Evolution Commons^™

Revision And Taxonomic Position Of Genera Afghanorthochirus Lourenço & Vachon, Baloorthochirus Kovařík, Butheolus Simon, Nanobuthus Pocock, Orthochiroides Kovařík, Pakistanorthochirus Lourenço, And Asian Orthochirus Karsch, With Descriptions Of Twelve New Species (Scorpiones, Buthidae), František Kovařík

Euscorpius

The genera Baloorthochirus, Butheolus, Orthochiroides, Pakistanorthochirus, and Asian species of the genus Orthochirus are revised and keys are presented to all species of Butheolus and Orthochiroides, Asian species of Orthochirus, and to species of all related genera. Afghanorthochirus Lourenço & Vachon, 1997 is synonymized with Orthochirus Karsch, 1892; Nanobuthus Pocock, 1895 is synonymized with Butheolus Simon, 1882; and Pakistanorthochirus Lourenço, 1997 is synonymized with Baloorthochirus Kovařík, 1996. Orthochiroides socotrensis sp. n. from Socotra Island and eleven new species of Orthochirus are described: O. afghanus sp. n., O. heratensis sp. n., O. jalalabadensis …

Antillean Scorpions Deposited At The Montana State University (Arachnida: Scorpiones), Luis F. De Armas

Euscorpius

A total of 71 specimens of Antillean scorpions belonging to three families [Liochelidae, Scorpionidae (Diplocentrinae), and Buthidae] and seven genera are deposited at the Montana Entomology Collection (MTEC) of the Montana State University. The most represented genus is Tityus C. L. Koch, 1836, which contains seven species from Dominican Republic. Those materials are from the following countries: Cuba, Dominican Republic, U.S. Virgin Islands, British Virgin Islands, Montserrat, and St. Kitts. The male of Tityus septentrionalis Armas & Abud Antun, 2004 is described for the first time.

A New Vaejovis From The Mogollon Highlands Of Northern Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey

Euscorpius

A new scorpion species, Vaejovis halli sp. nov., is described. This relatively small, brown new species is found on Mount Ord in the Mazatzal Mountains along the Mogollon Rim of northern Arizona. The new species appears most similar to V. vorhiesi Stahnke and V. deboerae Ayrey. The most distinguishing characteristic of this new species is the number of inner denticles (ID) found on the pedipalp fingers with six on the movable finger and usually five on the fixed finger, which more closely correlates with Vaejovis species from the mountains of southern Arizona rather than those geographically closer …

A Review Of The Genus Heterometrus Ehrenberg, 1828, With Descriptions Of Seven New Species (Scorpiones, Scorpionidae), František Kovařík

Euscorpius

The genus Heterometrus is revised and a key is presented to all species except H. tristis. Subgenera Chersonesometrus Couzijn, 1978, Gigantometrus Couzijn, 1978, Javanimetrus Couzijn, 1981 and Srilankametrus Couzijn, 1981 are synonymized with the subgenus Heterometrus Ehrenberg, 1828. Seven new species of Heterometrus are described: H. beccaloniae sp. n., H. mysorensis sp. n., H. rolciki sp. n., and H. ubicki sp. n. from India, H. nepalensis sp. n., from Nepal, H. sejnai sp. n. from Thailand, and H. cimrmani sp. n. from Thailand and Vietnam. The following species and subspecies are synonymized: H. ( …

A Note On The Scorpions From The Pirin Mountains, Southwestern Bulgaria (Scorpiones: Buthidae, Euscorpiidae), Rolando Teruel, Victor Fet, Luis F. De Armas

Euscorpius

In the present note, four scorpion species are recorded from the Pirin Mountains in southwestern Bulgaria, including the first records for Bulgaria of both Mesobuthus gibbosus (Brullé, 1832) (Buthidae) and the “mingrelicus complex” of the genus Euscorpius Thorell, 1876 (Euscorpiidae) (the latter based upon an undetermined species). Also, E. hadzii Caporiacco, 1950 and another undetermined species of Euscorpius belonging to the “carpathicus complex” are recorded from Pirin. The taxonomic affinities of the taxa are discussed, as well as some biogeographical considerations about the scorpions of this poorly studied mountainous area.

A Scorpion From A Peatbog: The First Arthropod Fossil From The Late Viséan Of The Moscow Coal Basin, Victor Fet, Michael E. Soleglad, Yulia V. Mosseichik, Dmitry E. Shcherbakov

Euscorpius

This note describes a fossil fragment from the Upper Viséan (ca. 330 Ma), of the Moscow Coal Basin (Novgorod Region, Russia). The fossil is identified as the cuticle of a single leg segment (left femur) of a scorpion (Arachnida: Scorpiones), without any further taxonomic placement. The anatomical details of the fragment are given, along with ecological considerations. It is the first fossil scorpion record from Russia.

Alloscorpiops Wongpromi Sp. N. From Thailand And Laos (Scorpiones: Euscorpiidae: Scorpiopinae), František Kovařík, Michael E. Soleglad, Ondřej Košulič

Euscorpius

Alloscorpiops wongpromi sp. n. from Thailand and Laos is described and compared with all other species of the genus Alloscorpiops Vachon, 1980. A key is presented to all species of the genus. A. wongpromi sp. n. is characterized mainly by a higher number of trichobothria on the patella, which has 33–37 external and 21–22 ventral trichobothria, and a dark-colored telson. The subgenus Laoscorpiops Lourenço, 2013 is synonymized with the nominotypical subgenus, because it was based on misinterpretation of the trichobothrial pattern, with two ventral accessory trichobothria on the chela regarded as part of the external Eb series.

Response Of Male Centruroides Vittatus (Scorpiones: Buthidae) To Aerial And Substrate-Borne Chemical Signals, Steffany B. Steinmetz, Karen C. Bost, Douglas D. Gaffin

Euscorpius

Chemical signaling promotes mate location throughout numerous animal taxa. In this study we investigated the possibility that striped scorpions, Centruroides vittatus, use chemical signaling in the form of odor plumes or substrate-borne deposits as communication channels. A Y-shaped arena was constructed to test scorpions’ use of air-borne chemical cues in the detection of potential mates. A second, circular behavioral choice chamber was used to test male scorpions’ responses to female deposits by direct substrate contact. Male scorpions showed no tendency to move toward the female in tests of air-borne chemical transmission but they did demonstrate behavior associated with chemical …

Scorpiofauna Of Kashan (Esfahan Province, Iran) (Arachnida: Scorpiones), Valerio Vignoli, František Kovařík, Pierangelo Crucitti

Euscorpius

This work contains the results of the zoological expedition of the Società Romana di Scienze Naturali to Kashan, in Iran. Four days of intensive field research provided interesting data on the community structure of scorpions of Kashan. Thirty-two scorpion specimens, all belonging to the family Buthidae, are listed and ecological notes are included. Iranobuthus krali Kovařík, 1997, Kraepelinia palpator (Birula, 1903) and Polisius persicus Fet, Capes & Sissom, 2001 were found for the first time in the examined localities. The species collected belong to different ecomorpho-types, characteristic for the particular geographic position of Kashan. We report presence of two species …

A Powerful New Light Source For Ultraviolet Detection Of Scorpions In The Field, Graeme Lowe, Steven R. Kutcher, Doug Edwards

Euscorpius

We describe a new portable solid-state ultraviolet (UV) light that should greatly improve the ease and effectiveness of fluorescent detection of scorpions in the field. The design is based on an array of InGaN light emitting diodes (LEDs). It is in-expensive, lightweight, simple to implement, energetically efficient and mechanically rugged. Scorpion cuticular fluorescence excited by an LED array is 13-fold brighter than excitation by previously available fluorescent tube lamps drawing the same current from the battery.

Mesobuthus Eupeus (C.L. Koch, 1839) (Scorpiones: Buthidae) In Turkey, Ayşegül Karataş, Ahmet Karataş

Euscorpius

Distribution of Mesobuthus eupeus (C.L. Koch, 1839) (Scorpiones: Buthidae) in Turkey is investigated and new distributional records are given. In addition, data on the variation of the number of pectinal teeth, and some ecological features of M. eupeus are provided.

Proceedings Of The 3rd Scorpiology Symposium (American Arachnological Society, 28th Annual Meeting), Victor Fet, Michael E. Soleglad

Euscorpius

Table of Contents

Victor Fet and Douglas D. Gaffin. Preface p. i

Karen C. Bost and Douglas D. Gaffin. Sand scorpion home burrow navigation in the laboratory p. 1

Douglas D. Gaffin and Mark E. Walvoord. Scorpion peg sensilla: are they the same or are they different? p. 7

J. Zachary Porterfield, Douglas D. Gaffin, Caitlin Porterfield and Curtis Johnston. Screening for scorpions: A non-invasive approach to tracking the movements of arachnids in sand p. 17

Tsunemi Yamashita. Surface activity, biomass, and phenology of the striped scorpion, Centruroides vittatus (Scorpiones: Buthidae) in Arkansas, USA p. 25

C. Neal McReynolds. Temporal …

Prey Capture Behavior In The East African Scorpions Parabuthus Leiosoma (Ehrenberg, 1828) And P. Pallidus Pocock, 1895 (Scorpiones: Buthidae), Jan O. Rein

Euscorpius

Prey capture behavior in Parabuthus leiosoma (Ehrenberg, 1828) and P. pallidus Pocock, 1895 was studied in the laboratory. The behavioral components involved in prey capture were identified and an ethogram is presented. The occurrence of the different prey capture components are analyzed and discussed.

A Review Of The Genus Isometrus Ehrenberg, 1828 (Scorpiones: Buthidae) With Descriptions Of Four New Species From Asia And Australia, František Kovařík

Euscorpius

Four new species of the genus Isometrus are described: Isometrus (Reddyanus) bilyi from Australia, I. (R.) petrzelkai from Vietnam and Thailand, I. (R.) problematicus from India and I. (R.) khammamensis from India. Keys are presented to all species of both subgenera of Isometrus. Lectotypes are designated for I. (I.) thurstoni Pocock, 1893 and I. (R.) assamensis Oates, 1888. I. (I.) sankariensis Tikader & Bastawade, 1983 is synonymized with I. (I.) thurstoni Pocock, 1893. First time records include I. (I.) thurstoni Pocock, …

High-Level Systematics And Phylogeny Of The Extant Scorpions (Scorpiones: Orthosterni), Michael E. Soleglad, Victor Fet

Euscorpius

A number of authors (e. g. Birula, 1917a, 1917b; Mello-Leitão, 1945; Stockwell, 1989) addressed above-level systematics of extant scorpions, and accepted the grouping of scorpion families in several superfamilies. At the same time, Kjellesvig-Waering (1986) classified all extant scorpions under the same superfamily, Scorpionoidea. Sissom (1990) and Fet et al. (2000) did not list any superfamilies, considering the systematic situation above family (and often at the family level as well) unresolved. Most recently, Lourenço (2000a) listed six superfamilies, largely following the unpublished but important study of Stockwell (1989). The goal of this paper is to address scorpion systematics and phylogeny …

The Scorpion Sternum: Structure And Phylogeny (Scorpiones: Orthosterni), Michael E. Soleglad, Victor Fet

Euscorpius

The structure of the sternum of all major Recent scorpion groups is analyzed in detail. Based on this analysis, two fundamental sternum types are identified, described and illustrated, type 1 and type 2. These sternum types are distinguished by criteria based on external and internal structural features. The sternum types described herein are offered as a replacement for the various characterizations used throughout the last 140 years which emphasize only gross overall shape and proportions. Phylogenetic and taxonomic ramifications of these new sternal types are discussed. The Carboniferous fossil scorpion Palaeopisthacanthus schucherti Petrunkevitch is assigned to sternum type 1 …

Genetic Analyses Determine Connectivity Among Cave And Surface Populations Of The Jamaican Endemic Freshwater Crab Sesarma Fossarum In The Cockpit Country, Manuel Stemmer, Christoph D. Schubart

International Journal of Speleology

The Jamaican freshwater crab Sesarma fossarum (Decapoda: Brachyura: Sesarmidae) is endemic to western central Jamaica where it occurs in cave and surface streams of karst regions. In the present study, we examine the population genetic structure of the species, providing evidence for intraspecific differentiation and genetic substructure among twelve sampled populations. Interestingly, crabs from caves appear genetically undistinguishable from representatives of nearby surface waters, despite previously observed and described morphometric differentiation. In contrast, genetic isolation takes place among populations from rivers and caves belonging to different watersheds. In one case, even populations from different tributaries of the same river were …

P-29 Diversity And Species Turnover Of Late Cenozoic Ground Squirrels In The Meade Basin, Ks, H. Thomas Goodwin

Celebration of Research and Creative Scholarship

The Meade Basin of southwestern Kansas preserves one of the richest fossil sequences from the late Cenozoic of North America. An ongoing study has documented a rich record of fossils, especially rodents; established episodes of especially rapid species turnover (when species appear or disappear from the local record); and has attempted to relate these processes to environmental change inferred from stable isotope and other proxies for ancient climate. Here, I report on fossil squirrels from the record. At least 7 genera and 13 species are present in the composite record. Species turnover occurs throughout the record, but an especially high …

Sediment Loss Of Santa Rosa Island Slopes: An Erosional Study, Michael Perez

STAR Program Research Presentations

Ranching began on Santa Rosa Island in the 1840’s, consequently introducing nonnative megafauna that put novel selective grazing pressures on endemic plant species. Their movement patterns also altered sediment integrity as the land was denuded of any stabilizing vegetation. Dense groves of island oak (Q. tomentella) are known to aid in sediment deposition and retention. The groves also function to collect water during periods of intense fog that are common to the island. This experiment sought to quantify the volume of sediment that has been lost on a south facing slope in the middle of the island that …

A Systematic Review Of The Soricimorph Eulipotyphla (Soricidae: Mammalia) From The Gray Fossil Site (Hemphillian), Tennessee, Joshua Doby

Electronic Theses and Dissertations

Due in part to the incompleteness of the Cenozoic fossil record in the eastern U.S., the evolution and immigration of shrews (Soricidae) is not well understood. A rich soricid fauna from the Gray Fossil Site (GFS), Washington County, TN, has enabled many new inferences to be made. There are 7 new species in 6 genera: Paenelimnoecus, “Blarinella”, Petenyia, Tregosorex, Crusafontina, and Gen et sp. nov. GFS species of the genera Paenelimnoecus, “Blarinella”, and Petenyia are the first occurrence of each genus in the New World. Tregosorex, Crusafontina, and the N.A. taxon Limnoecus all have their latest documented occurrence at the …

Two New Euscorpius Species From Central-Western Greece (Scorpiones: Euscorpiidae), Gioele Tropea, Victor Fet

Euscorpius

Two new Euscorpius species are described, based on specimens collected by P.M. Giachino & D. Vailati in central-western Greece in neighboring Aitoloakarnania (Western Greece) and Fokida (Central Greece) regional units. No Euscorpius specimens were previously available from this area. The first new species, Euscorpius giachinoi sp. n., is very similar to a recently described E. birulai Fet et al., 2014 from Euboea Island, and is also characterized by a low trichobothrial count (Pv = 7, et = 5), a low pectinal teeth count (Dp = 7 in males, 6 in females), and long-limbed features. The second species, …

Effects Of Proline And Glycine On The Cnidocyte Discharge Of Hydra Magnipapillata, Janine R. Appleton

Honors Theses and Capstones

The sense of taste enables animals to utilize environmental cues to detect favorable foods. Through specialized sensory receptors, Cnidarians employ stinging cells called cnidocytes to perform a variety of activities such as locomotion, capturing prey, inducing of feeding responses, and defense. Their discharge is highly regulated by mechanical and chemical signals that are mediated by a complex system including the opsin and taste pathways. Taste 1 Receptors (T1R) have previously been isolated in vertebrates but only until recently, have been noted in invertebrates. Receptors specific to L- amino acids corresponding to the taste sensation of umami, were studied to determine …

Systematics Of The African River Frog Genus Amietia (Anura: Pyxicephalidae) From Eastern Democratic Republic Of The Congo, Thornton Robert Larson

Open Access Theses & Dissertations

The African river frog genus Amietia is found near rivers and other lentic water sources throughout central, eastern and southern Africa. Because the genus includes multiple morphologically conservative species, taxonomic studies of river frogs have been limited. We sampled 49 individuals of Amietia from multiple localities in and near the Albertine Rift (AR) of Burundi, Democratic Republic of the Congo and Uganda. We utilized single-gene (16S) and concatenated (12S, 16S, cytochrome b and RAG1) gene-tree analyses and coalescent species-tree analyses to construct phylogenetic trees. Two divergence dating approaches were used in BEAST, including secondary calibration points with 12S, 16S and …

Phylogeny And Systematics Of Panaspis And Afroablepharus Skinks (Squamata: Scincidae) In The Savannas Of Sub-Saharan Africa, Maria Fernanda Medina

Open Access Theses & Dissertations

African snake-eyed skinks are relatively small lizards of the genera Panaspis and Afroablepharus. The allocation of these genera was frequently rearranged in the 20th century based on morphology, ecology and biogeography. Members of these genera occur primarily in savanna habitats throughout sub-Saharan Africa and include species that have highly conserved morphology, which poses a challenge for taxonomic studies. We sequenced two mitochondrial (16S and cyt b) and two nuclear genes (PDC and RAG1) from 91 Panaspis and Afroablepharus samples from various localities in eastern, central and southern Africa. Concatenated gene-tree and divergence dating analyses were conducted to infer phylogenies and …

Inter-And Intra-Population Variability Across The Transcriptome Of Lake Baikal’S Endemic Copepod With Ramifications For Adapting To Climate Change, Larry L. Bowman Jr

Electronic Theses and Dissertations

The future of Lake Baikal’s biodiversity is uncertain in response to climate change. Unlike its diverse benthos, Lake Baikal’s zooplankton is species poor, with up to 96% of its biomass being composed of a single Calanoid copepod species, Epischura baikalensis. This study characterizes the genetic differentiation and differential gene expression of E. baikalensis. Using partial-transcriptome sequences obtained by 454 Rosche and Illumina sequencing technologies, the genetic differentiation at inferred single nucleotide polymorphism (SNP) sites and differential gene expression in populations sampled from various parts of the lake were analyzed. The functional genomics of genes showed significant differential …

Fossil Crocodilians From The High Guajira Peninsula Of Colombia, And The History Of Neogene Crocodilian Diversity In Tropical South America, Jorge W. Moreno-Bernal

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

The greatest diversity of Cenozoic crocodilians occurred during the Miocene in equatorial South America. However, the origin of this high diversity and its relationship to environmental factors are poorly understood. Most described species come from localities assigned to Laventan (13.8-11.8 Ma) and Huayquerian (9.0-6.8 Ma) South American land mammal ages (SALMAS), whereas the record is sparse in the early to middle Miocene and after the latest Miocene and Pliocene. Field research in the Castilletes (early Miocene-Pliocene) Formation in the High Guajira Peninsula of Colombia provides new fossil data on the origin of Neotropical crocodylian diversity. The Castilletes Formation crops out …