1986) classified all extant scorpions under the same superfamily, Scorpionoidea. Sissom (1990) and Fet et al. (2000) did not list any superfamilies, considering the systematic situation above family (and often at the family level as well) unresolved. Most recently, Lourenço (2000a) listed six superfamilies, largely following the unpublished but important study of Stockwell (1989). The goal of this paper is to address scorpion systematics and phylogeny …

The Scorpion Sternum: Structure And Phylogeny (Scorpiones: Orthosterni), Michael E. Soleglad, Victor Fet

Euscorpius

The structure of the sternum of all major Recent scorpion groups is analyzed in detail. Based on this analysis, two fundamental sternum types are identified, described and illustrated, type 1 and type 2. These sternum types are distinguished by criteria based on external and internal structural features. The sternum types described herein are offered as a replacement for the various characterizations used throughout the last 140 years which emphasize only gross overall shape and proportions. Phylogenetic and taxonomic ramifications of these new sternal types are discussed. The Carboniferous fossil scorpion Palaeopisthacanthus schucherti Petrunkevitch is assigned to sternum type 1 …

Two New Euscorpius Species From Central-Western Greece (Scorpiones: Euscorpiidae), Gioele Tropea, Victor Fet

Euscorpius

Two new Euscorpius species are described, based on specimens collected by P.M. Giachino & D. Vailati in central-western Greece in neighboring Aitoloakarnania (Western Greece) and Fokida (Central Greece) regional units. No Euscorpius specimens were previously available from this area. The first new species, Euscorpius giachinoi sp. n., is very similar to a recently described E. birulai Fet et al., 2014 from Euboea Island, and is also characterized by a low trichobothrial count (Pv = 7, et = 5), a low pectinal teeth count (Dp = 7 in males, 6 in females), and long-limbed features. The second species, …

Population Level Responses To Direct Application Liming In Gyrinophilus Porphyriticus, Shelby Renea Timm

Theses, Dissertations and Capstones

Direct application liming (DAL) has been used to neutralize acidified streams to restore aquatic biota. This mitigation technique has been used globally for decades, yet little data exist on its effects on amphibian populations. My study investigated the effects of liming on amphibians by measuring variability in life histories of larval Gyrinophilus porphyriticus. I collected larvae from six streams in the Monongahela National Forest, West Virginia. I examined the effects of DAL on age structure, and I failed to detect a treatment effect. I used ANCOVAs to examine differences in body condition, body size, and gape size. I observed that …

Sampling Considerations For Amphibian Surveys: Evaluating Risks Of Committing Type I And Type Ii Errors, Kelli Marie Herrick

Theses, Dissertations and Capstones

Global amphibian declines pose a major threat to the world’s biodiversity. We examined the observation bias associated with volunteer based anuran surveys, such as the North American Amphibian Monitoring Protocol (NAAMP). We followed NAAMP protocol to examine if variation in the persons (1-3) in an observer unit affected observer error. We hypothesized that observation units with multiple observers have less observer bias and would better report anuran assemblages compared to single observers. Larger observer units had fewer incidences of false positive observations. Additionally, we attempted to determine which sampling method for the eastern hellbender (Cryptobranchus a. alleganiensis) had the highest …

Functional Studies Of Ptchd3 During Spermatogensis, Shaimar R. González Morales

Theses, Dissertations and Capstones

Paracrine factor Desert hedgehog (Dhh) is essential for mouse spermatogenesis. However, the specific receptor of Dhh during spermatogenesis is unknown. This study aims to test the hypothesis that Ptchd3, a male germ cell-specific gene acts as a receptor for Dhh in spermatogenesis. In this study, a transgenic mouse model with Ptchd3 gene deletion was first successfully established. Then, in vivo fertility assay and in vitro analysis were performed on Ptchd3 null mutant male mice. The data obtained from the in vivo fertility experiments indicates that there is no statistical significance in offspring litter number (p-value 0.7973) and litter size (p-value …

Development Of Culturing Methods For Native Mayfly Taxa For Use In Laboratory Toxicity Testing, Kevin Nicholas Rowsey

Theses, Dissertations and Capstones

Salinity is increasing in freshwaters throughout the world due to anthropogenic impacts with the implication of the increases to biological communities only now being realized. Declining mayfly populations in Appalachian streams have generated increased interest in elevated dissolved solids in that region and their effect on benthic macroinvertebrates, particularly the sensitive Ephemeroptera taxa. Field and laboratory evaluations have indicated high sensitivity of mayflies to elevated dissolved solids. The research described herein is part of an ongoing effort to provide opportunity for toxicity testing with native mayflies in the laboratory. A successful endeavor would establish mayfly sensitivity to total dissolved solids …

Effects Of Roads And Trails On Peaks Of Otter Salamander (Plethodon Hubrichti) And Eastern Red-Backed Salamander (Plethodon Cinereus) Movement Behavior, Cory Benjamin Goff

Theses, Dissertations and Capstones

Habitat fragmentation negatively affects movement and dispersal of wildlife by altering useable space and creating barriers. The objective of this study was to assess impacts of fragmentation by roads and trails on the movement behavior of two terrestrial, woodland salamander species. I evaluated whether proximity to roads and trails affected the movement behavior of two species: the Peaks of Otter Salamander (Plethodon hubrichti), a species of special concern, and the Eastern Red-backed Salamander (Plethodon cinereus). I examined movement behavior in three treatments: paved roads, gravel roads and hiking trails. I compared treatments to controls located at least 200m from treatment …

Monitoring The Land Cover Change In The Historic Range Of Cambarus Veteranus In West Virginia Using A 1973-2013 Landsat Time Series, Emma Martha Arneson

Theses, Dissertations and Capstones

The crayfish Cambarus veteranus is near extinction in its historic range of the Upper Guyandotte River watershed. The biggest threats to C. veteranus are mining and road construction. Mining has been a continuous activity in the southern coalfields where the crayfish has historically been found, yet little is known about how much land cover change the practice has done to the region. Crayfish act as important organisms within aquatic ecosystems and without them, those systems are often degraded. Quantifying the change in land cover is important to understanding threats to C. veteranus for future protection of the crayfish and its …