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Articles 1 - 30 of 61
Full-Text Articles in Fresh Water Studies
Flight Of The Freshwater Fish, Michael H. Wilson
Flight Of The Freshwater Fish, Michael H. Wilson
Capstones
Michael H. Wilson
Capstone Abstract
December 27, 2016
Flight of the Freshwater Fish
The Hudson River provides for millions of people as a path for commercial and private transportation, a source of food and energy, and perhaps most importantly for many living in the tri-state area as a destination for recreation and relaxation. The most overlooked feature of the river is how the wildlife shows clear signs of a changing climate and rapid environmental response to the impacts of global warming on the river.
Entire populations of fish species in the lower Hudson have been forced to leave the river …
Groundwater-Flow Model Of The Northern High Plains Aquifer In Colorado, Kansas, Nebraska, South Dakota, And Wyoming, Steven M. Peterson, Amanda T. Flynn, Jonathan (Jp) Traylor
Groundwater-Flow Model Of The Northern High Plains Aquifer In Colorado, Kansas, Nebraska, South Dakota, And Wyoming, Steven M. Peterson, Amanda T. Flynn, Jonathan (Jp) Traylor
United States Geological Survey: Water Reports and Publications
The High Plains aquifer is a nationally important water resource underlying about 175,000 square miles in parts of eight states: Colorado, Kansas, Oklahoma, Nebraska, New Mexico, South Dakota, Texas, and Wyoming. Droughts across much of the Northern High Plains from 2001 to 2007 have combined with recent (2004) legislative mandates to elevate concerns regarding future availability of groundwater and the need for additional information to support science-based water-resource management. To address these needs, the United States Geological Survey (USGS) began the High Plains Groundwater Availability Study to provide a tool for water-resource managers and other stakeholders to assess the status …
Umphlett Qci Dec 2016, Natalie A. Umphlett
Umphlett Qci Dec 2016, Natalie A. Umphlett
High Plains Regional Climate Center: Staff Publications
Highlights for the Basin
Temperature and Precipitation Anomalies
Drought Conditions
Large Fires Impact Region
Heavy Rains Bring Flooding to Montana
Isolated Impacts to Agriculture
3-Month Precipitation and Temperature Outlooks
Soil Moisture Conditions
Development Of A Risk Assessment Framework To Predict Invasive Species Establishment For Multiple Taxonomic Groups And Vectors Of Introduction, Alisha D. Davidson, Abigail J. Fusaro, Rochelle A. Sturtevant, Edward S. Rutherford, Donna R. Kashian
Development Of A Risk Assessment Framework To Predict Invasive Species Establishment For Multiple Taxonomic Groups And Vectors Of Introduction, Alisha D. Davidson, Abigail J. Fusaro, Rochelle A. Sturtevant, Edward S. Rutherford, Donna R. Kashian
Biological Sciences Faculty Research Publications
A thorough assessment of aquatic nonindigenous species’ risk facilitates successful monitoring and prevention activities. However, species- and vector-specific information is often limited and difficult to synthesize across a single risk framework. To address this need, we developed an assessment framework capable of estimating the potential for introduction, establishment, and impact by aquatic nonindigenous species from diverse spatial origins and taxonomic classification, in novel environments. Our model builds on previous approaches, while taking on a new perspective for evaluation across species, vectors and stages to overcome the limitations imposed by single species and single vector assessments. We applied this globally-relevant framework …
How To Collect Your Water Sample And Interpret The Results For The Livestock Analytical Package, Bradley J. Austin, Dirk Philipp, Mike Daniels, Brian E. Haggard
How To Collect Your Water Sample And Interpret The Results For The Livestock Analytical Package, Bradley J. Austin, Dirk Philipp, Mike Daniels, Brian E. Haggard
Arkansas Water Resource Center Fact Sheets
A plentiful supply of clean water is crucial for livestock health and productivity. To determine the quality of your livestock’s water resources, periodic sampling and analysis is needed. The Arkansas Water Resources Center (AWRC) in cooperation with the UA Cooperative Extension Service offers several analytical packages to assess the quality of your water resources. This document is intended to provide guidance to livestock owners on collecting water samples for analysis and understanding the results on your report provided by the AWRC’s Water Quality Laboratory (Lab). The information contained within this fact sheet should be used as general guidance, and the …
How To Collect Your Water Sample And Interpret The Results For The Fish Pond Analytical Package, Bradley J. Austin, Amit Sinha, Nathan Stone, W. Reed Green, Mike Daniels, Brian E. Haggard
How To Collect Your Water Sample And Interpret The Results For The Fish Pond Analytical Package, Bradley J. Austin, Amit Sinha, Nathan Stone, W. Reed Green, Mike Daniels, Brian E. Haggard
Arkansas Water Resource Center Fact Sheets
The Arkansas Water Resources Center (AWRC) in cooperation with the Cooperative Extension Service offers several analytical packages to assess the quality of your water resources. This document is intended to provide guidance to aquaculture producers and pond owners on the “Fish Pond Report” provided by the AWRC’s water quality laboratory. The information contained within this fact sheet should be used as general guidance, and the reader is encouraged to seek advice from Extension specialists regarding the interpretation of individual reports and water testing results that may be of concern. The Aquaculture Center for Excellence is at the University of Arkansas …
Identifying Groundwater - Dependent Wetlands Of The Broome Sandstone Aquifer In The La Grange Groundwater Area, Nicholas Wright, Richard J. George Dr, Robert Paul, Paul Raper
Identifying Groundwater - Dependent Wetlands Of The Broome Sandstone Aquifer In The La Grange Groundwater Area, Nicholas Wright, Richard J. George Dr, Robert Paul, Paul Raper
Resource management technical reports
This report identifies wetlands that are likely to be dependent on the Broome Sandstone aquifer within the La Grange groundwater allocation area. The Broome Sandstone aquifer is the dominant groundwater resource in the area. With potential agricultural growth of this area, it is necessary to understand, monitor and manage the aquifer and its dependent wetlands. This report describes how we created a watertable surface that was used to identify wetlands and to determine which of the previously mapped wetlands are likely to be sourced from the Broome Sandstone aquifer. A watertable surface was created from 148 points taken from bore …
2016 Columbia River Flowering Rush Survey, Rich Miller, Mark D. Sytsma
2016 Columbia River Flowering Rush Survey, Rich Miller, Mark D. Sytsma
Center for Lakes and Reservoirs Publications and Presentations
Flowering rush (Butomus umbellatus L.) early detection surveys were conducted in Lake Umatilla during the summer of 2016. Flowering rush is a Class A Noxious Weed in Washington State due to its aggressive nature, ecological and economic impacts, and limited distribution. Dense stands can outcompete native riparian plants, interfere with boating and fishing, and impede irrigation flows. Flowering rush has been reported in eight Washington counties (WSDA 2016) including a large infestation detected during 2008 in the Yakima River, Benton County. Several pioneering populations have since been detected further downstream in Lake Wallula and Lake Umatilla on the Columbia River …
Water Quality Reporting Limits, Method Detection Limits, And Censored Values: What Does It All Mean?, Bradley J. Austin, J. Thad Scott, Mike Daniels, Brian E. Haggard
Water Quality Reporting Limits, Method Detection Limits, And Censored Values: What Does It All Mean?, Bradley J. Austin, J. Thad Scott, Mike Daniels, Brian E. Haggard
Arkansas Water Resource Center Fact Sheets
The Arkansas Water Resources Center (AWRC) maintains a fee-based water-quality lab that is certified by the Arkansas Department of Environmental Quality (ADEQ). The AWRC Water Quality Lab analyzes water samples for a variety of constituents, using standard methods for the analysis of water samples (APHA 2012). The lab generates a report on the analysis, which is provided to clientele, and reports the concentrations or values as measured. Often times the concentrations or values might be very small, even zero as reported by the lab – what does this mean? How should we use this information? This document is intended to …
Database Analysis To Support Nutrient Criteria Development (Phase Ii), B. E. Haggard, M.A. Evans-White, L.B. Massey, E.M. Grantz
Database Analysis To Support Nutrient Criteria Development (Phase Ii), B. E. Haggard, M.A. Evans-White, L.B. Massey, E.M. Grantz
Arkansas Water Resources Center Technical Reports
The intent of this publication of the Arkansas Water Resources Center is to provide a location whereby a final report on water research to a funding agency can be archived. The Texas Commission on Environmental Quality (TCEQ) contracted with University of Arkansas researchers for a multiple year project titled “Database Analysis to Support Nutrient Criteria Development”. This publication covers the second of three phases of that project and has maintained the original format of the report as submitted to TCEQ. This report can be cited either as an AWRC publication (see below) or directly as the final report to TCEQ.
Database Analysis To Support Nutrient Criteria Development (Phase I), B. E. Haggard, J.T. Scott, M.A. Evans-White
Database Analysis To Support Nutrient Criteria Development (Phase I), B. E. Haggard, J.T. Scott, M.A. Evans-White
Arkansas Water Resources Center Technical Reports
The intent of this publication of the Arkansas Water Resources Center is to provide a location whereby a final report on water research to a funding agency can be archived. The Texas Commission on Environmental Quality (TCEQ) contracted with University of Arkansas researchers for a multiple year project titled “Database Analysis to Support Nutrient Criteria Development”. This publication covers the first of three phases of that project and has maintained the original format of the report as submitted to TCEQ. This report can be cited either as an AWRC publication (see below) or directly as the final report to TCEQ.
Beaver Lake Numeric Chlorophyll-A And Secchi Transparency Standards, Phases Ii And Iii: Uncertainty And Trend Analysis, J. Thad Scott, Brian E. Haggard, Zachary Simpson, Matthew Rich
Beaver Lake Numeric Chlorophyll-A And Secchi Transparency Standards, Phases Ii And Iii: Uncertainty And Trend Analysis, J. Thad Scott, Brian E. Haggard, Zachary Simpson, Matthew Rich
Arkansas Water Resources Center Technical Reports
The objective of Phases II and III of this study were to 1) assess the variation in chl‐a and ST across multiple spatial and temporal scales in Beaver Lake in order to validate the assessment method, and 2) quantify trends in chl‐a, ST, and nutrient (total phosphorus and total nitrogen) concentrations in Beaver Lake and the major inflowing rivers to verify any potential water quality impairment.
Database Analysis To Support Nutrient Criteria Development (Phase Iii), J. T. Scott, E.M. Grantz
Database Analysis To Support Nutrient Criteria Development (Phase Iii), J. T. Scott, E.M. Grantz
Arkansas Water Resources Center Technical Reports
The intent of this publication of the Arkansas Water Resources Center is to provide a location whereby a final report on water research to a funding agency can be archived. The Texas Commission on Environmental Quality (TCEQ) contracted with University of Arkansas researchers for a multiple year project titled “Database Analysis to Support Nutrient Criteria Development”. This publication covers the third of three phases of that project and has maintained the original format of the report as submitted to TCEQ. This report can be cited either as an AWRC publication (see below) or directly as the final report to TCEQ.
Droughtscape, Fall 2016, National Drought Mitigation Center
Droughtscape, Fall 2016, National Drought Mitigation Center
DroughtScape: Quarterly Newsletter of the National Drought Mitigation Center (2007–2023)
Contents
NDMC welcome two to team
Third quarter drought summary: Drought still spreading
Third quarter drought impacts: It was a hot, dry summer
Drought management framework for Africa approved
MENA region update
US Virgin Islands one step closer to its own drought monitor
Montana looks to improve watershed resilience
Tournament tackles hazards
Web tool will help officials make drought-related decisions
Dry Horizons launches
Climate Change Impacts On Freshwater Wetland Hydrology And Vegetation Cover Cycling Along A Regional Aridity Gradient, Philip A. Fay, Glenn R. Guntenspergen, Jennifer H. Olker, W. Carter Johnson
Climate Change Impacts On Freshwater Wetland Hydrology And Vegetation Cover Cycling Along A Regional Aridity Gradient, Philip A. Fay, Glenn R. Guntenspergen, Jennifer H. Olker, W. Carter Johnson
Natural Resource Management Faculty Publications
Global mean temperature may increase up to 6°C by the end of this century and together with precipitation change may steepen regional aridity gradients. The hydrology, productivity, and ecosystem services from freshwater wetlands depend on their future water balance. We simulated the hydrology and vegetation dynamics of wetland complexes in the North American Prairie Pothole Region with the WETLANDSCAPE model. Simulations for 63 precipitation × temperature combinations spanning 6°C warming and −20% to +20% annual precipitation change at 19 locations along a mid-continental aridity gradient showed that aridity explained up to 99% of the variation in wetland stage and hydroperiod …
Assessing Aquatic System Metabolism Response To Disturbance, Traci Carmen Hudson
Assessing Aquatic System Metabolism Response To Disturbance, Traci Carmen Hudson
Student Theses and Dissertations
Until recently, agricultural drainage ditches and urban stream systems have been underrepresented within aquatic research studies. This underrepresentation has led to gaps in knowledge within the area of ecosystem structure and function, characteristics of agriculture and urban stream stressors, and management strategies for those streams. Many of the factors (increased nutrients, changes to the quantity and composition of organic carbon inputs, light alterations, and sedimentation) that impair water bodies associated with agricultural drainage and urbanization also influence stream processes. Stream organic matter production and consumption measured as ecosystem metabolism, is a fundamental process comprised of gross primary production (GPP) and …
Water Quality Monitoring At Impaired Sites On The Cache River, Ar, Usa, Molly Elaine Kennon-Lacy
Water Quality Monitoring At Impaired Sites On The Cache River, Ar, Usa, Molly Elaine Kennon-Lacy
Student Theses and Dissertations
Cache River Watershed, AR has several impairments, including: dissolved lead (Pb) and copper (Cu), and total dissolved solids (TDS). Eight sites were monitored monthly measuring: pH, conductivity, DO, total suspended solids (TSS), TDS, turbidity, hardness, dissolved and total nutrients and metals (Pb and Cu), to add to ADEQ’s database. Flow was measured to determine if there was a relationship between measured parameters and periods of low or high flow. Filtration size (0.45 vs 0.20 µm) was compared to conclude if using different size filters on water samples affected concentrations of metals in water. Bioassays were performed using aquatic organisms on …
Umphlett Qci Sept 2016, Natalie Umphlett
Umphlett Qci Sept 2016, Natalie Umphlett
High Plains Regional Climate Center: Staff Publications
Highlights for the Basin
Temperature and Precipitation Anomalies
Streamflow Conditions
Challenging Season for Bird Breeding
Mixed Impacts to Agriculture this Summer
Unprecedented Fish Kill on Yellowstone River
3-Month Precipitation and Temperature Outlooks
Soil Moisture Conditions
Tidal-Fluvial And Estuarine Processes In The Lower Columbia River: Ii. Water Level Models, Floodplain Wetland Inundation, And System Zones, David A. Jay, Amy B. Borde, Heida Diefenderfer
Tidal-Fluvial And Estuarine Processes In The Lower Columbia River: Ii. Water Level Models, Floodplain Wetland Inundation, And System Zones, David A. Jay, Amy B. Borde, Heida Diefenderfer
Civil and Environmental Engineering Faculty Publications and Presentations
Spatially varying water-level regimes are a factor controlling estuarine and tidal-fluvial wetland vegetation patterns. As described in Part I, water levels in the Lower Columbia River and estuary (LCRE) are influenced by tides, river flow, hydropower operations, and coastal processes. In Part II, regression models based on tidal theory are used to quantify the role of these processes in determining water levels in the mainstem river and floodplain wetlands, and to provide 21-year inundation hindcasts. Analyses are conducted at 19 LCRE mainstem channel stations and 23 tidally exposed floodplain wetland stations. Sum exceedance values (SEVs) are used to compare wetland …
Presence Of The Cyanotoxin Microcystin In Arctic Lakes Of Southwestern Greenland, Jessica Trout-Haney, Zachary Wood, Kathryn Cottingham
Presence Of The Cyanotoxin Microcystin In Arctic Lakes Of Southwestern Greenland, Jessica Trout-Haney, Zachary Wood, Kathryn Cottingham
Dartmouth Scholarship
Cyanobacteria and their toxins have received significant attention in eutrophic temperate and tropical systems where conspicuous blooms of certain planktonic taxa release toxins into fresh water, threatening its potability and safe use for recreation. Although toxigenic cyanobacteria are not confined to high nutrient environments, bloom-forming species, or planktonic taxa, these other situations are studied les often studied. For example, toxin production in picoplankton and benthic cyanobacteria—the predominant photoautotrophs found in polar lakes—is poorly understood. We quantified the occurrence of microcystin (MC, a hepatotoxic cyanotoxin) across 18 Arctic lakes in southwestern Greenland. All of the focal lakes contained detectable levels of …
Velocity Profiling, Turbulence, And Chlorophyll Concentrations In The Bottom Boundary Layer Of Lake Michigan Near Muskegon, Michigan, Jonathan M. Benoit, Cary D. Troy, David J. Cannon
Velocity Profiling, Turbulence, And Chlorophyll Concentrations In The Bottom Boundary Layer Of Lake Michigan Near Muskegon, Michigan, Jonathan M. Benoit, Cary D. Troy, David J. Cannon
The Summer Undergraduate Research Fellowship (SURF) Symposium
The characterization of water flow and turbulence near lake beds is important for modelling environmental and ecological effects throughout a lake. In Lake Michigan, where invasive filter-feeding Quagga mussels dominate the lake bed, turbulence plays an important role in determining how much of chlorophyll is mixed down to the Quagga Mussels. Deep in Lake Michigan (44m) near Muskegon, MI, a large tripod was deployed, attached with an Acoustic Doppler Velocimeter, a fluorometer to measure chlorophyll concentrations, and a temperature sensor. Measurements were recorded from late May until early August by sampling velocities every hour in ten-minute bursts at 4 Hz, …
Analyzing Accuracy Of The Lufft Ws600 In Remotely Measuring Precipitation Events, Justin Gay
Analyzing Accuracy Of The Lufft Ws600 In Remotely Measuring Precipitation Events, Justin Gay
STAR Program Research Presentations
The goal of this project was to analyze the accuracy of the Lufft WS600 Weather Sensor in measuring the rate of both liquid and solid precipitation. Measurement accuracy, especially in remote locations, can be difficult to obtain and quantify. Wind, blowing debris, and atmospheric particles can all have the capacity to interfere with instruments that are not being continuously compared to manual observations. Access to quality precipitation data sets are important for both hydrologic and weather forecasting, climate monitoring, and understanding the role of water cycling through ecosystems. Commercially, weather sensors are heavily relied upon by the Federal Aviation Administration …
Optical Water Quality And Human Perceptions Of Rivers, Amie West
Optical Water Quality And Human Perceptions Of Rivers, Amie West
Graduate Theses and Dissertations
Understanding water quality dynamics in recreational rivers is integral in shaping management strategies that maintain ecosystem health, perceived value and appeal, and regional economic significance in a changing environment. Optical water quality describes the behavior of light in water as governed by its physical and chemical composition, and is among the strongest influences on human perceptions of water quality. Ethnohydrology is the study of culturally constructed knowledge and understanding of water. This work is the culmination of an interdisciplinary approach to water resources research—integrating optical water quality and ethnohydrology methods to recognize the intersection between measured water quality and visible …
Chlorine Demand Shows Thresholds And Hierarchy With Source Water Quality At Beaver Lake, Arkansas, Jaime M. Gile
Chlorine Demand Shows Thresholds And Hierarchy With Source Water Quality At Beaver Lake, Arkansas, Jaime M. Gile
Graduate Theses and Dissertations
This study investigated the effects of source water quality in Beaver Lake on the amount of chlorine (Cl) needed to develop decision support system to help guide chlorination practices in pre-treatment of source water. Chlorine demand assays were performed on water samples from Beaver Lake collected from the intake structure at Beaver Water District from March 2014 through August 2015, and using data from these assays, the two points of interest in this study were the Cl dose at which Cl residuals began to accumulate and the mean Cl demand occurring after that dose. Three methods of analysis were used …
Stream Microbial Communities As Potential Indicators Of River And Landscape Disturbance In North-Central Arkansas, Wilson Howard Johnson
Stream Microbial Communities As Potential Indicators Of River And Landscape Disturbance In North-Central Arkansas, Wilson Howard Johnson
Graduate Theses and Dissertations
In the past decade, 29 shale basins have been actively developed across 20 states for extraction of natural gas (NG) via horizontal drilling/hydraulic fracturing (=fracking). This includes ~5000 wells within the Fayetteville shale of north-central Arkansas. Development often impacts both river- and landscapes, and management requires catchment-level evaluations over time, with organismal presence/absence as indicators. For this study next-generation sequencing was used to identify/characterize microbial communities within biofilm of eight Arkansas River tributaries, so as to gauge potential catchment influences. Streams spanned a gradient of landscape features and hydrological flows, with four serving as ‘potentially impacted catchment zones’ (PICZ) and …
Trend Analysis Of Water Quality In Northwest Arkansas Streams Reflects Changes In The Watershed, Zachary Paul Simpson
Trend Analysis Of Water Quality In Northwest Arkansas Streams Reflects Changes In The Watershed, Zachary Paul Simpson
Graduate Theses and Dissertations
Watershed export of nutrients, sediments, and chemicals impacts receiving waters. Changes within the watershed (e.g., anthropogenic or climatic) can alter the transport of constituents in streams. Stream monitoring is crucial for understanding these effects. This study developed a potential improvement to flow-adjusting constituent concentrations in streams, an important step of analyzing monitoring data in lotic systems for trends. The method incorporates a K-fold cross-validation procedure to optimize a model explaining the relationship between the concentration and streamflow, thus providing a valuable tool to researchers in water quality. Additionally, two case studies were conducted on watersheds located in northwest Arkansas using …
Biogeochemistry Of A Saline, Alkaline, Terminal Lake Ecosystem In Transition; Walker Lake, Nevada, Katherine Lynn Willever
Biogeochemistry Of A Saline, Alkaline, Terminal Lake Ecosystem In Transition; Walker Lake, Nevada, Katherine Lynn Willever
UNLV Theses, Dissertations, Professional Papers, and Capstones
Walker Lake is a saline, alkaline, terminal lake ecosystem located in west-central Nevada. For over one hundred years, anthropogenic streamflow diversions within the Walker River Basin have ultimately led to little or no water reaching Walker Lake, the basin’s terminus for water flow. These diversions have resulted in a >46 meter decrease in the lake surface altitude and increases in salinity and dissolved salt constituents that have caused the elimination of native fish species. This study examines how the lack of freshwater inflow has altered the physical, chemical, and microbiological structure of Walker Lake during the lake’s ongoing desiccation.
Between …
Evaluating A Measure-Calculate Method For Determining Sediment Oxygen Demand In Lakes, Adrian Beirise
Evaluating A Measure-Calculate Method For Determining Sediment Oxygen Demand In Lakes, Adrian Beirise
Graduate Theses and Dissertations
A steady-state mass diffusion model used with simple measurable and calculable inputs for determining sediment oxygen demand (SOD) is compared to an intact core incubation (ICI) SOD method using samples from three lakes. The mass diffusion model coupled with inputs is known as the measure-calculate method (M-C) and is a potential alternative to existing methods for measuring SOD which are more complex, time-consuming, and costly. The M-C method requires inputs for volumetric sediment oxygen uptake (Ṅsed), sediment density and porosity, and water properties. Ṅsed was determined by suspending sediment in oxygen-saturated water with a DO probe and determining the steady …
Electrodeionization Versus Electrodialysis: A Clean-Up Of Produced Water In Hydraulic Fracturing, Brigitte Marie Rodgers
Electrodeionization Versus Electrodialysis: A Clean-Up Of Produced Water In Hydraulic Fracturing, Brigitte Marie Rodgers
Graduate Theses and Dissertations
Electrodeionization (EDI) is a widely studied process ranging from applications in wastewater clean-up in the food and beverage industry to purifying organic compounds. To date, there are no apparent studies on applying this technology to produced wastewater recovered from hydraulic fracking sites. Water consumption within hydraulic fracturing sites can reach in the upwards of millions of gallons per site, so a need for a water recycling process becomes necessary within areas where water requirements are scarce. Implementation of an EDI module that is capable of handling high salt solutions from produced wastewater in subsequent fracturing practices will decrease overall water …
Overview Of The Phytoplankton And Water Quality Of Lake Maumelle For 2009, Rich Shaver
Overview Of The Phytoplankton And Water Quality Of Lake Maumelle For 2009, Rich Shaver
Theses and Dissertations
Since 1989 the United States Geological Survey (USGS) and Central Arkansas Water (CAW) have collected water quality and phytoplankton samples from Lake Maumelle in Central Arkansas. These samples were collected from three locations within the reservoir. Phytoplankton samples for 2009 were identified to genus and counted. The phytoplankton did not follow a typical succession pattern for north temperate lakes. Starting in late winter, Chlorophyta tended to be the dominant division followed by Cyanophyta in the fall. Diversity was relatively high at all sampling locations and no differences among sites were detected. Total phosphorus appeared to be the limiting nutrient. Based …