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Full-Text Articles in Population Biology

Fall Field Report, August - November 2017, W. Ross Silcock Dec 2017

Fall Field Report, August - November 2017, W. Ross Silcock

Nebraska Bird Review

This fall season will be remembered as the fall with the mega-rarities. There were five, one of which, Pacific Wren, is a first Nebraska record. Based on occurrence dates, Hurricane Irma appears to have influenced the appearance of two of the megas, Black Vulture and Magnificent Frigatebird, neither of which provided opportunities for observers to view them, but both well-documented with photographs. There were an extraordinary TWO sightings of Harris’s Hawk, which together with the presence as of this writing (December 19) of one in northeast Kansas and a view of the species map in eBird strongly suggest a single …


Ogallala Fall Field Days, Sept. 15 - 17, 2017, Janis Paseka Dec 2017

Ogallala Fall Field Days, Sept. 15 - 17, 2017, Janis Paseka

Nebraska Bird Review

The 2017 NOU Fall Field Days took place in and around Ogallala on Sept. 15 - 17 with 55 in attendance. The weather on Friday was hot and windy, making finding birds a challenge. Overnight a cold front came through causing birders to shiver on Saturday and Sunday mornings, but the payoff was easier birding and some interesting species. Field trips were led by William Flack, Clem Klaphake, Wayne Mollhoff, Don Paseka, Ross Silcock and T. J. Walker. Locations included Cedar Point Biological Station, Lake Ogallala SRA, various beaches along the shore of Lake McConaughy SRA, Ash Hollow State Historical …


Evaluating The Ecological Status Of The Introduced Nile Monitor (Varanus Niloticus) In Florida: Forecasting Presence And Population Expansion Using Computational Geographic Information Systems, Noah G. Cohen Nov 2017

Evaluating The Ecological Status Of The Introduced Nile Monitor (Varanus Niloticus) In Florida: Forecasting Presence And Population Expansion Using Computational Geographic Information Systems, Noah G. Cohen

HCNSO Student Theses and Dissertations

The Nile monitor (Varanus niloticus) is a large, carnivorous lizard that has become a notorious invasive species in Florida, USA. Initially released in the 1980s from the pet trade, the species has since established at least three breeding populations and spread throughout much of southern Florida. While current control efforts have failed to eradicate V. niloticus, it is important to attain a better understanding of its invasive dynamics to guide and inform better control strategies. In this study, available georeferenced records of V. niloticus in Florida were compiled and linked to a habitat classification map to evaluate ecotype preferences. Factored …


A Study Of The Urban Red Fox (Vulpes Vulpes) Population In Baton Rouge, Louisiana Using Social Media, Ahsennur Soysal Nov 2017

A Study Of The Urban Red Fox (Vulpes Vulpes) Population In Baton Rouge, Louisiana Using Social Media, Ahsennur Soysal

LSU Master's Theses

Foxes are timid yet resourceful animals that are integrated into many urban environments. Because they are elusive, collecting information about the number of urban foxes, their diet and spatial distribution, their interactions with the ecological community in their urban habitat, as well as residents’ response to them, is difficult. Involving stakeholders to participate in the data collection on wildlife via citizen science on social media is one way to overcome this complication, while simultaneously engaging residents in the ecology happening around them. Therefore, we used social media as the platform to engage the public to document and map the foxes …


The North American Quails, Partridges, And Pheasants, Paul A. Johnsgard Oct 2017

The North American Quails, Partridges, And Pheasants, Paul A. Johnsgard

Zea E-Books Collection

This book documents the biology of six species of New World quails that are native to North America north of Mexico (mountain, scaled, Gambel’s, California, and Montezuma quails, and the northern bobwhite), three introduced Old World partridges (chukar, Himalayan snowcock, and gray partridge), and the introduced common (ring-necked) pheasant. Collectively, quails, partridges, and pheasants range throughout all of the continental United States and the Canadian provinces. Two of the species, the northern bobwhite and ring-necked pheasant, are the most economically important of all North American upland game birds. All of the species are hunted extensively for sport and are highly …


First Report Of A Population Of Western Blacknose Dace (Rhinichthys Obtusus) In The Brushy Creek System Of The Black Warrior River Drainage, Alabama, Eric Bauer, Malorie M. Hayes Sep 2017

First Report Of A Population Of Western Blacknose Dace (Rhinichthys Obtusus) In The Brushy Creek System Of The Black Warrior River Drainage, Alabama, Eric Bauer, Malorie M. Hayes

Southeastern Fishes Council Proceedings

Alabama is home to the southernmost populations of Rhinichthys obtusus, the Western Blacknose Dace. Within Alabama, R. obtusus is found in the Tennessee, Coosa, and Black Warrior River basins, but its presence in the Black Warrior River drainage has been limited. Until now, R. obtusus in the Black Warrior drainage has only been reported as collections of 1 to 4 specimens at a time in the Sipsey Fork drainage. Herein, we report two novel occurrences of R. obtusus in the headwaters of the Brushy Creek system in the Black Warrior River drainage including a singleton and a large population. …


Diet And Habitat Analysis Of Barn Owls (Tyto Alba) Nesting At Ponca State Park, Ann Spilker, Joseph A. Gubanyi Sep 2017

Diet And Habitat Analysis Of Barn Owls (Tyto Alba) Nesting At Ponca State Park, Ann Spilker, Joseph A. Gubanyi

Nebraska Bird Review

In 2015 Barn Owls (Tyto alba) nested on the floor of a grain bin in Ponca State Park, Dixon County. Seven nestlings were observed (Figure 1). It is believed that 6 fledged (one carcass was found on the floor of the bin September 26). Barn Owls were not observed the following spring (Jan Johnson, personal communication). Because there are few records of Barn Owls in the region, we collected data to better understand the Barn Owl nesting in Ponca State Park. Barn Owl prey and their importance to reproductive success has been well documented (Marti et al. 2005; Gubanyi et …


The Nebraska Bird Review, Volume 85 September 2017 Number 3 Sep 2017

The Nebraska Bird Review, Volume 85 September 2017 Number 3

Nebraska Bird Review

Summer Field Report, June - July 2017, by W. Ross Silcock …94

Diet and Habitat Analysis of Barn Owls (Tyto alba) Nesting at Ponca State Park, by Ann Spilker and Joseph Gubanyi … 121

2016 (28th) Report of the NOU Records Committee, by Mark A. Brogie …128

Subscription and Organization Information … 143


Summer Field Report, June - July 2017, W. Ross Silcock Sep 2017

Summer Field Report, June - July 2017, W. Ross Silcock

Nebraska Bird Review

Changes in breeding ranges and numbers of Nebraska breeding birds are the primary focus of a Summer Seasonal Report. Every year there are changes to be noted; this year is no exception. Although most of the phenomena listed below are examples of dispersing individuals without breeding evidence, these often indicate breeding range expansion. White-winged Doves are becoming numerous in southern and western Nebraska, although breeding in far eastern Nebraska was only the 5th report for the state. Chuckwill’s- widow continues to creep westward in the south; the westernmost report to date was of one that crossed the 100-mile gap from …


2016 (28th) Report Of The Nou Records Committee, Mark A. Brogie Sep 2017

2016 (28th) Report Of The Nou Records Committee, Mark A. Brogie

Nebraska Bird Review

The functions and methods of the Nebraska Ornithologists’ Union Records Committee (NOURC) are described in its bylaws (NOURC 2010). The committee’s purpose is to provide a procedure for documenting unusual bird sightings and to establish a list of all documented birds for Nebraska. Accidental and casual species for which the NOURC seeks documentation (NOURC Review List) can be found at the NOU website: www.NOUbirds.org. All records mentioned here are available to interested persons at the NOU archives at the University of Nebraska State Museum (UNSM), Lincoln, NE. Interested parties should contact the current NOU Librarian, whose address can be found …


Habitat Associations With Small Mammal Communities At Wilson's Creek National Battlefield, Stephanie Anne Ellison Aug 2017

Habitat Associations With Small Mammal Communities At Wilson's Creek National Battlefield, Stephanie Anne Ellison

Graduate Theses/Dissertations

The purpose of this study was to aid the National Park Service at Wilson’s Creek National Battlefield with the reported habitat monitoring and management goals through vegetation and wildlife surveys within the park. I provided a description of two major habitat types that are of ecological concern, which included non-native ruderal grasslands and upland deciduous woodlands and forests. I evaluated small mammal communities to determine factors that may affect the detection of individual species and examined habitat associations with occupancy, as small mammals are good indicators of habitat quality. My study highlights the need to manage invasive species such as …


The Nebraska Bird Review, Volume 85 June 2017 Number 2 Jun 2017

The Nebraska Bird Review, Volume 85 June 2017 Number 2

Nebraska Bird Review

Spring Field Report, March - May 2017, by W. Ross Silcock … 54

Greater Sandhill Crane (Antigone canadensis tabida) Copulation Detected Along the Big Bend of the Platte River, South-Central Nebraska, by Andrew J. Caven and Emma M. Brinley Buckley … 83

NOU Spring Field Days and Annual Meeting in Auburn, May 19-21, 2017, by Janis Paseka … 85-90

Subscription and Organization Information … 91


Greater Sandhill Crane (Antigone Canadensis Tabida) Copulation Detected Along The Big Bend Of The Platte River, South-Central Nebraska, Andrew J. Caven, Emma M. Brinley Buckley Jun 2017

Greater Sandhill Crane (Antigone Canadensis Tabida) Copulation Detected Along The Big Bend Of The Platte River, South-Central Nebraska, Andrew J. Caven, Emma M. Brinley Buckley

Nebraska Bird Review

On 9 March 2017 at 0805 hrs, two A. canadensis tabida were observed copulating on a sandy island within the south channel of the Platte River, Hall County, Nebraska (40.790982°N, -98.404635°W, WGS84; 581 m elev.). Two biologists witnessed the copulation at a distance of about 175 m looking to the south from within an overnight viewing blind on the north bank of the south channel of the Platte River, utilizing an 80 mm spotting scope (20x60x magnification). The sequence of events previous to the copulation followed very closely the observations reported by Tacha (1988). The copulation was proceeded by the …


Nou Spring Field Days And Annual Meeting, Auburn, May 19-21, 2017, Janis Paseka Jun 2017

Nou Spring Field Days And Annual Meeting, Auburn, May 19-21, 2017, Janis Paseka

Nebraska Bird Review

The annual meeting of the Nebraska Ornithologists’ Union was held at the Senior Center in Auburn on May 19-21, 2017, and was organized by Robin Harding and Don and Janis Paseka. About 70 birders attended on a weekend that began wet but finished with more birder-friendly conditions. Our Friday evening program was a presentation by Kevin Holliday, Superintendent of Indian Cave State Park. He mentioned the park’s features including the cave and petroglyphs, St. Deroin cemetery, 22 miles of trails, Missouri River overlook and reconstructed town. He also highlighted the many events held in the park during the year, including …


Mitigating The Conflict Between Pitfall-Trap Sampling And Conservation Of Terrestrial Subterranean Communities In Caves, Peter Kozel, Tanja Pipan, Nina Šajna, Slavko Polak, Tone Novak May 2017

Mitigating The Conflict Between Pitfall-Trap Sampling And Conservation Of Terrestrial Subterranean Communities In Caves, Peter Kozel, Tanja Pipan, Nina Šajna, Slavko Polak, Tone Novak

International Journal of Speleology

Subterranean habitats are known for their rich endemic fauna and high vulnerability to disturbance. Many methods and techniques are used to sample the biodiversity of terrestrial invertebrate fauna in caves, among which pitfall trapping remains one of the most frequently used and effective ones. However, this method has turned out to be harmful to subterranean communities if applied inappropriately. Traditionally, pitfall traps have been placed in caves solely on the ground. Here we present an optimized technique of pitfall trapping to achieve a balance between sampling completeness and minimal disturbance of the fauna in the cave. Monthly we placed traps …


Spring Field Report, March-May 2017, W. Ross Silcock May 2017

Spring Field Report, March-May 2017, W. Ross Silcock

Nebraska Bird Review

In general, this was a routine spring, except for remarkable numbers reported of several species, and a notable paucity of most species of wood-warblers. A huge Snow Goose flock in the east was estimated to contain 500,000- 1,000,000 birds; other notable numbers were 2000 Tree Swallows and 5000 Cliff Swallows, both record counts, 44 Black-capped Chickadees, 300 House Wrens, 62 Fox and 76 Song Sparrows on the same day at a single location, 168 Dark-eyed (Slate-colored) Juncos, and 10,000 Common Grackles. Perhaps the most notable phenomenon this spring was the dearth of migrant wood-warblers. Joel Jorgensen constructed graphics based on …


Development Of A Wild Infant Woolly Monkey Social Interactions, Time Allocation And Behavior Of A Wild Lagothrix Lagotricha Poeppigii Infant And Her Mother, Saba Zewdie Apr 2017

Development Of A Wild Infant Woolly Monkey Social Interactions, Time Allocation And Behavior Of A Wild Lagothrix Lagotricha Poeppigii Infant And Her Mother, Saba Zewdie

Independent Study Project (ISP) Collection

The lowland woolly monkey (Lagothrix lagotricha poeppigii), like all species of woolly monkey, is extremely threatened due to anthropologic activities such as deforestation, petroleum drilling and commercial hunting across their habitat. Due to their low reproductive rate and low reproductive success in captivity, the captive population is decreasing every year as habitat loss increases. In late February of 2017 an infant female was born on the island of Sumak Allpa, a monkey rehabilitation organization and reserve in the western Ecuadorian Amazon. The woolly monkeys are completely independent on the reserve; and as such her birth marks the first …


What, Where, Why: A Survey Of Felidae Populations At Enashiva Nature Refuge, Tanzania, Lucrecia Aguilar Apr 2017

What, Where, Why: A Survey Of Felidae Populations At Enashiva Nature Refuge, Tanzania, Lucrecia Aguilar

Independent Study Project (ISP) Collection

As felid populations worldwide continue to deteriorate due to human activities, understanding how felid species utilize various landscapes, along with what factors affect such use or disuse, becomes essential to the preservation of these species. While previous research has examined felid populations around the world, many species and locations remain understudied. This study surveyed felid species at Enashiva Nature Refuge (ENR) in the Serengeti-Mara ecosystem to determine (1) what felid species are present at ENR, (2) where these species generally occur, and (3) why felids reside at ENR in low or high numbers. Through the use of opportunistic camera trapping, …


2016 Christmas Bird Counts, Don Paseka Mar 2017

2016 Christmas Bird Counts, Don Paseka

Nebraska Bird Review

Fourteen Nebraska counts were held during the 2016 CBC season, which is one less than last year. The Harrison and Crawford counts were cancelled this year, but in the opposite corner of the state, a new count was started that includes Indian Cave State Park. An unusually mild November and a seasonable start to December were followed by a modest cold snap during the second week of the month. The Harlan County count was run early in the period on the 15th with ample open water, and it produced a record number of species (90) for that circle. But in …


Annual Color Photo Section 2017, The Nebraska Bird Review Mar 2017

Annual Color Photo Section 2017, The Nebraska Bird Review

Nebraska Bird Review

For our color photo section this year we asked especially for photos of birds in unexpected places. The images on this page and the following two pages show birds in surprising locations or unusual geographic ranges.

32 images

Canyon Wren in wheel of the photographer’s Subaru at Mary Sue Shoemaker’s ranch in central Cherry Co, 20 Nov 2016. 5th record of this species for Nebraska. Photo by Joel Jorgensen.

Great Horned Owl at water feature in the photographer’s yard, Dixon Co., 18 April 2017. Photo by Jan Johnson.

Brambling, 4th record in Nebraska, coming to the feeders at the home …


Winter Field Report, December 2016 To February 2017, W. Ross Silcock Mar 2017

Winter Field Report, December 2016 To February 2017, W. Ross Silcock

Nebraska Bird Review

This was an interesting winter from several standpoints. The trend of unexpected winter occurrences and earlier arrivals in “spring” continued. Mid-winter records of several species were notable: Rock Wren, Marsh Wren, American Pipit, Pine Warbler, and Chipping Sparrow. Rarities (see below) that could be considered in this category as well were Golden-crowned Sparrow and Lesser Goldfinch. For earlier arrivals, there was a noticeable influx 14-18 Feb of various species that provided rather early dates for those species. In this group were Blue-winged Teal, Ruddy Duck, Double-crested Cormorant, American White Pelican, and Turkey Vulture.

Some amazing numbers were reported, including a …


Brambling In Bellevue, Sarpy County, Loren Padelford, Babs Padelford Mar 2017

Brambling In Bellevue, Sarpy County, Loren Padelford, Babs Padelford

Nebraska Bird Review

As we were having breakfast on January 8, 2017, we spotted an unusuallooking bird with House Sparrows in the tree above our deck. After consulting the National Geographic field guide, we determined it was a Brambling. Soon it was feeding on the deck where we were able to get some photos. It was first spotted at 9:25 a.m. and again at 10:30. We put the information on NEBirds and by late morning birders began to arrive.The first group saw the bird at 12:02. By early afternoon we had standing room only in our family room. The bird appeared again at …


Geographic Distribution: Acrantophis Dumerili (Duméril’S Madagascan Ground Boa). Usa. Florida., Louis A. Somma, Kenneth L. Krysko, Laurence L. Conner Jan 2017

Geographic Distribution: Acrantophis Dumerili (Duméril’S Madagascan Ground Boa). Usa. Florida., Louis A. Somma, Kenneth L. Krysko, Laurence L. Conner

Papers in Herpetology

The geographic distribution of Acrantophis dumerili, a nonindigenous introduced species in Florida, with new records, is presented.

Acrantophis dumerili is indigenous to southern and southwestern Madagascar. We provide the first vouchers for Miami-Dade and Bay counties, which were likely escaped or released pets. Florida already has three established species of booid snakes (one boa, two pythons), all introduced through the pet trade pathway. Although there is no clear evidence for an established population of A. dumerili in Florida at this time, the Bay County individuals in a rural area in the Florida panhandle are a particular concern given that two …


Genomic Perspectives On Amphibian Evolution Across Multiple Phylogenetic Scales, Paul Michael Hime Jan 2017

Genomic Perspectives On Amphibian Evolution Across Multiple Phylogenetic Scales, Paul Michael Hime

Theses and Dissertations--Biology

Genomes provide windows into the evolutionary histories of species. The recent accessibility of genome-scale data in non-model organisms and the proliferation of powerful statistical models are now providing unprecedented opportunities to uncover evolutionary relationships and to test hypotheses about the processes that generate and maintain biodiversity. This dissertation work reveals shallow-scale species boundaries and population genetic structure in two imperiled groups of salamanders and demonstrates that the number and information content of genomic regions used in species delimitation exert strong effects on the resulting inferences. Genome scans are employed to test hypotheses about the mechanisms of genetic sex determination in …


A Comparative Analysis Of Geometric Morphometrics Across Two Pseudemys Turtle Species In East Central Virginia, Kristin C. Dillard Jan 2017

A Comparative Analysis Of Geometric Morphometrics Across Two Pseudemys Turtle Species In East Central Virginia, Kristin C. Dillard

Theses and Dissertations

The phylogeny of the turtle genus Pseudemys is poorly understood. In Virginia, many turtles have been found with indicator traits of both eastern river cooters (Pseudemys concinna concinna) and northern red bellied cooters (Pseudemys rubriventris). This study explores morphological evidence for hybridization between the two species across three riverine sites in east central Virginia.

Museum voucher groups for each species were analyzed for relative shell height and plastron length. The shape of the plastral scutes and upper jaw were analyzed using landmark-based morphometric software. These metrics were compared with measurements taken from 188 field-caught Pseudemys specimens. …


Using Citizen Science And Remote Sensing Data To Model Hummingbird Migration, Sarah R. Supp, Laura J. Graham, Frank A. La Sorte, Tina A. Cormier, Gil Bohrer, Donald Powers, Susan Wethington, Kevin Guay, Patrick Jantz, Scott Goetz, Catherine H. Graham Oct 2016

Using Citizen Science And Remote Sensing Data To Model Hummingbird Migration, Sarah R. Supp, Laura J. Graham, Frank A. La Sorte, Tina A. Cormier, Gil Bohrer, Donald Powers, Susan Wethington, Kevin Guay, Patrick Jantz, Scott Goetz, Catherine H. Graham

Annual Symposium on Biomathematics and Ecology Education and Research

No abstract provided.


Harpy Eagle (Harpia Harpyja) Conservation: Juvenile Behavior And Diet At A Nest Near Pijibasal, Darien, Hannah Rodgers Oct 2016

Harpy Eagle (Harpia Harpyja) Conservation: Juvenile Behavior And Diet At A Nest Near Pijibasal, Darien, Hannah Rodgers

Independent Study Project (ISP) Collection

The Harpy Eagle (Harpia harpyja), the largest and most powerful bird of prey in the Americas, is declining throughout its range in Neotropical forests. Hunting and deforestation threaten populations in the most important remaining eagle habitat in Central America, the Darien province of Panama. These eagles may have the longest postfledging juvenile dependency of any raptor, though juvenile behavior during this period is poorly studied. This study monitored an 11-month-old juvenile for 7 days at a nest near the indigenous community of Pijibasal in order to study behavior and diet. Researchers recorded the juvenile’s location and behavior, studied prey remains …


Population Characteristics Of Human-Commensal Rodents Present In Households From Mérida, Yucatán, México, Jesús Alonso Panti-May, Silvia F. Hernández-Betancourt, Marco A. Torres-Castro, Carlos Machaín-Williams, Nohemi Cigarroa-Toledo, Lorenzo Sodá, Gabriela López-Manzanero, Josué R. Meza-Sulú, Victor M. Vidal-Martínez Sep 2016

Population Characteristics Of Human-Commensal Rodents Present In Households From Mérida, Yucatán, México, Jesús Alonso Panti-May, Silvia F. Hernández-Betancourt, Marco A. Torres-Castro, Carlos Machaín-Williams, Nohemi Cigarroa-Toledo, Lorenzo Sodá, Gabriela López-Manzanero, Josué R. Meza-Sulú, Victor M. Vidal-Martínez

MANTER: Journal of Parasite Biodiversity

Anthropocommensal rodents live in close proximity to humans in many habitats around the world. They are a threat to public health because of the pathogens they carry. Recent studies in Mérida, Yucatán, México, have shown that commensal rodents harbor potential zoonotic pathogens such as bacteria, helminths, and viruses. In this study, we describe reproductive and demographic parameters of house mice and black rats present in households from Mérida, Yucatán, México, a municipality located in a tropical region in southern México. Rodents were trapped in 142 households within the municipality of Mérida from 2011 to 2014. A total of 832 rodents …


The Origin And Expansion Of The Eastern Red Fox, Adrienne Egge Kasprowicz May 2016

The Origin And Expansion Of The Eastern Red Fox, Adrienne Egge Kasprowicz

LSU New Orleans Theses and Dissertations

When new populations are first identified in a region there are multiple potential sources: introduction of a non-native species, extra-range expansion of a nearby population, or demographic growth of a previously unnoticed species. Red foxes were absent or rare in the mid-eastern portion United States until the late 1800s. Their origins potentially include natural population increase/expansion, translocations from Europe, and, eventually, 20th century fur farming. In this study I attempt to identify the relative impact of native expansion versus human mediated introductions of both colonial era European foxes and early 20th century fur-farm foxes on the establishment of red …


Molecular Analysis Confirming The Introduction Of Nile Crocodiles, Crocodylus Niloticus Laurenti 1768 (Crocodylidae), In Southern Florida, With An Assessment Of Potential For Establishment, Spread, And Impacts., Michael R. Rochford, Kenneth L. Krysko, Frank J. Mazzotti, Matthew W. Shirley, Mark W. Parry, Joseph A. Wasilewski, Jeffrey S. Beauchamp, Christpher R. Gillette, Edward F. Metzger Iii, Michiko A. Squires, Louis A. Somma Apr 2016

Molecular Analysis Confirming The Introduction Of Nile Crocodiles, Crocodylus Niloticus Laurenti 1768 (Crocodylidae), In Southern Florida, With An Assessment Of Potential For Establishment, Spread, And Impacts., Michael R. Rochford, Kenneth L. Krysko, Frank J. Mazzotti, Matthew W. Shirley, Mark W. Parry, Joseph A. Wasilewski, Jeffrey S. Beauchamp, Christpher R. Gillette, Edward F. Metzger Iii, Michiko A. Squires, Louis A. Somma

Papers in Herpetology

The state of Florida, USA, has more introduced herpetofauna than any other governmental region on Earth. Four species of nonnative crocodilians have been introduced to Florida (all since 1960), one of which is established. Between 2000–2014 we field-collected three nonnative crocodilians in Miami-Dade County, Florida, and one in Hendry County, Florida. We used DNA barcoding and molecular phylogenetics to determine species identification and native range origin. Also, we described diet, movement, and growth for one crocodile. Our molecular analyses illustrated that two of the crocodiles we collected are most closely related to Nile Crocodiles (Crocodylus niloticus) from South Africa, suggesting …