Evolution Commons^™

Micro- And Macroevolution Of Drab Plumage Color In Birds, Rafael Marcondes

LSU Doctoral Dissertations

I address the evolution of drab bird colors at micro- and macroevolutionary scales. I use as a study system the Furnariida, a clade of >600 Neotropical passerine species that, despite tremendous ecological and morphological diversity, are colored almost exclusively in shades of brown and grey produced by melanin pigments.

In chapters two and three, I took a macroevolutionary approach and showed that (1) plumage colors in the Furnariida evolve at similar rates in a monochromatic clade, males of a dichromatic clade, and females of a dichromatic clade; (2) ventral plumage color evolves faster than dorsal; (3) bird species that occupy …

Habitat Characteristics Of Two Scorpion Species, Liocheles Australasiae (Fabricius, 1775) And Isometrus Maculatus (De Geer, 1778) In Miyako Islands, Japan, Kazusa Kawai

Euscorpius

Scorpion surveys were conducted in the five islands: Miyakojima, Ikemajima, Kurimajima, Irabujima, and Shimojishima (Miyako Islands, part of Nansei/Ryukyu Islands of Japan). Eight individuals of Liocheles australasiae (Hormuridae) and 16 individuals of Isometrus maculatus (Buthidae) were observed in a total of 10 points on four islands, except Ikemajima. The localities of both species are recorded in this survey, focusing on altitude, distance from the nearest coast, and sunlight. The lowest temperatures in the habitat are also examined, and the cold tolerance of the species is discussed.

Scorpions Of The Horn Of Africa (Arachnida: Scorpiones). Part Xxvi. Records Of Hottentotta Polystictus (Pocock, 1896), With Descriptions Of H. Haudensis Sp. N. And H. Nigrimontanus Sp. N. (Buthidae) From Somaliland, František Kovařík, Graeme Lowe

Euscorpius

The distribution of Hottentotta polystictus (Pocock, 1896) is summarized according to recently confirmed records. Two new species, Hottentotta haudensis sp. n. and H. nigrimontanus sp. n. from the Somaliland, are described, compared with H. polystictus, and fully illustrated with color photos showing their morphology, habitus and collection areas. H. haudensis sp. n. is the smallest known species of the genus Hottentotta Birula, 1908, while H. nigrimontanus sp. n. is larger than H. polystictus.

Predation Of Ananteris Spp. (Scorpiones: Buthidae) By Ants And A Social Wasp (Hymenoptera: Formicidae, Vespidae) In Panama, Central America, Roberto Miranda, Luis F. De Armas, Roberto A. Cambra

Euscorpius

Two events of predation on Ananteris spp. were observed in the Parque Nacional Darién, Darién Province, Panama. The first case involved the social wasp Agelaia centralis (Cameron, 1907) (Vespidae); the second one, a ponerine ant Leptogenys cf. cuneata Lattke, 2011 (Formicidae). This is the first time that predation by a social wasp on a scorpion is observed. They also represent the first recorded predators for Panamanian Ananteris species.

Fetilinia Dentator Gen. Et Sp. N. From Pakistan (Scorpiones: Buthidae), Graeme Lowe, František Kovařík

Euscorpius

We describe from northern Pakistan a new buthid genus Fetilinia gen. n., monotypic with type species F. dentator sp. n., belonging to the Palaearctic ‘Buthus’ group. It bears some similarities to Kraepelinia Vachon, 1974, in having enlarged metasomal dentition, and to Orthochirus Karsch, 1891, and related genera in having a trapezoidal carapace and small abbreviated pedipalps.

Using Audio Recording Devices And Multiseason Occupancy Modelling To Evaluate Clark's Nutcracker (Nucifraga Columbiana) Occurrence In Whitebark Pine Habitats Of Washington's Cascade Mountains, Lauren Taracka

Undergraduate Honors Theses

The Clark’s nutcracker (Nucifraga columbiana) and whitebark pine (Pinus albicaulis), both native to the harsh subalpine habitats of Washington’s Cascade Mountains, are being negatively impacted by white pine blister rust (Cronartium ribicola), the mountain pine beetle (Dendroctonus ponderosae), and climate change. As temperatures continue to warm, while many species at lower elevations can move upward in elevation to suit their temperature needs, subalpine species do not have that option. Whitebark pine is a keystone tree species - numerous animals consume its high fat, high energy seeds; it’s a nurse tree, providing shelter …

Lepidopteran Granivory Reduces Seed Counts In A Rare Species Of Riparian Scour Prairies, Cheyenne Moore, Angela J. Mcdonnell, Scott Schuette, Christopher T. Martine

Faculty Journal Articles

In Pennsylvania Baptisia australis var. australis is found along only four waterways: the Allegheny River, Youghiogheny River, Clarion River, and Red Bank Creek. Because of its limited distribution and small number of extant populations, the species is considered state-threatened in Pennsylvania. In addition, the riparian prairie habitat that Pennsylvania Baptisia australis var. australis is restricted to is also in decline and considered vulnerable. Because of these conservation concerns, insights into the natural history of the taxon in the state is valuable and will inform conservation efforts. Field surveys and fruit collections along the Allegheny River and herbarium collections were used …

Alfred Russel Wallace Notes 8: Wallace's Earliest Exposures To The Writings Of Alexander Von Humboldt, Charles H. Smith

Faculty/Staff Personal Papers

For many years it has been known that the German geographer and naturalist Alexander von Humboldt (1769-1859) exercised a considerable influence on Alfred Russel Wallace’s (1823-1913) decision to become a travelling naturalist. Almost no attention, however, has been given to the real possibility that von Humboldt exerted an even greater influence on Wallace’s general worldview. In this note, Wallace’s probable earliest contacts with von Humboldt’s writings are described.

A Desert Tortoise–Common Raven Viable Conflict Threshold, Kerry L. Holcomb, Peter S. Coates, Brian G. Prochazka, Timothy Shields, William I. Boarman

Human–Wildlife Interactions

Since 1966, common raven (Corvus corax; raven) abundance has increased throughout much of this species’ Holarctic distribution, fueled by an ever-expanding supply of anthropogenic resource subsidies (e.g., water, food, shelter, and nesting substrate) to ecoregion specific raven population carrying capacities. Consequently, ravens are implicated in declines of both avian and reptilian species of conservation concern, including the California (USA) endangered and federally threatened Mojave desert tortoise (Gopherus agassizii; desert tortoise). While ravens are a natural predator of desert tortoises, the inter-generational stability of desert tortoise populations is expected to be compromised as annual juvenile survival is …

Evaluation Of The Diversity Of Durum Wheat Coleoptera (Triticum Durum Desf.) In The Region Of Sigus Oum El Bouaghi (Eastern Algeria), Amokrane Assia, Khammar Hichem, Hadjab Ramzi, Saheb Menouar

Journal of Bioresource Management

The Oum El Bouaghi region in Eastern Algeria, long considered as a cereal-growing area is nowadays a durum wheat production region par excellence. Although the damage caused by Coleoptera is very significant, studies on the knowledge of their diversity are few and remain limited for some entomological groups. Our work is the first step to evaluate the diversity of Coleoptera and the long-term impact of taking biological management measures against harmful fauna in favour of more environment friendly agriculture. To assess the beetle community, different sampling methods were combined (Barber trap, coloured traps, mowing net and sight hunting). Evaluation of …

A New Cryptic Species Of Scorpiops Peters, 1861 (Scorpiones: Scorpiopidae) From The Northern Western Ghats, India, Shauri Sulakhe, Shubhankar Deshpande, Nikhil Dandekar, Makarand Ketkar, Anand Padhye, Deshabhushan Bastawade

Euscorpius

A new cryptic species of Scorpiops (Scorpiopidae) is described from northern Western Ghats of India with integrated taxonomic approach. Scorpiops telbaila sp. n. is closely related to S. tenuicauda and differs from all species of Scorpiops in morphological features and raw genetic divergence of 5.4-14.1 %.

Euscorpius Thracicus Sp. N. (Scorpiones: Euscorpiidae) From Bulgaria, František Kovařík, Graeme Lowe, Markéta Byronová, František Šťáhlavský

Euscorpius

Euscorpius thracicus sp. n. from eastern Bulgaria is described, fully complemented with color photographs of both live and preserved specimens, as well as their habitats. This species is described based on morphology and the cytochrome oxidase I (COI) DNA barcoding marker. In addition to the analyses of external morphology and hemispermatophore, we also describe the karyotype of E. thracicus sp. n. (2n=92).

Revision Of Genus-Group Taxa In The Family Scorpiopidae Kraepelin, 1905, With Description Of 15 New Species (Arachnida: Scorpiones), František Kovařík, Graeme Lowe, Mark Stockmann, František Šťáhlavský

Euscorpius

The scorpiopids comprise a large scorpion family, distributed widely across the Oriental region, that currently encompasses 82 species placed under 8 genera and one subgenus. A variety of morphological characters have been used in the diagnoses of these genera or subgenera. Some characters were introduced by Vachon (1980) and Soleglad & Sissom (2001) when less than a few dozen species were known, while other more recent characters have not been fully evaluated. We critically review all characters that have been applied to generic or subgeneric taxonomy of the family, as well as several others hitherto neglected. We analyze their variation …

Two New Chaerilus From Thailand And Laos (Scorpiones: Chaerilidae), František Kovařík, Graeme Lowe, Mark Stockmann, František Šťáhlavský

Euscorpius

Chaerilus kautti sp. n. from Thailand and C. pulcherrimus sp. n. from Laos are described and fully illustrated with color photographs of live and preserved specimens. C. kautti sp. n. is characterized by the unique combination of two characters: movable finger of pedipalp with 11 imbricated rows of granules; and pedipalp chela length/width ratio in male 4.55. C. pulcherrimus sp. n. is characterized by the unique combination of four characters: movable finger of pedipalp with 8 imbricated rows of granules; total length 16–19 mm; pedipalp chela length/width ratio in male 2.57, in female 2.69; and sternite VII smooth. In addition …

Quantifying Structure And Variation In Complex Phylogenetic Data, Genevieve Geraldine Mount

LSU Doctoral Dissertations

Identifying the source and structure of variation in nature is crucial to understanding fundamental aspects of evolution. Despite a recent plethora of genetic and morphological data, many interesting questions about the relationships between different groups remain unresolved. My dissertation evaluates three approaches for identifying and quantifying the variation within phylogenetic datasets. Characterizing variation within datasets and across analytical methods gives insight into biologically interesting characters, unusual evolutionary processes, and areas for model improvement.

Network-based community detection approaches offer a powerful tool to describe variation in phylogenetic signal across the genome (i.e., gene tree variation). In Chapter 2, I investigate the …

Effect Of Seasons And Scorpion Size On The Foraging And Diet Of The Striped Bark Scorpion, Centruroides Vittatus (Buthidae: Scorpiones) In Blackbrush Habitat Of South Texas, C. Neal Mcreynolds

Euscorpius

Diet and foraging success of the striped bark scorpion, Centruroides vittatus, in South Texas are influenced by both scorpion size and season of the year. In the ten-year study of the striped bark scorpions in the blackbrush habitat of south Texas, the diet was variable with caterpillars (Lepidoptera) as the main prey for all seasons and all size classes of scorpions. The proportion of caterpillars did vary significantly with size class of scorpion and months of the year with intermediate size scorpions capturing more caterpillars during January–April than other size classes or months of the year. The proportion of …

Scorpions Of The Horn Of Africa (Arachnida: Scorpiones). Part Xxv. Description Of Pandinurus Awalei Sp. N. And The Male Of Pandiborellius Somalilandus (Kovařík, 2012), With Remarks On Recent Synonymies (Scorpionidae: Pandininae), František Kovařík, Graeme Lowe, Hassan Sh A. Elmi

Euscorpius

A new species Pandinurus awalei sp. n. is described from Somaliland. The male of Pandiborellius somalilandus (Kovařík, 2012) is also described for the first time and sexual dimorphism of the species is defined. The habitus, morphology and habitats of both species are illustrated in detail with color and UV fluorescence images, including both live and preserved specimens. The species Pandinurus intermedius (Borelli, 1919) and Pandipalpus lowei (Kovařík, 2012) of Pandininae that were synonymized by Prendini & Loria (2020) are reinstated as valid species.

Reddyanus Justi Sp. N. From Laos (Scorpiones: Buthidae), František Kovařík, Graeme Lowe, František Šťáhlavský

Euscorpius

A new species Reddyanus justi sp. n. from Laos is described, fully complemented with color photographs of live and preserved specimens. The new species is characterized by the irregular quadrilateral shape of the subaculear tubercle in males. In addition to analysis of external morphology and hemispermatophore, we also describe the karyotype (2n=12)

Two New Species Of Chiromachetes (Scorpiones: Hormuridae) From The Northern Western Ghats, India, Shauri Sulakhe, Shubhankar Deshpande, Nikhil Dandekar, Makarand Ketkar, Gaurang Gowande, Anand Padhye, Deshabhushan Bastawade

Euscorpius

Two new species of Chiromachetes Pocock, 1899 (Hormuridae) are described from the northern Western Ghats of India using an integrated taxonomic approach. Chiromachetes parakrami sp. n. and C. ramdasswamii sp. n. are closely related and differ from each other and C. sahyadriensis by morphological features and raw genetic divergence of 7.9–9.4 %.

Тaxonomic Position Of Orthochirus Olivaceus (Karsch, 1881), The Type Species Of The Genus Orthochirus Karsch, 1892 (Scorpiones: Buthidae), František Kovařík, Victor Fet, Manal Siyam

Euscorpius

We address the taxonomic position of Orthodactylus olivaceus Karsch, 1881, the type species of the genus Orthochirus Karsch, 1892. For over a century, this taxon was considered a synonym of Orthochirus scrobiculosus (Grube, 1873); however, the latter was recently limited to Central Asia (Turkmenistan) (Kovařík et al., 2020). Analysis of Karsch’s holotype confirmed that Orthochirus olivaceus (Karsch, 1881), is a valid species. Its type locality published as “Sicily” is clearly incorrect. It is a senior synonym of Butheolus aristidis Simon, 1882, syn. n., described from Sudan (Nubia). We confirm its distribution in northern Sudan and southern Egypt.

Further Review Of Orthochirus Karsch, 1892 (Scorpiones: Buthidae) From Asia: Taxonomic Position Of O. Melanurus, O. Persa, O. Scrobiculosus, And Description Of Six New Species, František Kovařík, Victor Fet, Ersen A. Yağmur

Euscorpius

We describe six new species of Orthochirus: O. birulai sp. n. (Pakistan), O. formozovi sp. n. (Afghanistan, Iran, Tajikistan, Turkmenistan), O. grosseri sp. n. (Uzbekistan), O. kryzhanovskyi sp. n. (Pakistan), O. nordmanni sp. n. (Afghanistan), and O. sejnai sp. n. (Iran). Descriptions are complemented with color photographs of preserved specimens. The identities of Orthochirus melanurus (Kessler, 1874) (Kazakhstan, Uzbekistan), O. persa (Birula, 1900), stat. n. (Afghanistan, Iran), and O. scrobiculosus (Grube, 1873) (Turkmenistan) are reexamined, based on detailed study of the type specimens; lectotypes of all three species are designated. We demonstrate for the first time that the name …

Description Of The Male Of Wernerius Mumai (Sissom, 1993) From Western Arizona, With Data On Reproduction (Scorpiones: Vaejovidae), Richard F. Ayrey, Brandon T. Myers

Euscorpius

A male of Wernerius mumai (Sissom, 1993) is described for the first time, collected at the type locality (Gold Road, Black Mountains, Mohave County, Arizona). Originally placed in the genus Vaejovis, this species later was transferred by Soleglad & Fet (2008) to the genus Wernerius. It is one of the smallest vaejovid species known. The pedipalp fixed finger usually has 6 ID denticles and the movable finger has 7. The most unique characteristic of this species is the long, pointed, subaculear tubercle.

Integrative Species Delimitation And Taxonomic Status Of The Scorpion Genus Vaejovis Koch, 1836 (Vaejovidae) In The Santa Catalina Mountains, Arizona, Emma E. Jochim, Lillian-Lee M. Broussard, Brent E. Hendrixson

Euscorpius

Scorpions belonging to the Vaejovis vorhiesi species complex are widely distributed throughout the southwestern United States and northern Mexico. Most species are endemic to single mountain ranges but two species, Vaejovis deboerae Ayrey, 2009 and V. brysoni Ayrey & Webber, 2013, have been documented from the Santa Catalina Mountains in Arizona. We reevaluated the taxonomic diversity of these scorpions by integrating data from several different sources. Phylogenetic analyses indicate that scorpions in the Santa Catalina Mountains are monophyletic but comprise two divergent mitochondrial lineages that overlap at the type locality of V. deboerae. We failed to detect congruence between …

Interrelationships Among Monorchiid Trematodes With Special Emphasis On Some Northwestern Atlantic Genera, Apryle Panyi

Master's Theses

The Monorchiidae Odhner, 1911 is a cosmopolitan family of flukes (Trematoda: Digenea) comprising species that parasitize the digestive tract of estuarine and marine fishes as adults. Compared with other oceans, recent morphological or molecular taxonomic work conducted on monorchiid species from the northwestern Atlantic Ocean has been sparse (Manter, 1931; Overstreet, 1969; Andres et al., 2018; Wee et al., 2018, 2019, 2020). Therefore, the present work investigated the interrelationships of some monorchiids from the northwestern Atlantic Ocean with emphasis on several genera and investigated if Lasiotocus minutus (Manter, 1931) Thomas, 1959 constitutes a complex of cryptic species. New morphological and …

Five New Species Of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) From Albania, Greece, North Macedonia, And Serbia, František Kovařík, František Šťáhlavský

Euscorpius

Five new species are described: Euscorpius bonacinai sp. n. (Albania), E. janstai sp. n. (North Macedonia), E. kabateki sp. n. (Greece), E. sadileki sp. n. (Serbia), and E. scheraboni sp. n. (Greece), fully complemented with color photographs. New species are distinguished from all other species of the genus on the basis of five taxonomic characters. Two other new characters, shape of the pedipalp chela fingers and number of chromosomes, are presented and discussed. In addition to the analyses of external morphology, we also describe karyotypes of two species: E. janstai sp. n. (2n=112) and E. sadileki sp. n. (2n=90).

Notes On The Diet And Reproduction Of The Bark Scorpion Centruroides Gracilis (Scorpiones: Buthidae) On Utila Island, Honduras, Anna M. Davison, Tom W. Brown, Cristina Arrivillaga

Euscorpius

The brown bark scorpion Centruroides gracilis (Latreille, 1804) (Buthidae) is an abundant and widespread species, however, fairly little is known about its ecology and natural history. We include several observations from Utila Island, Honduras, that expand on the known literature regarding the diet and reproductive behavior of C. gracilis. We report several prey items for this opportunistic species, which include invertebrates such as spiders (including tarantulas), centipedes, katydids, and crickets; as well as a case of cannibalism between two adult females C. gracilis. We suggest that such cannibalism may be driven by high population densities and/or strong intraspecific …

A New Lesser Antillean Scorpion Of The Genus Didymocentrus Kraepelin, 1905 (Scorpiones: Diplocentridae), Rolando Teruel, Karl Questel

Euscorpius

A new species of the diplocentrid scorpion of the genus Didymocentrus Kraepelin, 1905 is herein described from the Lesser Antillean island of Martinique. It long remained misidentified as Didymocentrus lesueurii (Gervais, 1844), the only species of this genus inhabiting the adjacent island of Saint Lucia. The direct comparison of adults of both sexes from these two islands revealed solid morphological characters, which warrant their recognition as two different taxa. Each one is endemic from a single major island, where it is widespread even in its satellite islets. This addition raises the known composition of the genus to 12 species, 10 …

Six New Species Of Orthochirus Karsch, 1892 From Iran (Scorpiones: Buthidae), František Kovařík, Shahrokh Navidpour

Euscorpius

Six new scorpion species from Iran, Orthochirus hormozganensis sp. n. (Hormozgan Province), O. kermanensis sp. n. (Kerman Province), O. kucerai sp. n. (Kerman Province), O. masihipouri sp. n. (Bushehr Province), O. semnanensis sp. n. (Semnan Province), and O. vignolii sp. n. (Yazd Province) are described, compared with other Iranian Orthochirus species, and fully illustrated with color photographs. A key and a distribution map of Orthochirus of Iran, Turkey, and Iraq (18 species) are included.

Scorpions Of Puerto Rico And Its Satellite Islands (Scorpiones: Buthidae, Diplocentridae): An Аnnotated List, Key For Genera, And Bibliography, Luis De Armas

Euscorpius

An annotated list of the scorpions of Puerto Rico and its satellite islands (mainly Vieques, Culebra, Caja de Muertos, Magueyes, Desecheo, and Mona), a key for the seven genera present in those insular territories, a key for the three species of Microtityus, and the basic bibliography on taxonomy and natural history are provided. Of the 16 confirmed species described from these Greater Antillean islands, four are known only from a single sex and only one or two adult specimens. Seven species (six of them from the genus Tityus), are endemic to the Puerto Rico Island; Vieques has two …

A New Species Of Isometrus (Scorpiones: Buthidae) From Southern India, Shauri Sulakhe, Nikhil Dandekar, Shomen Mukherjee, Malay Pandey, Makarand Ketkar, Anand Padhye, Deshabhushan Bastawade

Euscorpius

A new species of Isometrus (Buthidae) is described from India using integrated taxonomic approach. Isometrus kovariki sp. n. is closely related to I. thurstoni, and differs in morphological features and raw genetic divergence of more than 9%.