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Articles 151 - 180 of 835
Full-Text Articles in Botany
Phylogeny Of The Australian Solanum Dioicum Group Using Seven Nuclear Genes: Testing Symon’S Fruit And Seed Dispersal Hypotheses., Christopher T. Martine, Ingrid E. Jordon-Thaden, Angela J. Mcdonnell, Jason T. Cantley, Daniel S. Hayes, Morgan D. Roche, Emma S. Frawley, Ian S. Gilman, David C. Tank
Phylogeny Of The Australian Solanum Dioicum Group Using Seven Nuclear Genes: Testing Symon’S Fruit And Seed Dispersal Hypotheses., Christopher T. Martine, Ingrid E. Jordon-Thaden, Angela J. Mcdonnell, Jason T. Cantley, Daniel S. Hayes, Morgan D. Roche, Emma S. Frawley, Ian S. Gilman, David C. Tank
Faculty Journal Articles
The dioecious and andromonoecious Solanum taxa (the “S. dioicum group”) of the Australian Monsoon Tropics have been the subject of phylogenetic and taxonomic study for decades, yet much of their basic biology is still unknown. This is especially true for plant-animal interactions, including the influence of fruit form and calyx morphology on seed dispersal. We combine field/greenhouse observations and specimen-based study with phylogenetic analysis of seven nuclear regions obtained via a microfluidic PCR-based enrichment strategy and high-throughput sequencing, and present the first species-tree hypothesis for the S. dioicum group. Our results suggest that epizoochorous trample burr seed …
Mixed Responses To Woody Encroachment In Icelandic Herbaceous Plant Morphology And Phenology, Cait Battle-Mcdonald
Mixed Responses To Woody Encroachment In Icelandic Herbaceous Plant Morphology And Phenology, Cait Battle-Mcdonald
Independent Study Project (ISP) Collection
Climate change and afforestation efforts are promoting tree growth into traditionally treeless locations in Iceland. This study examined this woody encroachment’s effect on the phenology and morophology of eight plants (Agrostis aequivalvis, Equisetum pratense, Equisetum arvense, Alchemilla alpina, Ranunculus acris, Lupinus nootkatensis, Anthriscus sylvestris, and Taraxacum sp.) in Kjarnaskógur forest, Akureyri, Iceland. An initial transplantation experiment testing trees’ effect on A. aequivalvis, E. pratense, R. acris, and A. sylvestris failed due to plant death. A subsequent observational study examined trees’ effect on E. pratense, A. alpina, R. acris, L. nootkatensis, A. sylvestris, and Taraxacum by observing morphological characteristics (leaf size, …
African Affinity Mask, Giselle M. Strauss
Distribution And Population Structure Of The Invasive Nitellopsis Obtusa (Desv. In Loisel.) J. Groves And Native Species Of Characeae In The Northeast U.S.A., Robin Sleith
Dissertations, Theses, and Capstone Projects
Freshwater ecosystems are some of the most biologically diverse environments on Earth. Billions of humans rely on functioning freshwater ecosystems for drinking water and many other services. These ecosystems are increasingly threatened by human impacts including nutrient pollution, invasive species, and climate change. Here I contribute four research chapters that investigate freshwater diversity and ecosystem threats using the Characeae, a family of freshwater green macroalgae, as a study system. Characeae are a diverse and ancient group with more than 500 extant species and a fossil history spanning at least 250 million years. These algae are macrophytes in freshwater ecosystems, and …
Pollinator-Mediated Yield Across A Heterogeneous Landscape In Eastern South Dakota, Shane Stiles
Pollinator-Mediated Yield Across A Heterogeneous Landscape In Eastern South Dakota, Shane Stiles
Oak Lake Field Station 30th Anniversary Retreat Presentations
No abstract provided.
Floristic Quality Assessments Of Remnant Natural Areas In The Greater Grand Rapids, Michigan Region: Evaluating Botanical Change Since The 1890s, Alan W. Stockdale, Garrett Crow, David P. Warners
Floristic Quality Assessments Of Remnant Natural Areas In The Greater Grand Rapids, Michigan Region: Evaluating Botanical Change Since The 1890s, Alan W. Stockdale, Garrett Crow, David P. Warners
Faculty and Professional Research
In 1901, Emma J. Cole, a beloved high school teacher and distinguished botanist, published Grand Rapids Flora. Cole’s floristic inventory remains the most comprehensive botanical study of the greater Grand Rapids, Michigan region, even though this area has experienced significant land- scape alterations since her publication. To address this knowledge gap, in 2014 we embarked on a multiple-year project aimed to discover and reassess the sites Cole described, as well as to catalog other natural remnant locations within the Grand Rapids area. This paper reports botanical invento- ries that were completed at eight sites within Kent County during 2015 and …
Quantifying Signpost Usage By Captive Male White-Tailed Deer, Cassie L. Auxt, Eric S. Michel, Jonathan A. Jenks
Quantifying Signpost Usage By Captive Male White-Tailed Deer, Cassie L. Auxt, Eric S. Michel, Jonathan A. Jenks
The Prairie Naturalist
White-tailed deer (Odocoileus virginianus) use rubbing of signpost structures to communicate during the breeding season. Rubbing of signpost structures allows deer to communicate via visual and chemical cues, which allows them to establish dominance hierarchies and maintain hierarchal status throughout the breeding season (Moore and Marchinton 1974, Miller et al. 1981, Hewitt 2011). Once a living tree is rubbed, the exposed light-colored sapwood creates a stark contrast in wooded areas, increasing visibility and further enticing deer to investigate the structure (Oehler et al. 1995). Anatomically, the tubular apocrine sudoriferous glands of white-tailed deer are located at the antler …
Island Invasion: The Silent Crisis In Hawaii, Sophia Janssen
Island Invasion: The Silent Crisis In Hawaii, Sophia Janssen
Pomona Senior Theses
Keeping out invasive species may, upon first review, seem like a trivial environmental cry from ecologists and deep environmentalists; a belated wish to return to an undeveloped world where nature was pristine. However invasive species create problems that impact all of us and can have far more severe consequences than changing a stunning landscape. These problems are heightened in islands like Hawaii, where the fragile ecosystems have developed over centuries of evolution and adaptation. The introduction of a disease-carrying mosquito can put the people of Hawaii at risk to many vector-born illnesses and create an epidemic, taking human life. The …
Relative Competitive Abilities, Intersexual Overyielding, And Population Sex Ratio Changes In A Bryophyte, Zachary Stanley
Relative Competitive Abilities, Intersexual Overyielding, And Population Sex Ratio Changes In A Bryophyte, Zachary Stanley
Theses and Dissertations--Biology
Unequal sex ratios are widespread in dioecious plants and understanding their cause is important to understanding fundamental aspects of their population dynamics, and yet what causes biased sex ratios in plants is still poorly understood. Competition experiments have been used in plants to predict the outcome of species interactions, but they have rarely been used to help explain sex ratio bias. This study used a response surface competition design to measure the relative competitive abilities of the sexes of the bryophyte Marchantia inflexa (a thallus liverwort of Marchantiaceae) to predict the outcome of competition before the onset of sexual reproductive …
Three Sister Crops: Understanding American Indian Agricultural Practices Of Corn, Beans And Squash, Sara Colombe, Madhav P. Nepal, Larry B. Browning, Matthew L. Miller, P. Troy White
Three Sister Crops: Understanding American Indian Agricultural Practices Of Corn, Beans And Squash, Sara Colombe, Madhav P. Nepal, Larry B. Browning, Matthew L. Miller, P. Troy White
iLEARN Teaching Resources
American Indians have practiced an inter-planting system to produce corn, beans, and squash, for generations. These crops are known as the “Three Sisters”. In this lesson developed for secondary agriscience curriculum, students will understand the past, current and future production practices of the three important crops. Students will also apply their knowledge to understand the crop selection process and relate to the changing environment.
Review: Canids Of The World: Wolves, Wild Dogs, Foxes, Jackals, Coyotes, And Their Relatives. José R. Castelló., Jonathan (Jon) Way
Review: Canids Of The World: Wolves, Wild Dogs, Foxes, Jackals, Coyotes, And Their Relatives. José R. Castelló., Jonathan (Jon) Way
The Prairie Naturalist
Canids of the World is one of those reads where just when you think that something cannot be beat, this book comes out. What I mean by that is that I have read many works on wolves and other canids, most recently Nate Blakeslee’s American Wolf (Way 2017), and I have enjoyed and been enthralled with many of them. But Canids of the World may take the cake for being the most impressive book available on canids. For a modest price of just under $30.00, you can own a book that has over 600 amazing high-resolution glossy photographs of every …
New Breeding Record And Location For Wilson’S Phalarope (Phalaropus Tricolor) In The Nebraska Great Plains, Usa, Madison O. Sutton, Nico Arcilla
New Breeding Record And Location For Wilson’S Phalarope (Phalaropus Tricolor) In The Nebraska Great Plains, Usa, Madison O. Sutton, Nico Arcilla
The Prairie Naturalist
Wilson’s phalarope (Phalaropus tricolor; Scolopacidae) is a migratory shorebird that relies on interior wetlands for foraging and breeding (Colwell and Jehl 1994, van Gils et al. 2018). Its global population status is unclear (Colwell and Jehl 1994, Lesterhuis and Clay 2010), and is variously listed as declining (Morrison et al. 2006, van Gils et al. 2018), increasing (Andres 2009, BirdLife International 2018), and exhibiting a long-term decline but recent stability (Sauer et al. 2011, Andres et al. 2012). Its global population estimate of 1.5 million birds has not been updated for 30 years, since 1988 (Colwell and Jehl 1994, Lesterhuis …
Review: Behavior Of The Golden Eagle: An Illustrated Ethogram. David H. Ellis (Illustrated By N. John Schmitt)., Jeremy E. Guinn
Review: Behavior Of The Golden Eagle: An Illustrated Ethogram. David H. Ellis (Illustrated By N. John Schmitt)., Jeremy E. Guinn
The Prairie Naturalist
Author David H. Ellis and illustrator N. John Schmitt deliver precisely what is promised in Behavior of the Golden Eagle: An Illustrated Ethogram. This “little volume,” as the author coins it, represents a single source for defining, identifying, and describing behaviors of the Golden Eagle (Aquila chrysaetos). With the inclusion of Schmitt’s exceptional drawings, the book is a piece of art, as well as the most useful manual describing Golden Eagle behaviors. Nearly four decades earlier, Ellis (1979) authored the very first Golden Eagle ethogram—the set of repeated standard behaviors for a species called action patterns—and in his new book, …
Review: Edges And Rushes Of Minnesota: The Complete Guide To Species Identification, Welby R. Smith; Photography By Richard Haug, Edward S. Dekeyser
Review: Edges And Rushes Of Minnesota: The Complete Guide To Species Identification, Welby R. Smith; Photography By Richard Haug, Edward S. Dekeyser
The Prairie Naturalist
Even the most seasoned individual with a plant identification background can relate to the difficulty of identifying sedges and rushes to the species level. Historically, one has had to rely on dichotomous keys to identify a sedge or rush species in the field. After hours of frustration, a person ends up collecting the plant and, if lucky, bringing the collection back to a herbarium where it can be compared to known specimens. I have been collecting and identifying sedge and rush species for over 25 years, and author Welby Smith along with photographer Richard Haug have published what I believe …
Red-Bellied Snake (Storeria Occipitomaculata) Copulation In South Dakota, Drew R. Davis
Red-Bellied Snake (Storeria Occipitomaculata) Copulation In South Dakota, Drew R. Davis
The Prairie Naturalist
Storeria occipitomaculata is a small, terrestrial species of snake that occurs across much of eastern North America (Ernst and Barbour 1989, Ernst 2002, Ernst and Ernst 2003). Due to this widespread distribution, S. occipitomaculata faces varied climates that likely result in regional differences in reproductive phenology. Much of what is known about the reproductive ecology of S. occipitomaculata has been documented from the southeastern portion of its range in North America (South Carolina: Semlitsch and Moran 1984, North Carolina: Willson and Dorcas 2004), though Blanchard (1937) studied a population from northern Michigan. The exact reproductive timing in this …
Review: North American Ducks, Geese & Swans: Identification Guide. Frank S. Todd., Kevin M. Ringelman
Review: North American Ducks, Geese & Swans: Identification Guide. Frank S. Todd., Kevin M. Ringelman
The Prairie Naturalist
The North American Duck, Geese & Swans: Identification Guide, released in 2018, is the culmination of a lifetime of waterfowl photography by the late Frank Todd. Indeed, this book stands apart from other identification guides as being entirely photo-driven, with minimal introductory material, notations of field marks, or descriptions of the various waterfowl species. It is small enough to be carried in the field (6.5” × 9” × 0.5”), but most readers will find it less useful than a standard bird identification book (Sibley 2014), even for waterfowl. As the title suggests, the Todd guide focuses on North American species, …
Preparing An Effective Poster Presentation, Christopher N. Jacques
Preparing An Effective Poster Presentation, Christopher N. Jacques
The Prairie Naturalist
Greetings GPNSS members! By the time you read this editorial, many of have been experiencing the fury unleashed by ‘Old Man Winter’ and may find yourself counting the days until warmer spring temperatures return once again to the Great Plains. Until then, just a couple more months of bitter cold temperatures, strong winds, and blowing and drifting snow. But not to worry, winter also provides the cold weather enthusiasts among us a chance to enjoy a range of outdoor recreational opportunities, a chance to reflect on the previous year in review, exciting professional and personal opportunities ushered in by the …
Comparison Of Northern Flying And Red Squirrel Phylogenies With Focus On The Insular United States, Alyssa M. Kiesow, Hugh B. Britten
Comparison Of Northern Flying And Red Squirrel Phylogenies With Focus On The Insular United States, Alyssa M. Kiesow, Hugh B. Britten
The Prairie Naturalist
Northern flying squirrel (Glaucomys sabrinus) and red squirrel (Tamiasciurus hudsonicus) populations are endemic to northern North America, including the Black Hills. The Black Hills populations are considered disjunct from other populations within their range. We examined insular populations to determine whether arboreal squirrels in the Black Hills each represent a unique population. We trapped and collected ear samples from northern flying and red squirrels in the Black Hills and in areas of Montana, Wyoming, Idaho, Utah, Minnesota, and Wisconsin to infer population phylogenies with special consideration of the Black Hills population. Microsatellite loci and two mtDNA sequences were used for …
Notes: Two-Headed White-Tailed Deer Fetus, William F. Jensen, Daniel M. Grove, Ryan J. Herigstad, William J. Haase
Notes: Two-Headed White-Tailed Deer Fetus, William F. Jensen, Daniel M. Grove, Ryan J. Herigstad, William J. Haase
The Prairie Naturalist
On 6 April 2018 a female white-tailed deer (Odocoileus virginianus dakotensis) was hit and killed by a vehicle along HWY 1806 in rural Morton County, North Dakota, USA (N46o 38.617; W100o 42.901). Based upon dental eruption and wear (Severinghaus 1949), the female was estimated to be a >4.5 years-of-age. A male fetus with parapagus diprosopus (i.e., shared face) apparently was expelled from the adult female, and a domestic canine (Canis lupus familiaris) carried the fetus to a private residence. The resident contacted the North Dakota Game and Fish Department (NDGFD) to turn in the fetus. On 10 April 2018, NDGFD …
Metabolic Gas Emissions From Prairie Soil Containing Foraging Termites, Charles E. Konemann, B.M. Kard, Tom A. Royer, Mark E. Payton
Metabolic Gas Emissions From Prairie Soil Containing Foraging Termites, Charles E. Konemann, B.M. Kard, Tom A. Royer, Mark E. Payton
The Prairie Naturalist
Differences in subterranean termite metabolic gas emissions are readily observed in laboratory experiments. However, in natural field ecosystems a primary difficulty in measuring subterranean termite gases is non-homogeneous distribution of foraging termites in soil. Our field experiment was designed to aggregate foragers of the 'eastern subterranean termite', Reticulitermes flavipes Kollar (EST), in one of four flux chamber configurations placed on a tallgrass prairie throughout 2014 and 2015. We used differently configured flux chambers to measure metabolic gas emissions from soil with or without foraging termites on The Joseph H. Williams Tallgrass Prairie Preserve (TGPP) in north-central Oklahoma. Foraging termitesaggregated in …
The Prairie Naturalist: The Journal Of The Great Plains Natural Science Society Volume 50 No. 2
The Prairie Naturalist: The Journal Of The Great Plains Natural Science Society Volume 50 No. 2
The Prairie Naturalist
TABLE OF CONTENTS
43 EDITOR’S NOTE
RESEARCH ARTICLES
46 Metabolic Gas Emissions from Prairie Soil Containing Foraging Termites Charles E. Konemann, B. M. Kard, Tom A. Royer, and Mark. E. Payton
59 Comparison of northern flying and red squirrel phylogenies with focus on the insular United States Alyssa M. Kiesow, and Hugh B. Britten
NOTES
70 Two-Headed White-Tailed Deer Fetus
72 Red-Bellied Snake (Storeria occipitomacilata) Copulation in South Dakota
74 New Breeding Record and Location for Wilson's Phalarope (Phalaropus tricolor) in the Nebraska Great Plains, USA
BOOK REVIEWS
76 Sedges and Rushes of Minnesota: The Complete Guide to Species Identification. …
A Catalog Of Orchid Species And Their Distribution In Mazumbai Forest Reserve, Tanga Region, Tanzania, Sierra Loomis
A Catalog Of Orchid Species And Their Distribution In Mazumbai Forest Reserve, Tanga Region, Tanzania, Sierra Loomis
Independent Study Project (ISP) Collection
Although orchids are common in Mazumbai Forest Reserve (MFR), there are limited resources on where they grow and how to identify them. The primary objective of this study is to create an updated catalog of orchids in MFR, using species abundance, richness and diversity of the orchids as indicators of where orchids prefer to grow in MFR. Orchid species vary greatly as altitude changes, making it important to survey orchids across different altitudes. Therefore, Mazumbai Forest was split into five different altitudinal bands spanning 1400 – 1900m and six 100 x 20m plots in each band were surveyed for both …
Genetic Diversity In An Invasive Clonal Plant? A Historical And Contemporary Perspective, Elliot D. Weidow
Genetic Diversity In An Invasive Clonal Plant? A Historical And Contemporary Perspective, Elliot D. Weidow
LSU New Orleans Theses and Dissertations
Introduced populations of Eichhornia crassipes (Pontederiaceae) possess extremely low levels of genetic diversity due to severe bottleneck events and clonal reproduction. While populations elsewhere have been well studied, North American populations of E. crassipes remain understudied. We used Amplified Fragment Length Polymorphism markers to assess genetic diversity and population structure in North American E. crassipes populations. Patterns of diversity over the past fifty years were analyzed using herbarium specimens. Furthermore, we sampled populations across the Gulf Coast of the United States throughout a year to determine contemporary genetic diversity and assess potential seasonal effects. Genetic diversity was found to be …
Oaks (Family: Fagaceae) Diversity From Moist Temperate Forests Of Lesser Himalayas, Pakistan, Sher Wali, Kishwar Sultana, Fida Muhammad Khan
Oaks (Family: Fagaceae) Diversity From Moist Temperate Forests Of Lesser Himalayas, Pakistan, Sher Wali, Kishwar Sultana, Fida Muhammad Khan
Journal of Bioresource Management
A detailed survey was conducted in five national parks and game reserves of Azad Jammu and Kashmir, Pakistan to assess the biodiversity of family Fagaceae. Identification of the species was done using literary sources and comparison of samples with herbaria from Quaid-i-Azam University, Islamabad and Pakistan Museum of Natural History, Islamabad, Pakistan. A total of eight different species belonging to two genera were identified, including Quercus spp., Castanea sativa, Quercus glauca, Quercus baloot, Quercus incana, Quercus velutina, Quercus alba and Quercus dilatata. The greatest biodiversity was found in Banjosa Game Reserve (BGR) and Tolipir National Park (TNP), while Pir …
Lekking Behavior Of A Sharp-Tailed Grouse In South-Central Nebraska, Kelsey C. King, Andrew J. Caven, Keith Geluso
Lekking Behavior Of A Sharp-Tailed Grouse In South-Central Nebraska, Kelsey C. King, Andrew J. Caven, Keith Geluso
The Prairie Naturalist
Sharp-tailed grouse (Tympanuchus phasianellus; STGR) occur throughout much of the northern Great Plains, and historically were suspected to have ranged as far south as Kansas (Connelly et al. 1998). Sharp-tailed grouse were the dominant grouse in Nebraska 200 years ago, but early settlers and their associated agricultural practices extirpated STGR from much of their former range, including Kansas and southern Nebraska (Sisson 1976, Johnsgard 2016). During the period of initial land conversion, the greater prairie-chicken (T. cupido; GRPC) filled the void, benefiting from small plots of cropland providing winter forage, and outcompeting the STGR (Johnsgard and Wood 1968, Svedarsky et …
A Short Narrative For Writing For The Prairie Naturalist, Christopher N. Jacques
A Short Narrative For Writing For The Prairie Naturalist, Christopher N. Jacques
The Prairie Naturalist
The abstract should be written upon completion of the paper. A well-written abstract is a single short (≤1 line/page of text), concise, and includes 1) an introductory sentence justifying why the study was conducted, 2) a statement of the principal objectives or hypotheses tested during the study, 3) a brief description of pertinent methods, 4) a summary of significant results, 5) a punchy conclusion, and 6) management implications (i.e., utility of results explaining how, when, where, and by whom data or interpretations can be applied; Krausman and Cox 2017). Keep in mind that abstracts are read more than authors than …
The Prairie Naturalist: The Journal Of The Great Plains Natural Science Society Volume 50 No. 1
The Prairie Naturalist: The Journal Of The Great Plains Natural Science Society Volume 50 No. 1
The Prairie Naturalist
TABLE OF CONTENTS
1 EDITOR’S NOTE
RESEARCH ARTICLES
3 Spatiotemporal Distribution of Waterfowl Disease Outbreaks in Kansas, USA Thomas A. Becker, Adam A. Ahlers, Shane Hesting, and David A. Haukos
14 Wild Turkey Occupancy in a Prairie Landscape Joshua C. Courlas and R. Scott Lutz
24 Age and Growth of Cottonwood Trees along the Missouri River, North Dakota Jonathan M. Friedman, Fisher R. Ankeny, and Marshall Wolf
NOTES
34 Hoary Bat Impaled on Barbed Wire Fence
37 Lekking Behavior of a Sharp-Tailed Grouse in South-Central Nebraska
BOOK REVIEWS
40 Force of Nature: George Fell, Founder of The Natural Areas Movement. …
Age And Growth Of Cottonwood Trees Along The Missouri River, North Dakota, Jonathan M. Friedman, Fisher R. Ankney, J. Marshall Wolf
Age And Growth Of Cottonwood Trees Along The Missouri River, North Dakota, Jonathan M. Friedman, Fisher R. Ankney, J. Marshall Wolf
The Prairie Naturalist
The relict plains cottonwood (Populus deltoides subsp. monilifera) forest along the Missouri River between Lakes Sakakawea and Oahe includes trees as large as two meters in diameter. We cored 24 of these trees to determine their age and suitability for flow reconstruction. Because most of the trees were rotten in the center, we developed a method to estimate the date of the center ring that accounts for the increase in ring width toward the center. Estimated center ring dates were as early as 1806. Cottonwood growth at a dry site was correlated with April–August flow prior to construction of Lake …
Wild Turkey Occupancy In A Prairie Landscape, Joshua C. Courlas, R. Scott Lutz
Wild Turkey Occupancy In A Prairie Landscape, Joshua C. Courlas, R. Scott Lutz
The Prairie Naturalist
We investigated wild turkey (Meleagris gallopavo) distribution in prairie landscapes in North Dakota using occupancy modeling in two stages. In 2012, we extensively surveyed ecoregions across the state and in 2013, intensively surveyed the ecoregion with the highest probability of occupancy. Occupancy models from the statewide survey indicated wild turkeys were sparse in ecoregions with primarily agricultural landscapes, were found associated with wooded riparian cover, and found most frequently in the Missouri River Plateau ecoregion. In the Missouri River Plateau, our occupancy models identified that an additive model including mean patch area of cropland fields and spatial aggregation of forest …
Book Reviews: Force Of Nature: George Fell, Founder Of The Natural Areas Movement. Arthur Melville Pearson., James R. Herkert
Book Reviews: Force Of Nature: George Fell, Founder Of The Natural Areas Movement. Arthur Melville Pearson., James R. Herkert
The Prairie Naturalist
Arthur Melville Pearson’s Force of Nature is a book that tells two very interesting and intertwined stories. One is a story of how perseverance and determination can drive an individual to accomplish great things. And the other is a story of how the modern day natural areas movement came to be. Fortunately for those interested in conservation and natural areas protection, those two stories were combined in the life of George Fell (1916-1994). In Fell’s home state of Illinois, where his impact is widely known, his name is synonymous with conservation. In other places where his impact is less well …