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Articles 151 - 180 of 208
Full-Text Articles in Terrestrial and Aquatic Ecology
The Spatial And Temporal Dimensions Of Plant–Soil Feedback In Native Perennial Monocultures, Bret Lang
The Spatial And Temporal Dimensions Of Plant–Soil Feedback In Native Perennial Monocultures, Bret Lang
Electronic Theses and Dissertations
Plant-soil feedbacks (PSFs) describe how plants change their soil environment in ways that can either negatively or positively affect subsequent plant growth. In doing so, they offer an opportunity to connect ecological processes with agricultural practice. My dissertation examines PSFs created by native perennial monocultures, commonly used for native seed production, across time and space, and through soil legacies. Specifically, I focus on how plant species identity shapes both soil nutrient and soil microbial community responses. To address these questions, I conducted field experiments with monocultures of five forb species native to South Dakota and established plots with these species …
The Nesting Ecology Of Southern Two-Lined Salamanders (Eurycea Cirrigera) In Urban Streams, Leah T. Rittenburg, Emma G. Downing, Todd Pierson
The Nesting Ecology Of Southern Two-Lined Salamanders (Eurycea Cirrigera) In Urban Streams, Leah T. Rittenburg, Emma G. Downing, Todd Pierson
Faculty Articles
Urbanization alters many aspects of stream ecosystems, and its effects upon wildlife will likely increase as more watersheds become developed. This may be particularly true for species with complex life histories, which could be vulnerable in different life stages. Eurycea cirrigera (Green) (Southern Two-lined Salamanders) are common members of many stream communities in the southeastern United States, and they prefer to nest in cobble with little sedimentation—a microhabitat that is threatened by stream degradation. We collected data on the nesting ecology of this amphibian species in urban streams in Atlanta, Georgia, to evaluate microhabitat availability, nest-site selection, and the influence …
Carbon Dynamics Across The Mangrove-Tidal Flat Ecotone In A Tropical Seascape, E. S. Yando, I. J.B. Alemu, L. Kiah Eng, T. M. Sloey, M. Van Breugel, Bhatia N. Friess
Carbon Dynamics Across The Mangrove-Tidal Flat Ecotone In A Tropical Seascape, E. S. Yando, I. J.B. Alemu, L. Kiah Eng, T. M. Sloey, M. Van Breugel, Bhatia N. Friess
Biological Sciences Faculty Publications
Mangroves and other blue carbon ecosystems are important for climate through the storage and sequestration of carbon. Most research estimates carbon dynamics by subsampling central portions of the ecosystem and are then upscales using habitat maps to patch, landscape, or regional levels. However, heterogeneity is rarely accounted for - particularly at edges. Further, adjacent tidal flat ecosystems have only recently been considered for carbon storage, but may be undervalued if areas where they transition with vegetated systems are not accounted for. To address the lack of information at coastal ecotones, we examined both soil carbon storage and carbon dioxide flux …
Using Environmental Dna Metabarcoding To Investigate Diversity And Seasonal Variation Of Fishes In Humboldt Bay, California, Johnathon M. Richardson
Using Environmental Dna Metabarcoding To Investigate Diversity And Seasonal Variation Of Fishes In Humboldt Bay, California, Johnathon M. Richardson
Cal Poly Humboldt theses and projects
This study applied environmental DNA (eDNA) metabarcoding to characterize fish diversity and temporal patterns in fish assemblage structure within Humboldt Bay, California. This work advances ecological understanding of Humboldt Bay’s ichthyofauna, regional natural history and ecosystem function. Over one year (February 2023–January 2024), triplicate surface-water samples were collected every two weeks from a fixed site in Humboldt Bay (n = 78). DNA was extracted from filters and amplified using MiFish 12S primers in six replicate PCRs per sample. Amplicon concentrations were normalized, pooled and sequenced on an Illumina MiSeq platform (2×150 bp). Sequences were processed using a custom bioinformatics pipeline, …
Just Keep Swimming: Outmigration Survival Of Upper Klamath Basin Juvenile Chinook Salmon, James M. Whelan
Just Keep Swimming: Outmigration Survival Of Upper Klamath Basin Juvenile Chinook Salmon, James M. Whelan
Cal Poly Humboldt theses and projects
The removal of four impassable hydroelectric dams on the Klamath River provides salmonids with the opportunity to recolonize their historic range in the upper Klamath River watershed. However, successful reestablishment of salmon populations requires juvenile salmon born in Oregon's upper tributaries to safely migrate downstream through the river's upper reaches. This stretch of the Klamath River, immediately downstream of Upper Klamath Lake, presents multiple migration challenges, including two flow-control dams not scheduled for removal (Link River Dam and Keno Dam), significant water diversions, and water quality issues. To evaluate these challenges, I conducted experimental releases of radio-tagged juvenile Chinook salmon …
Investigating The Mechanisms Of Freeze Tolerance In Lemna Minor, Tamara Bouchard
Investigating The Mechanisms Of Freeze Tolerance In Lemna Minor, Tamara Bouchard
Honors Theses and Capstones
Freezing presents a significant abiotic stress to aquatic plants, yet little is known about the mechanisms by which species like Lemna minor (duckweed) survive subzero temperatures in nature. This study investigates whether L. minor employs freeze-avoidance or freeze-tolerance strategies and explores the potential role of its microbiome in enhancing cold resilience. Using both field-collected fronds from naturally frozen ponds and controlled freezing experiments in the lab, I assessed survival through viability staining and post-thaw survival. Results showed that L. minor can survive natural ice encasement and that prior cold acclimation significantly improves tissue survival under induced freezing. Trypan blue staining …
Journal Of Earth And Life Science, Academic Year 2024-2025, Megan Boche, Nathan D. Pyburn, Ben Hoover, Jayden R. Alsleben, Hector Montoya, Jr, Reed Cumberland, Rose Kuhlmann, Ray Anderson, Ellie Isakson, Seth Sisneros-Martinez, Zanden Mellen, Quady Bernu, Cian Verbridge, Benjamin D. Strei, Madelaine Adamich, Arian Garcia, Tegwen Romportl, Sandra J. Marcu, Bemidji State University
Journal Of Earth And Life Science, Academic Year 2024-2025, Megan Boche, Nathan D. Pyburn, Ben Hoover, Jayden R. Alsleben, Hector Montoya, Jr, Reed Cumberland, Rose Kuhlmann, Ray Anderson, Ellie Isakson, Seth Sisneros-Martinez, Zanden Mellen, Quady Bernu, Cian Verbridge, Benjamin D. Strei, Madelaine Adamich, Arian Garcia, Tegwen Romportl, Sandra J. Marcu, Bemidji State University
Journal of Earth and Life Science
No abstract provided.
Size Structure Of Black Crappie And Bluegill In Northern, Eastern, Western, And Southern Minnesota, Ray Anderson
Size Structure Of Black Crappie And Bluegill In Northern, Eastern, Western, And Southern Minnesota, Ray Anderson
Journal of Earth and Life Science
Fish size structure is an important factor in a fishery population. Panfish such as black crappie Pomoxis nigromaculatus and bluegill Lepomis macrochirus play a major role in the health of fisheries. Five lakes from southern, northern, eastern, and western Minnesota were selected for analysis. Data were collected using standard sampling methods, including gill nets, electrofishing, and fyke nets. Length-frequency distributions were created to assess size structure in each region. Relative abundance was measured using catch per unit effort. Data analysis included a one-way ANOVA to compare Proportional Size Distribution (PSD) values and the distribution of fish sizes among regions. Chi-square …
Comparison Of Hoop Nets To Trap Nets In Sampling Panfish, Jayden R, Alsleben
Comparison Of Hoop Nets To Trap Nets In Sampling Panfish, Jayden R, Alsleben
Journal of Earth and Life Science
Panfish including bluegill Lepomis macrochirus and black crappie Pomoxis nigromaculatus are gamefish species of significance found in Minnesota. Hoop nets and trap nets are two types of gear that can be used to sample these fish. Fisheries managers and anglers alike are interested in the best tactics to capture panfish. The behavior of fish, including bluegill and black crappie, change throughout the sampling season and make it difficult to assume accurate representations. Trap nets have historically been adequate in sampling near shore populations, but questions surrounding the gear have allowed other options to be pursued. The objective of this study …
The Relationship Between Plant Density And Microplastic Concentration In The Water In Lake Bemidji, Mn, Arian Garcia
The Relationship Between Plant Density And Microplastic Concentration In The Water In Lake Bemidji, Mn, Arian Garcia
Journal of Earth and Life Science
Microplastics, plastic particles smaller than 5 mm, are an increasing environmental threat, particularly due to their widespread distribution and toxicity. While much research has focused on larger freshwater and marine ecosystems, the presence and impacts of microplastics in smaller inland lakes remain less understood. This study investigates the relationship between plant density and microplastic concentrations in Lake Bemidji, a smaller freshwater lake. A total of 30 water samples were collected across varying vegetation conditions, ranging from open water (0%) to 90% plant coverage. Microplastics were identified and categorized into six morphological types: fibers, fragments, films, spheres/beads, and foam. The samples …
A Sample Of Fish Communities In Beaver Ponds Of Northern Minnesota, Reed Cumberland, Rose Kuhlmann
A Sample Of Fish Communities In Beaver Ponds Of Northern Minnesota, Reed Cumberland, Rose Kuhlmann
Journal of Earth and Life Science
The North American beaver Castor canadensis is an ecosystem engineer and a keystone species to the environment. Beaver inhabit ponds and construct dams to build ponds that provide habitat for certain fish. Little information is provided on the species of fish that live in beaver ponds. The objective of this study is to sample the fish communities in beaver ponds while comparing the effectiveness of two styles of minnow traps. Twelve beaver ponds were selected in the Bemidji area to be sampled. The quantity and species caught in each style of trap were recorded to compare effectiveness of each trap …
Effects Of Round-Up On The Environment, Sandra J. Marcu
Effects Of Round-Up On The Environment, Sandra J. Marcu
Journal of Earth and Life Science
Many people around the world have used and still currently use Roundup but are unaware of the effects it has on the environment. Roundup is a spray on application weedkiller that is widely used around the world today both residentially and commercially. It enables its user to grow a garden or a field of crops with a no-tilling approach to eliminate weeds. It is a well-known and popular choice for killing weeds that has been around since the mid 1970’s (Oca, 2017). John Franz, a Monsanto scientist discovered that glyphosate (main ingredient in Roundup) was an herbicide or weedkiller, and …
The Effectiveness Of The Minnesota Buffer Law And Its Impacts On People, Ellen P. Isakson
The Effectiveness Of The Minnesota Buffer Law And Its Impacts On People, Ellen P. Isakson
Journal of Earth and Life Science
In January 2015, the then-governor of Minnesota Mark Dayton announced new legislation that would require 50-foot vegetation buffers along all waterways in Minnesota; this legislation would come to be known as the Minnesota buffer law, exact requirements being “perennial vegetative buffers of up to 50 feet along lakes, rivers, and streams and buffers of 16.5 feet along ditches” (Albert, 2017; MN Board of Water and Soil Resources [MN BWSR] 4 ). A buffer in the context of vegetation and natural resource management is a strip of vegetation around a body of water “designed to intercept stormwater runoff and minimize soil …
Does Depth Affect Zebra Mussel Length In Scalp Lake?, Zanden Mellen
Does Depth Affect Zebra Mussel Length In Scalp Lake?, Zanden Mellen
Journal of Earth and Life Science
Since its first sighting in 1996, zebra mussels Dreissena polymorpha have been confirmed in hundreds of Minnesota lakes. Not only were they found in lakes, but they are also found in rivers too. With lack of effective treatments to impede their spread, zebra mussels continue to colonize new areas causing ecological as well as economic harm. The objective of this study was to see if there was a relationship between a zebra mussel length and the depth, they are found using shell length (mm) and depth (cm). The comparison of zebra mussel length and depth was done by performing a …
Age Class Is Not Related To Size Or Composition Of Home Ranges Among Suburban White-Tailed Deer During Parturition, Nathan D. Pyburn
Age Class Is Not Related To Size Or Composition Of Home Ranges Among Suburban White-Tailed Deer During Parturition, Nathan D. Pyburn
Journal of Earth and Life Science
How white-tailed deer use the landscape in an urban environment can help us understand the effects of urbanization on deer populations. White-tailed deer form observable matriarchal hierarchies in studied populations. This study examines the matriarchal structure, home range size, and home range composition of 25 collared white-tailed deer within the city limits of Bemidji, MN. The results found no significant difference between age class and home range size or composition. However, movement metrics from yearling and mature age classes were more similar than the juvenile age class females. I propose this is because social acceptance between mature females was higher …
Relationship Of Utricle And Trap Door Size On Common Bladderworts Utricularia Macrorhiza, Tegwen Romportl
Relationship Of Utricle And Trap Door Size On Common Bladderworts Utricularia Macrorhiza, Tegwen Romportl
Journal of Earth and Life Science
The common bladderwort Utricularia macrorhiza is a carnivorous aquatic plant that captures organisms with its bladders. This study was aimed at setting the stage for future research which might determine if U. macrorhiza could be a biological method of control for invasive species. The objective of this study was to determine the relationship between utricle and trap door sizes and to gather data on the size ranges of these two morphological features. A regression analysis was used to obtain results which showed there was enough evidence to suggest that the size of bladderwort trap doors have a significant, positive relationship …
Investigating The Relationships Between Biotic And Abiotic Factors Affecting Migratory Behavior In Westslope Cutthroat Trout Within The Pend Oreille System, Garrett M. Duncan
Investigating The Relationships Between Biotic And Abiotic Factors Affecting Migratory Behavior In Westslope Cutthroat Trout Within The Pend Oreille System, Garrett M. Duncan
EWU Masters Thesis Collection
Salmonids express several different life histories from migrating to oceans and back or staying in their natal streams as a resident. Migratory individuals are a vital part of the overall success of populations due to the potential for increased genetic diversity and increased fecundity they offer over resident individuals. This research was completed in three basins including Priest River, Pack River, and Lightning Creek which are all connected to the Pend Oreille Basin and are important producers of Westslope Cutthroat Trout (Oncorhynchus clarkii lewisi). Thirty-nine sites were sampled between the three basins. Results were compared between these tributaries that have …
In The Clear: Timber Harvest Informs Elk Summer Nutrition And Resource Selection In A Densely Forested Landscape, Trevor C. Weeks
In The Clear: Timber Harvest Informs Elk Summer Nutrition And Resource Selection In A Densely Forested Landscape, Trevor C. Weeks
Graduate Student Theses, Dissertations, & Professional Papers
As wildlife and habitats face unprecedented challenges in the form of landscape conversion, shifting disturbance regimes, and climate change, it becomes increasingly important for land and wildlife managers to find avenues for collaboration as they seek to maintain habitat to support robust wildlife populations. Timber harvest may provide one such avenue for collaboration. It has been shown to increase the quality and quantity of ungulate nutritional resources by reverting stands to early successional stages. Additionally, timber harvest may allow for a more precise application of such effects, as managers choose where and how to harvest timber. However, the exact effects …
Evaluating Dietary Consequences On Body Mass And Thermal Heat Flux Of North American Bison (Bison Bison) Yearling Heifers, Kendall Marie Wald
Evaluating Dietary Consequences On Body Mass And Thermal Heat Flux Of North American Bison (Bison Bison) Yearling Heifers, Kendall Marie Wald
Electronic Theses and Dissertations
No abstract provided.
Characterizing Nursery Habitat For Juvenile Invasive Carp (Hypophthalmichthys Spp.) On The Ohio River, Rebecca Hiller
Characterizing Nursery Habitat For Juvenile Invasive Carp (Hypophthalmichthys Spp.) On The Ohio River, Rebecca Hiller
Graduate Theses, Dissertations, and Problem Reports (ETD)
Since their introduction to US waters, Silver Carp (Hypophthalmichthys molitrix) and Bighead Carp (H. nobilis) have rapidly spread throughout the Mississippi River basin and its major tributaries, but their invasion has stalled within the Ohio River. Recent research has suggested this slowed spread could be due to a lack of suitable nursery habitat in the Ohio River, affecting recruitment of invasive carp. This study aims to characterize the habitats juvenile carp are utilizing by investigating three main lines of inquiry: that juvenile carp habitat use is limited by (1) the flow regime of the Ohio River …
Presence Of Potential Nursery Habitat For The Lemon Shark In Mississippi Coastal Waters, Lindsay K. Bomgardner, P. O. Grammer, Angie Hoover, Anna Millender, Jeremy Higgs, Jill M. Hendon, Michael J. Andres
Presence Of Potential Nursery Habitat For The Lemon Shark In Mississippi Coastal Waters, Lindsay K. Bomgardner, P. O. Grammer, Angie Hoover, Anna Millender, Jeremy Higgs, Jill M. Hendon, Michael J. Andres
Gulf and Caribbean Research
No abstract provided.
Oyster Horizons: Spatial Prioritization For Oyster Restoration At Alabama Living Shorelines, Trinity Henry-Jack, Aaron Bland, Christa Russell, Ronald Baker
Oyster Horizons: Spatial Prioritization For Oyster Restoration At Alabama Living Shorelines, Trinity Henry-Jack, Aaron Bland, Christa Russell, Ronald Baker
Gulf and Caribbean Research
No abstract provided.
Selective Nutrient Incorporation May Underestimate Heterotrophy Of A Mixotrophic Reef-Building Coral, Connor R. Love, Marleen Stuhr, Michael D. Fox, Veronica Z. Radice, Maoz Fine, Kelton W. Mcmahon, David L. Valentine
Selective Nutrient Incorporation May Underestimate Heterotrophy Of A Mixotrophic Reef-Building Coral, Connor R. Love, Marleen Stuhr, Michael D. Fox, Veronica Z. Radice, Maoz Fine, Kelton W. Mcmahon, David L. Valentine
Biological Sciences Faculty Publications
Mixotrophic reef-building corals acquire nutrition via photosynthate translocation from endosymbiotic microalgae and by heterotrophic prey/particle capture. Heterotrophy promotes resistance to, and recovery from, environmental stress, but quantifying coral heterotrophy remains difficult due to complex resource exchanges within the coral holobiont. We interrogated the response of multiple biomarkers to coral heterotrophy using fatty acid profiling and stable isotope analysis of Stylophora pistillata grown along a controlled feeding gradient from pure autotrophy to pure heterotrophy. We found that fatty acids and nitrogen were effectively incorporated into both coral host and symbiont tissues, while carbon, which is the primary target for conventional heterotrophy …
“Nice To See You”: Addressing Plant Awareness Disparity With Locally Specific Ecological And Cultural Plant Education, Cecilia Muecke
“Nice To See You”: Addressing Plant Awareness Disparity With Locally Specific Ecological And Cultural Plant Education, Cecilia Muecke
Pomona Senior Theses
Plants form the basis of life on Earth. However, due to the phenomenon of plant awareness disparity, a self-perpetuating cycle whereby plants are overlooked on an individual and global level ensues. Lack of plant awareness leads to a lessened emphasis on plants in sustainability and conservation as well as biology education, resulting in negative impacts on biodiversity and food security. The severity of plant awareness disparity is dependent on cultural context, and Indigenous and/or traditional societies are less likely to experience plant awareness disparity because of they are typically more connected to plants for their everyday needs. Native plants are …
Précis Of The Edge Of Sentience: Risk And Precaution In Humans, Other Animals, And Ai, Jonathan Birch
Précis Of The Edge Of Sentience: Risk And Precaution In Humans, Other Animals, And Ai, Jonathan Birch
Animal Sentience
We often face grave practical decisions that seem to hinge on whether a system is sentient. This family of cases includes invertebrate animals, people who are unresponsive after brain injury, fetuses, neural organoids, and now AI technologies. We must decide what to do despite ongoing disagreement about the nature of sentience. In our state of uncertainty, we should pragmatically transform the question from “Is it sentient?” to “Is it a sentience candidate, an investigation priority, or neither?”. When a system is a sentience candidate, it is negligent to fail to consider precautions. We should instead evaluate precautions for their proportionality …
Assemblage Structure Of Large Carnivorous Coral Reef Fishes At Shallow And Mesophotic Depths In Two Of The Largest And Most Isolated Marine Protected Areas In The Philippines, Rene A. Abesamis, Mikaela L. Salvador, Ryan Murray, Kymry Delijero, Segundo F. Conales, Jean T. Utzurrum, Christopher E. Bird, David T. Gauthier
Assemblage Structure Of Large Carnivorous Coral Reef Fishes At Shallow And Mesophotic Depths In Two Of The Largest And Most Isolated Marine Protected Areas In The Philippines, Rene A. Abesamis, Mikaela L. Salvador, Ryan Murray, Kymry Delijero, Segundo F. Conales, Jean T. Utzurrum, Christopher E. Bird, David T. Gauthier
Biological Sciences Faculty Publications
Overfishing remains a threat to coral reef fishes globally, with large carnivores often the most vulnerable. Marine protected areas (MPAs) can restore fish populations and biodiversity, but their impact has been understudied in mesophotic coral ecosystems (MCEs – extensions of shallow reefs from 30 to 150 m). We analyzed baited remote underwater videos recorded in 2016 to investigate the assemblage structure of large carnivorous fishes at shallow (4–12 m) and mesophotic (45–96 m) depths in two of the largest and most isolated MPAs in the Philippines: Tubbataha Reefs Natural Park (TRNP), a fully no-take MPA enacted in 1988; and Cagayancillo, …
Aspen Stands Enrich Bird Communities Following A Wildfire In A Montane, Mixed Conifer Forest, Cody Q. Lane
Aspen Stands Enrich Bird Communities Following A Wildfire In A Montane, Mixed Conifer Forest, Cody Q. Lane
Graduate Student Theses, Dissertations, & Professional Papers
Fire is a major ecological force in most terrestrial systems, but the responses of animal communities to fire vary markedly across ecosystems due to variability in fire regimes and vegetation communities. Quaking aspen (Populus tremuloides) is unique within coniferous forests of western North America for its ability to regenerate following wildfire via vegetative resprouting. As managers employ techniques to restore fire regimes and conserve declining aspen forests, it becomes important to understand how aspen alters postfire communities. Our objectives were to (1) assess how aspen interacted with fire severity and years since fire to influence avian species and …
Spatial And Temporal Patterns Of Nutrient Limitation In A Western Montana River, Matthew T. Nichols
Spatial And Temporal Patterns Of Nutrient Limitation In A Western Montana River, Matthew T. Nichols
Graduate Student Theses, Dissertations, & Professional Papers
Enhanced loading of nitrogen (N) and phosphorus (P) is pervasive in rivers around the world. As a consequence of nutrient-rich conditions, many well-lit rivers experience severe algal blooms, which threaten water quality and aquatic habitats. Nutrient enrichment in the upper Clark Fork River, located in western Montana, has increased the productivity of benthic algae, Cladophora glomerata, which routinely forms extensive blooms during summer growing seasons (July – September). Addressing eutrophication in rivers begins with identifying the nutrients that contribute toward algal productivity. I deployed nutrient diffusing substrates (NDS), collected bi-weekly surface water samples, and conducted algal surveys at four …
Assessing Genetic Diversity And Population Structure In The Imperiled Aconitum Reclinatum (White Monkshood), An Endemic Wildflower Of Central Appalachia, Diamanda A. Zizis
Assessing Genetic Diversity And Population Structure In The Imperiled Aconitum Reclinatum (White Monkshood), An Endemic Wildflower Of Central Appalachia, Diamanda A. Zizis
Master’s Theses
Aconitum reclinatum, white monkshood (Ranunculaceae), is a Pennsylvania critically imperiled plant species found in five states of the Appalachian region of the United States: Pennsylvania, West Virginia, Virginia, North Carolina, and Tennessee. This plant is adapted to cool, moist environments but faces threats from increasing habitat loss, deer browse, logging, and changes in rainfall and temperature associated with climate change. Comprehensive knowledge of A. reclinatum genetic population structure remains limited. To address this, we collaborated with Natural Heritage programs within the range of A. reclinatum to sample populations across the species distribution and generate a genotyping-by-sequencing data set of …
Balancing Timber Harvest And Biodiversity: Impacts Of Selective Logging On Wildlife In The Chiquibul Forest Reserve, Belize, Kilynn Groen
Balancing Timber Harvest And Biodiversity: Impacts Of Selective Logging On Wildlife In The Chiquibul Forest Reserve, Belize, Kilynn Groen
Undergraduate Theses, Professional Papers, and Capstone Artifacts
Selective logging is often promoted as a sustainable forestry practice that maintains biodiversity and ecosystem function, yet its ecological consequences remain under examined, particularly in tropical forests. This study investigates how selective logging affects wildlife communities in the Chiquibul Forest Reserve, Belize. Using remote camera trap data from 22 sites (9 unlogged forest sites and 13 logged (stump) sites), we assessed species richness, trophic structure, and environmental predictors of wildlife abundance. While overall wildlife captures and species richness did not differ significantly between forest and stump habitats, principal component analysis revealed strong community-level separation, with stump habitats supporting distinct species …