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Articles 1861 - 1890 of 2249

Full-Text Articles in Population Biology

An Assessment Of The Prevalence Of The Bass Tapeworm, Proteocephalus Ambloplitis, And Other Parasites In Five Species Of Basses From River And Lake Ecosystems In West Virginia, J. Alyce Bradbury Jan 1998

An Assessment Of The Prevalence Of The Bass Tapeworm, Proteocephalus Ambloplitis, And Other Parasites In Five Species Of Basses From River And Lake Ecosystems In West Virginia, J. Alyce Bradbury

Theses, Dissertations and Capstones

Two-hundred and fifteen basses were collected from four river sites (Racine Locks and Dam, Robert C. Byrd Locks and Dam; Ashland, KY—Ohio River; London Locks and Dam—Kanawha River) and five reservoir sites (Beech Fork Lake, East Lynn Lake, Summersville Lake, Burnsville Lake, and Stonewall Jackson Lake,). The five species of bass caught were: largemouth bass, Micropterus salmoides; smallmouth bass, Micropterus dolomieui; spotted bass, Micropterus punctulatus; white bass, Morone chrysops; and hybrid bass, Morone chyrsops x Morone saxatilis. Basses were collected by three methods: 1) local anglers (hook and line), 2) gill netting surveys; and 3) …


The Use Of Emergent Rocks As Refugia For The Cheat Mountain Salamander, Plethodon Nettingi Green, Beth Anne Pauley Jan 1998

The Use Of Emergent Rocks As Refugia For The Cheat Mountain Salamander, Plethodon Nettingi Green, Beth Anne Pauley

Theses, Dissertations and Capstones

Plethodon nettinqi was listed as a threatened species in 1989 by the United States Fish and Wildlife Service (Pauley, 1991). Its total range is within 5 counties in eastern West Virginia. There are fewer then 60 disjunct populations known and most populations are above 3,000 ft. and are associated with emergent rocks or narrow ravines with Rhododendron. It is hypothesized the P. netting! survived lumbering practices at the turn of the century by taking refuge beneath large emergent rocks and narrow ravines with Rhododendron. This study examined environmental factors associated with emergent rocks that might regulate the distribution of P. …


Ecological Life History Of Caenis Amica Hagen (Emphemeroptera: Caenidae) From The Mitigated Area Of Green Bottom Wildlife Management Area, Cabell County, West Virginia, Gail Lynn Perrine Jan 1998

Ecological Life History Of Caenis Amica Hagen (Emphemeroptera: Caenidae) From The Mitigated Area Of Green Bottom Wildlife Management Area, Cabell County, West Virginia, Gail Lynn Perrine

Theses, Dissertations and Capstones

The ecological life history of a population of Caenis amica Hagen from the Green Bottom Wildlife Management Area was studied from November 1996 to November 1997. Water quality data and 2,505 larvae were collected from the newly mitigated area of Green Bottom Swamp. Total body lengths of larvae were measured using a computer-digitizing program. These results indicated a bivoltine life cycle. Most larvae were found in the spring and summer months of April, June, and July. The highest peak of larvae (1,035) occurred in July. The pH values ranged from 7.0 to 8.0 with a mean of 7.6. Temperatures ranged …


Chondrocranial Anatomy And Skeletogenesis In Dendrobates Auratus, Rafael O. De Sá, Shannon Hill Jan 1998

Chondrocranial Anatomy And Skeletogenesis In Dendrobates Auratus, Rafael O. De Sá, Shannon Hill

Biology Faculty Publications

The larval chondrocranium and visceral skeleton of Dendrobates auratus ls described and compared with those of other dendrobatids. Four characters, i.e. lack of fusion between orbital cartilages and otic capsules, wide processus muscularis palatoquadrati, lack of processus pseudopterygoideus, and lack of processus anterolateralis hyalis, represent derived conditions for Dendrobates within Dendrobatidae. Cranial and postcranial ossification sequences are reported for D. auratus and Epipedobates tricolor. Skeletogenesis is earlier in E. tricolor, but the overall pattern of ossification is similar in the two species.


Habitat Selection, Reproductive Success, And Impaling Patterns Of A Migratory Population Of Loggerhead Shrikes, Jeffrey D. Esely Jr. Jan 1998

Habitat Selection, Reproductive Success, And Impaling Patterns Of A Migratory Population Of Loggerhead Shrikes, Jeffrey D. Esely Jr.

Masters Theses

I studied habitat selection and reproductive success of a migratory population of Loggerhead Shrikes (Lanius ludovicianus migrans) as well as the temporal and spatial patterns of impaling exhibited by this population during the nesting season. The selection of breeding habitat was investigated at different spatial scales. Thirty-eight (67%) of 57 nests were located in osage orange (Maclura pomifera) trees. Shrike territories (area ≤ 200 m from nest site) contained significantly more pasture, fenceline, and perch sites, but less com, alfalfa, and forest than random sites. At larger spatial scales, more grassland was detected within 300 m …


Draft: Plan For The Management Of The Houtman Abrolhos Islands Fish Habitat Protection Area (Schedule 1), Abrolhos Islands Management Advisory Council, Fisheries Department Of Western Australia Dec 1997

Draft: Plan For The Management Of The Houtman Abrolhos Islands Fish Habitat Protection Area (Schedule 1), Abrolhos Islands Management Advisory Council, Fisheries Department Of Western Australia

Fisheries Management Papers

This document is a separate draft plan which has been prepared to meet the requirements of Section 117 of the Fish Resources Management Act 1994. It contains a description of the area; outlines the purposes for which the area is to be set aside; and lists the recommendations contained in Fisheries Management Paper No. 104, Management of the Houtman Abrolhos System, which relate to the management of the proposed Fish Habitat Protection Area.


Management Of The Houtman Abrolhos System, Abrolhos Islands Management Advisory Council, Fisheries Department Of Western Australia Dec 1997

Management Of The Houtman Abrolhos System, Abrolhos Islands Management Advisory Council, Fisheries Department Of Western Australia

Fisheries Management Papers

In November 1995 Cabinet resolved that the Minister for Fisheries would establish the Abrolhos Islands Management Advisory Committee (AIMAC) pursuant to Section 42 of the Fish Resources Management Act 1994. The role of AIMAC is to advise the Minister for Fisheries about the management of the Abrolhos Islands and the adjoining State Territorial Waters, as detailed in Section 2.2.3.1 of this report. The AIMAC has overseen the preparation of this report on behalf of the Minister for Fisheries. This draft report contains a brief description of the Abrolhos System and its special values. It also states goals and objectives for …


Management Options For Pilbara Demersal Line Fishing, Gaye Looby Nov 1997

Management Options For Pilbara Demersal Line Fishing, Gaye Looby

Fisheries Management Papers

Fisheries Western Australia licensing arrangements allow all vessels with an unrestircted fishing boat licence ('wetline licence') to take scalefish throughout the State. These arrangements have led to a high level of excess capacity in the licensed 'wetline' fishing fleet to exploit the scalefish resources. The declaration of the Kimberley Interim Line Fishery and forthcoming declaration the Northern Demersal Scalefish Fishery have resolved this problem for the Kimberley waters. However, the high level of excss capacity in the 'wetline' fleet to exploit the waters of the Pilbara remains a problem Preview access and develop a management plan for the Pilbara demersal …


Abalone Aquaculture In Western Australia. Discussion Paper And Draft Policy Guidelines., Cameron Westaway, Jeff Norriss Oct 1997

Abalone Aquaculture In Western Australia. Discussion Paper And Draft Policy Guidelines., Cameron Westaway, Jeff Norriss

Fisheries Management Papers

The purposes of this discussion paper is to highlight relevant issues associated with abalone aquaculture development and outline a set of draft policy guidelines. The paper is being disseminated to members of the aquaculture industry, commercial and recreational fishing bodies and the broader community to allow comment.


Discussion Paper: Future Management Of The Aquatic Charter Industry In Western Australia, Tour Operators Fishing Working Group Sep 1997

Discussion Paper: Future Management Of The Aquatic Charter Industry In Western Australia, Tour Operators Fishing Working Group

Fisheries Management Papers

The Tour Operators Fishing Working Group (TOFWG) was established by the Hon Monty House MLA, Minister for Fisheries, in recognition of the growing importance of this element of the tourism industry and in recognition of the need to ensure that the development of this industry occurs in an ecologically sustainable framework across the State. This discussion paper is the result of extensive consultation with the aquatic charter operators' industry by the TOFWG. Proposals outlined here for community discussion are intended to establish a management framework within which the aquatic charter operators’ industry may continue to develop, and which will ensure …


Draft: Optimising The Worth Of The Lobster Catch: Options And Issues, Marec Pty Ltd Sep 1997

Draft: Optimising The Worth Of The Lobster Catch: Options And Issues, Marec Pty Ltd

Fisheries Management Papers

The purpose of this document is to contribute to informed discussion on alternative measures which might be adopted to optimise the worth of the lobster catch. In doing so, it provides a number of possible initiatives which could be more closely examined for adoption. Details of issues are provided which should be taken into consideration when assessing the various proposals. The industry as it is today is examined and assessed, and its achievements analysed. The possible initiatives proposed stem from this research. As a consequence of this document, the rock lobster industry could undertake to proceed with a detailed examination …


Marine Farm Planning And Consultation Processes In Western Australia, Dave Everall Aug 1997

Marine Farm Planning And Consultation Processes In Western Australia, Dave Everall

Fisheries Management Papers

The development of a sustainable aquaculture industry in Western Australia is considered to be a high priority by the Government. The marine farming industry has a bright future in Western Australia, but there is a need to balance its developing needs for suitable waters with those of existing activities, and with community expectations for accountability and transparency in the resolution of equity and resource sharing decisions. The purpose of this report is to review the existing planning and consultation processes for pearling and aquaculture in WA and recommend how they may be improved and incorporated into a single process. It …


The Annual Reproductive Cycle Of Oryzomys Palustris In A Virginia Tidal Marsh, Erin A. Dreelin Jul 1997

The Annual Reproductive Cycle Of Oryzomys Palustris In A Virginia Tidal Marsh, Erin A. Dreelin

Biological Sciences Theses & Dissertations

The objectives of my study were to determine when the marsh rice rat (Oryzomys palustris) begins and ends reproduction during the year, if there are seasonal changes in litter size, and the weight at which sexual maturity is reached. Monthly samples of rice rats were trapped from May 1995 to May 1996 in Northampton County, Virginia. Rice rats were trapped with Fitch live traps, euthanized in the field, and brought to the lab for necropsy. Data from two other studies of rice rats in Northampton County were also examined. Potential breeders did not reproduce with equal intensity throughout the …


Ecological Relationships Between The Marsh Rice Rat (Oryzomys Palustris) And The Meadow Vole (Microtus Pennsylvanicus) In Two Virginia Tidal Marshes, Christopher P. Bloch Jul 1997

Ecological Relationships Between The Marsh Rice Rat (Oryzomys Palustris) And The Meadow Vole (Microtus Pennsylvanicus) In Two Virginia Tidal Marshes, Christopher P. Bloch

Biological Sciences Theses & Dissertations

Small mammals were trapped for 23 months on two live trap grids in tidal marshes in Northampton County, on the Eastern Shore of Virginia, to examine ecological relationships between the marsh rice rat (Oryzomys palustris) and the meadow vole (Microtus pennsylvanicus). Population density, survival rates, capture probability, recruitment, residence time, and home range size were calculated for each species. Patterns of reproductive activity and fluctuations in mean weight were also observed.

There were considerable differences in population dynamics between the two sites. At Grid 2, where vegetative cover was more abundant, population densities of rice rats and meadow voles …


The Aquaculture Of Non-Endemic Species In Western Australia, Tina Thorne Jun 1997

The Aquaculture Of Non-Endemic Species In Western Australia, Tina Thorne

Fisheries Management Papers

This Management Paper has been drafted in liaison with the Department of Environmental Protection in accordance with the principles of risk assessment and taking into account translocation issues of non-endemic species such as redclaw (Cheras quadricarninatus). It is considered that the implementation of this policy will protect the natural environment and native crayfish species. It will also allow for the development of a redclaw industry, and provide the first step towards self regulation of that industry.


Management Arrangements For The Western Rock Lobster Fishery For The 1997/98 Season, F. B. Prokop May 1997

Management Arrangements For The Western Rock Lobster Fishery For The 1997/98 Season, F. B. Prokop

Fisheries Management Papers

This paper deals with the proposed management strategies for the 1997/98 seasons. The purpose of this paper is to present the views of RLIAC on management arrangements for the 1997/98 rock lobster season and to indicate the strategic directions that long term management of this important fishery might take so as to ensure both its biological sustainability and the optimisation of product value for both the State and the participants in the fishery.


Management Of The South Coast Purse Seine Fishery, J Fowler, K Donohue, R.C. J. Lenanton, M. Moran, D Gaughan Mar 1997

Management Of The South Coast Purse Seine Fishery, J Fowler, K Donohue, R.C. J. Lenanton, M. Moran, D Gaughan

Fisheries Management Papers

This paper has presented a strategy for moving the management of the small pelagics fishery off the south coast to management of the whole fishery, which facilitates a more biologically sound and economically stable fishery.


A Pricing Policy For Fisheries Agencies, Management Committee Of Scfa, Standing Committee Of Fisheries And Aquaculture (Sfca), Department Of Primary Industries And Energy Mar 1997

A Pricing Policy For Fisheries Agencies, Management Committee Of Scfa, Standing Committee Of Fisheries And Aquaculture (Sfca), Department Of Primary Industries And Energy

Fisheries Management Papers

The intent of this paper is to inform members of Standing Committee for Fisheries and Aquaculture (SCFA) and the members of the Ministerial Council for Fisheries, Forestry and Aquaculture of the methods of fee setting that are currently applied by fisheries agencies in Australia, and alternative pricing policies for fisheries agencies.


Natural History Of The Red-Spotted Newt, Notophthalmus Viridescens Viridescens (Rafinesque), In West Virginia, Jennifer M. Piascik Jan 1997

Natural History Of The Red-Spotted Newt, Notophthalmus Viridescens Viridescens (Rafinesque), In West Virginia, Jennifer M. Piascik

Theses, Dissertations and Capstones

Many studies have been conducted on Notophthalmus viridescens viridescens but none have occurred in West Virginia. There are variations in the life history of newts depending on location. In this study, I studied a population of newts in Wayne County, West Virginia from February 1996 to June 1997. I examined population structure, seasonal activity, and reproductive biology.

The population structure can be divided into four classes: egg, larvae, eft, and adult. Eggs at Shoals were deposited in mid-May and larvae were present from mid-June to early September. Larvae began to transform at an approximate total length of 32.8 mm. Red …


Observations On Reproduction, Isozyme Patterns, And Meristics In Mosquitofish (Pisces: Poeciliidae) From West Virginia, Matt Mcreynolds Jan 1997

Observations On Reproduction, Isozyme Patterns, And Meristics In Mosquitofish (Pisces: Poeciliidae) From West Virginia, Matt Mcreynolds

Theses, Dissertations and Capstones

Prior to 1988, two subspecies of the mosquitofish Gambusia affinis (G. a. affinis and G. a. holbrooki) were recognized along the Gulf and Atlantic Coasts of the United States. Based on electrophoretic analysis, subsequent investigators recognized G. holbrooki and G. affinis as separate species. Populations in drainages west of Mobile Bay were considered G. affinis, and those east of this divide being G. holbrooki. The only mosquitofish population in West Virginia is found in the Meadow River wetlands (1392 ha). This wetland complex is located at the western end of Greenbrier County in the Allegheny Mountain Province. …


Diet And Growth Of Larval And Juvenile Grass Pickerel, Esox Americanus Vermiculatus, And Central Mudminnow, Umbra Limi, In The Green Bottom Wildlife Management Area, Cabell County, West Virginia, Erica Midkiff Jan 1997

Diet And Growth Of Larval And Juvenile Grass Pickerel, Esox Americanus Vermiculatus, And Central Mudminnow, Umbra Limi, In The Green Bottom Wildlife Management Area, Cabell County, West Virginia, Erica Midkiff

Theses, Dissertations and Capstones

The grass pickerel, Esox americanus vermiculatus, is listed as Undetermined on the Vertebrate Species List of Concern in West Virginia. The central mudminnow, Umbra limi, is a disjunct population in Green Bottom Swamp. The lentic, vegetated areas that are required for spawning for the grass pickerel and the central mudminnow have been reduced by residential, agricultural, and industrial developments. Green Bottom Swamp, a naturally occurring wetland of 58 ha, and a nearby mitigated wetland of 29 ha, provides spawning habitat for the grass pickerel and the mudminnow. Fishes were collected in beds of Potamogetan crispus and Ceratophyllum demursum …


Habitat Use By Migrant Whooping Cranes In Saskatchewan, Brian W. Johns, Eric J. Woodsworth, Ed A. Driver Jan 1997

Habitat Use By Migrant Whooping Cranes In Saskatchewan, Brian W. Johns, Eric J. Woodsworth, Ed A. Driver

Proceedings of the North American Crane Workshop

We investigated habitat use by migrating whooping cranes (Grus americana) in Saskatchewan between 1986 and 1990. At foraging and roosting sites and a sample of randomly selected sites we measured various habitat parameters. Palustrine and lacustrine wetlands were the most frequently used wetlands. Temporary and seasonal wetlands were the primary roost habitat during spring migration, and semi-permanent and permanent wetlands were frequently used during the fall. Cranes were attracted to areas of higher than average wetland density. Land use within 2 km of roost sites was intensively modified by man. Cereal crops were the most used feeding areas. …


Availability Of Stopover Habitat For Migrant Whooping Cranes In Nebraska, Dale W. Stahlecker Jan 1997

Availability Of Stopover Habitat For Migrant Whooping Cranes In Nebraska, Dale W. Stahlecker

Proceedings of the North American Crane Workshop

Four stratified random samples of 512 National Wetland Inventory (NWI) maps within the central Nebraska portion of the Wood Buffalo-Aransas whooping crane (Grus americana) migration corridor were used to evaluate the availability of wetland roost sites. Wetlands were eliminated as potential roosts if visibility was obscured by vegetation or slope, if certain human activities occurred within 100-800 ro, or if water < 30 em deep was not available. Seasonal emergent wetlands, available as roosts primarily in spring, dominated all samples, particularly in the nortb. Sixty-five percent of all wetlands >0.04 ha passed map review and 52 % passed when ground-truthed. NWI map review was a good predictor of both suitability (63 % correct) and unsuitability (73 % correct). More than one-half of all open and emergent …


Woody Vegetation Expansion And Continuing Declines In Open Channel Habitat On The Platte River In Nebraska, Paul J. Currier Jan 1997

Woody Vegetation Expansion And Continuing Declines In Open Channel Habitat On The Platte River In Nebraska, Paul J. Currier

Proceedings of the North American Crane Workshop

Aerial videography in 1988 and 1994 of the same selected segments of the Big Bend reach of the Platte River in central Nebraska was compared. During this 6-year period, channel area declined 4 to 41 % (median 26%). In river segments where the channel had already narrowed significantly and at sites where channel habitat is actively managed declines were less (17-18%). These substantial changes occurred during a time that included a relatively low flow period (1990-92). Flows during the summer gertttination and establishment period (mid-May through August) for cottonwood (Populus deltoides) and willow (Salix spp.), the …


Home Range And Habitat Use By Isolation-Reared Sandhill Cranes, Wenrui Duan, Theodore A. Bookhouts, Richard P. Urbanek Jan 1997

Home Range And Habitat Use By Isolation-Reared Sandhill Cranes, Wenrui Duan, Theodore A. Bookhouts, Richard P. Urbanek

Proceedings of the North American Crane Workshop

We isolation-reared, placed radio transmitters on, and released 38 greater sandhill cranes (Grus canadensis tabida) on Seney National Wildlife Refuge (Seney) in 1988-90 to develop procedures for initiating new populations of cranes. Here we report on habitat selection by breeding birds with functional radio transmitters. Home ranges established by 6 cranes in 1992, when they were 2 to 4 years old, averaged 199±50.8 (SE) ha (harmonic mean method, 75 % utilization) and were of 2 types: feeding grounds separated from nesting habitat and feeding grounds adjacent to nesting habitat. Home ranges consisted of 36% emergent palustrine wetlands, 28% …


Recent Advances In Scheduling Strategies And Practical Techniques In Crane Artificial Insemination, Scott R. Swengel, Monica L. Tuite Jan 1997

Recent Advances In Scheduling Strategies And Practical Techniques In Crane Artificial Insemination, Scott R. Swengel, Monica L. Tuite

Proceedings of the North American Crane Workshop

We analyzed fertility patterns of 339 eggs laid 1985-93 at the International Crane Foundation by cranes whose sole means of fertility was artificial insemination (AI). Ten of 15 crane species were studied. Fertility rates increased significantly (P < 0.05) with (I) higher sperm density, (2) greater sperm motility, and (3) 2 vs. 1 vs. 0 inseminations 4-7 days before oviposition. Semen sample size was inversely related to sperm density and had no strong relationship with egg fertility. Inseminations less than 2.5-3.0 days before an egg is laid are probably too late to contribute to fertilizing the egg but may contribute substantially to fertilizing the next egg. We also describe practical knowledge gained during the past 10 years that apply to certain species or situations. Wattled cranes (Bugeranus carunculatus) have a long fertile period post-AI (up to 16 days), but may require more careful methods and special AI scheduling strategies. We also describe techniques for AI just after oviposition that are especially beneficial for Siberian cranes (Grus leucogeranus), and methods for maximizing sample delivery to the female.


Sandhill Crane Hunts In The Rio Grande Valley And Southwest New Mexico, C. Gregory Schmitt, Barry Hale Jan 1997

Sandhill Crane Hunts In The Rio Grande Valley And Southwest New Mexico, C. Gregory Schmitt, Barry Hale

Proceedings of the North American Crane Workshop

Annual limited permit hunting seasons for sandhill cranes (Grus canadensis) were initiated in southwestern New Mexico (SWNM) in 1982 and in the middle Rio Grande Valley (MRGV) in 1986. Within the MRGV and SWNM are wintering areas for 3 migratory subspecies, including the lesser sandhill crane (G. c. canadensis) (lessers). Canadian sandhill crane (G. c. rowani) (Canadians), and greater sandhill crane (G. c. tabida) (greaters). We summarized annual season dates, bag limits, requirements for permits, procedures for examination of crane specimens at check stations, number of hunters, estimated harvest, mean harvest per …


Results Of An Experiment To Lead Cranes On Migration Behind Motorized Ground Vehicles, David H. Ellis, Brian Clauss, Tsuyoshi Watanabe, R. Curt Mykut, Matthew Kinloch, Catherine H. Ellis Jan 1997

Results Of An Experiment To Lead Cranes On Migration Behind Motorized Ground Vehicles, David H. Ellis, Brian Clauss, Tsuyoshi Watanabe, R. Curt Mykut, Matthew Kinloch, Catherine H. Ellis

Proceedings of the North American Crane Workshop

Ten greater sandhill cranes (Grus canadensis tabida) , trained to enter and ride in a specially-equipped truck, were transported at ca 80 days of age from their rearing site at Patuxent Wildlife Research Center (patuxent), Maryland, to a reintroduction site located within the species' former breeding range in northern Arizona. After 5 additional weeks of training, these juvenile cranes were led south ca 600 km to a wintering area on the Arizona/Mexico border. Nine of the 10 survived the trek, 495 km of which were flown, although only a few cranes flew every stage of the route. Their …


Corrigendum Crane Migration In Northern New Mexico, Dale W. Stahlecker Jan 1997

Corrigendum Crane Migration In Northern New Mexico, Dale W. Stahlecker

Proceedings of the North American Crane Workshop

These tables were not included in the above article. They are published here so that interested readers will have access to them while reading the results and discussion presented in that paper. It is now well documented that sandhill cranes (Grus canadensis) from outside the Rocky Mountain Population stage in spring and autumn in the San Luis Valley of southern Colorado (Benning et al. 1997). Therefore, the high autumn counts reported in the 1988 paper and discussed in relation to Table 3 could also have included cranes from other populations.


Survival And Habitat Use Of Greater Sandhill Crane Colts On Modoc National Wildlife Refuge, California, Kevin J. Desroberts Jan 1997

Survival And Habitat Use Of Greater Sandhill Crane Colts On Modoc National Wildlife Refuge, California, Kevin J. Desroberts

Proceedings of the North American Crane Workshop

Radiotelemetry was used to monitor 13 (1990) and 14 (1992) greater sandhill crane (Grus canadensis tabida) colts on Modoc National Wildlife Refuge, California, to determine causes of mortality, brood habitat utilization, and effects of habitat management on colt survival. Colt survival rates were 0.36 (1990) and 0.54 (1992). Coyotes (Canis latrans) killed 3 colts and mink (Mustela vison) killed 4; I colt died because of a bacterial infection (Staphylococcus aureous). Broods used 6 different habitat types and 79% used more than I type. Irrigated meadows (74%), cultivated uplands (53 %), and marsh …