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Full-Text Articles in Evolution

Scorpions Of The Horn Of Africa (Arachnida, Scorpiones). Part Xvii.Revision Of Neobuthus, With Description Of Seven New Species From Ethiopia, Kenya And Somaliland (Buthidae), František Kovařík, Graeme Lowe, Ahmed Ibrahim Awale, Hassan Sh Abdirahman Elmi, Ali Abdi Hurre Dec 2018

Scorpions Of The Horn Of Africa (Arachnida, Scorpiones). Part Xvii.Revision Of Neobuthus, With Description Of Seven New Species From Ethiopia, Kenya And Somaliland (Buthidae), František Kovařík, Graeme Lowe, Ahmed Ibrahim Awale, Hassan Sh Abdirahman Elmi, Ali Abdi Hurre

Euscorpius

New information about the taxonomy and distribution of the genus Neobuthus Hirst, 1911 is presented, based on material recently collected mainly from Somaliland, but also Djibouti and Kenya. Emended diagnoses are proposed for N. berberensis Hirst, 1911 and N. ferrugineus (Kraepelin, 1898) sensu stricto. New species described are: N. amoudensis sp. n. from Ethiopia and Somaliland; N. erigavoensis sp. n., N. factorio sp. n., N. gubanensis sp. n., N. maidensis sp. n., and N. montanus sp. n. from Somaliland; and N. kloppersi sp. n. from Kenya. This doubles the number of species in Neobuthus, unearthing a rich diversification …

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Revision Of The Central Asian Scorpion Genus Anomalobuthus Kraepelin, 1900, With Descriptions Of Three New Species And A Generic Synonymy (Scorpiones: Buthidae), Rolando Teruel, František Kovařík, Victor Fet Dec 2018

Revision Of The Central Asian Scorpion Genus Anomalobuthus Kraepelin, 1900, With Descriptions Of Three New Species And A Generic Synonymy (Scorpiones: Buthidae), Rolando Teruel, František Kovařík, Victor Fet

Euscorpius

We revise the Central Asian endemic genus Anomalobuthus Kraepelin, 1900, which was considered monotypic for more than 100 years until the recent addition of a second species from Iran (Teruel et al., 2014). We redefine the generic diagnosis of Anomalobuthus and reveal that it is composed of no less than six species, three of which are described as new: A. krivochatskyi, sp. n. (central Uzbekistan and extreme southern Kazakhstan), A. lowei, sp. n. (southeastern Kazakhstan), and A. pavlovskyi, sp. n. (south-central Kazakhstan and extreme northern Turkmenistan). The monotypic genus Psammobuthus Birula, 1911 (described from the Ferghana Valley …

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Courtship Behavior, Communication, And Copulation In Tigrosa Annexa, Samuel White Nov 2018

Courtship Behavior, Communication, And Copulation In Tigrosa Annexa, Samuel White

Scholars Week

The evolution of multimodal communication, where signalers use multiple signal components in multiple sensory modalities, has become the subject of investigation by many researchers. Signaling puts males at risk of predation, so why do males of some species evolve extra signals that may increase this risk? In some wolf spider species, males incorporate many visual and vibrational signals into a display that they use to attract a female for mating. Female spiders are often aggressive toward courting males and so the male display also functions to decrease the odds of cannibalism. Female wandering spiders deposit silk containing pheromones that communicate …

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Notes On The Genera Buthacus, Compsobuthus, And Lanzatus With Several Synonymies And Corrections Of Published Characters (Scorpiones: Buthidae), František Kovařík Oct 2018

Notes On The Genera Buthacus, Compsobuthus, And Lanzatus With Several Synonymies And Corrections Of Published Characters (Scorpiones: Buthidae), František Kovařík

Euscorpius

The taxonomic position of Buthacus armasi Lourenço, 2013, B. clevai Lourenço, 2001, B. huberi Lourenço, 2001, B. maliensis Lourenço & Qi, 2007, B. nigerianus Lourenço & Qi, 2006, Compsobuthus andresi Lourenço, 2004, C. simoni Lourenço, 1999, C. tassili Lourenço, 2010, C. tofti Lourenço, 2001, C. williamsi Lourenço, 1999, and Sabinebuthus elegans Lourenço, 2001 is revised and fictitious characters in their original descriptions are discussed and corrected. Buthacus armasi Lourenço, 2013 is synonymized with Buthacus leptochelys (Ehrenberg, 1829) syn. n., B. huberi Lourenço, 2001 is confirmed to be a synonym of Buthacus occidentalis Vachon, 1953, B. maliensis Lourenço & Qi, …

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Three New Chaerilus From Malaysia (Tioman Island) And Thailand (Scorpiones: Chaerilidae), With A Review Of C. Cimrmani, C. Sejnai, And C. Tichyi, František Kovařík, Graeme Lowe, František Šťáhlavský Sep 2018

Three New Chaerilus From Malaysia (Tioman Island) And Thailand (Scorpiones: Chaerilidae), With A Review Of C. Cimrmani, C. Sejnai, And C. Tichyi, František Kovařík, Graeme Lowe, František Šťáhlavský

Euscorpius

Chaerilus majkusi sp. n. from Malaysia (Tioman Island), C. neradorum sp. n. and C. stockmannorum sp. n. from Thailand are described and fully illustrated with color photographs of live and preserved specimens, as well as of their habitat. They are compared to the species C. cimrmani Kovařík, 2012, C. sejnai Kovařík, 2005, and C. tichyi Kovařík, 2000, which we also illustrate with color photographs of live unpublished specimens. Hemispermatophores of C. cimrmani, C. majkusi sp .n., C. stockmannorum sp. n., and C. tichyi are illustrated and compared, and we also describe the karyotypes of C. cimrmani, C. …

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Review Of The Genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), With Descriptions Of Barbaracurus Gen. N. And Two New Species From Oman And Yemen, František Kovařík, Graeme Lowe, František Šťáhlavský Aug 2018

Review Of The Genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), With Descriptions Of Barbaracurus Gen. N. And Two New Species From Oman And Yemen, František Kovařík, Graeme Lowe, František Šťáhlavský

Euscorpius

The genus Babycurus Karsch, 1886 sensu lato is split into two genera, a strictly African genus Babycurus, and the new genus Barbaracurus gen. n., which mainly includes species from the Horn of Africa and Arabian Peninsula. Two new species Barbaracurus winklerorum sp. n. from Oman and B. yemenensis sp. n. from Yemen are described, compared with other species, and fully illustrated with color photos of morphology, habitus, live specimens and collection localities. Males of Barbaracurus somalicus (Hirst, 1907) comb. n. and Barbaracurus zambonellii (Borelli, 1902) comb. n. are recorded for the first time and fully illustrated. Babycurus ornatus …

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A New Scorpion Species From Kenya, Gint Childsi Sp. N. (Scorpiones: Buthidae), Frantíšek Kovařík Jul 2018

A New Scorpion Species From Kenya, Gint Childsi Sp. N. (Scorpiones: Buthidae), Frantíšek Kovařík

Euscorpius

Gint childsi sp. n. from Kenya is described and compared with other species of the genus. Additional information is provided on the taxonomy and distribution of the genus Gint, fully complemented with color photos of preserved specimens of both sexes of the new species, as well as of their habitat.

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Two New Hottentotta Species From Iran, With A Review Of Hottentotta Saulcyi (Scorpiones: Buthidae), Frantíšek Kovařík, Ersen Aydin Yağmur, Mohammed Moradi Jul 2018

Two New Hottentotta Species From Iran, With A Review Of Hottentotta Saulcyi (Scorpiones: Buthidae), Frantíšek Kovařík, Ersen Aydin Yağmur, Mohammed Moradi

Euscorpius

Two new buthid species from Iran, Hottentotta navidpouri sp. n. (Hormozgan Province) and H. sistanensis sp. n. (Sistan and Baluchestan Province) are described, compared with H. saulcyi (Simon, 1880), and fully illustrated with color photos. The two new species differ from H. saulcyi mainly in shape of chela, which is strongly elongated in both new species.

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A New Species Of Microtityus From The British Virgin Islands, West Indies, And New Localities For Other Scorpions (Scorpiones: Buthidae, Scorpionidae), Luis F. De Armas Jul 2018

A New Species Of Microtityus From The British Virgin Islands, West Indies, And New Localities For Other Scorpions (Scorpiones: Buthidae, Scorpionidae), Luis F. De Armas

Euscorpius

Microtityus (Parvabsonus) eustatia sp. n. is herein described from the British Virgin Islands (West Indies): Eustatia Island (type locality), Virgin Gorda Island, and Camanoe Island, based on seven specimens (three males and four females). The new species closely resembles M. waeringi Francke & Sissom, 1980 from St. John Island and St. Thomas Island, U.S. Virgin Islands, differing mainly by the fixed finger of pedipalp having ten rows of denticles (nine in M. waeringi) and more attenuated metasoma. Also, new localities are recorded for Heteronebo yntemai Francke & Sissom, 1980 (Scorpionidae) and Centruroides griseus (C. L. Koch, 1844) …

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Review Of Microbuthus With Description Of M. Satyrus Sp. N. (Scorpiones, Buthidae) From Oman And Yemen, Graeme Lowe, František Kovařík, Mark Stockmann, František Šťáhlavský Jun 2018

Review Of Microbuthus With Description Of M. Satyrus Sp. N. (Scorpiones, Buthidae) From Oman And Yemen, Graeme Lowe, František Kovařík, Mark Stockmann, František Šťáhlavský

Euscorpius

The taxonomy of the genus Microbuthus is reviewed, and a new species from Oman and Yemen, M. satyrus sp. n., is described and fully illustrated with color photographs of live and preserved specimens, as well as of its habitat. It is compared to the closely similar species M. litoralis, which we also illustrate. Synonymy of the type species M. pusillus Kraepelin, 1898 with M. litoralis (Pavesi, 1885) is confirmed, and the species is recorded for the first time from Yemen. Hemispermatophores of M. satyrus sp. n., M. gardneri Lowe, 2010, and …

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A New Species Of Vaejovis From The Patagonia Mountains, Southern Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey Jun 2018

A New Species Of Vaejovis From The Patagonia Mountains, Southern Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey

Euscorpius

A new scorpion species, Vaejovis patagonia sp. nov. is described and placed in the “vorhiesi” group. This small, dark brown species is found near Patagonia, Arizona. It is geographically closest to V. troupi Ayrey & Soleglad, V. grahami Ayrey & Soleglad and V. vorhiesi Stahnke. Those three species are found in a triangle surrounding the Patagonia Mountains, the locality of Vaejovis patagonia. The pedipalp fixed finger has 5 ID denticles and the movable finger has 6, like most, but not all, of the other southern Arizona Vaejovis. Carapace of female is shorter than metasomal segment V.

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The Genera Butheolus Simon, 1882 And Xenobuthus Gen. Nov. (Scorpiones: Buthidae) In Oman, Graeme Lowe Jun 2018

The Genera Butheolus Simon, 1882 And Xenobuthus Gen. Nov. (Scorpiones: Buthidae) In Oman, Graeme Lowe

Euscorpius

The genus Butheolus Simon, 1882 is revised based on new material from Dhofar Province in Oman. B. gallagheri Vachon, 1980 is redescribed, and a related new species, B. harrisoni sp. n., is also described. The species B. anthracinus (Pocock, 1895) is redescribed and moved to a new genus Xenobuthus gen. n., that is differentiated from Butheolus by size, pedipalp finger dentition, setation, granulation and hemispermatophore structure, and a related new species, X. xanthus sp. n., is also described. Revised diagnoses are provided for the genus Butheolus, and for the species B. …

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Radical Social Ecology As Deep Pragmatism: A Call To The Abolition Of Systemic Dissonance And The Minimization Of Entropic Chaos, Arielle Brender May 2018

Radical Social Ecology As Deep Pragmatism: A Call To The Abolition Of Systemic Dissonance And The Minimization Of Entropic Chaos, Arielle Brender

Student Theses 2015-Present

This paper aims to shed light on the dissonance caused by the superimposition of Dominant Human Systems on Natural Systems. I highlight the synthetic nature of Dominant Human Systems as egoic and linguistic phenomenon manufactured by a mere portion of the human population, which renders them inherently oppressive unto peoples and landscapes whose wisdom were barred from the design process. In pursuing a radical pragmatic approach to mending the simultaneous oppression and destruction of the human being and the earth, I highlight the necessity of minimizing entropic chaos caused by excess energy expenditure, an essential feature of systems that aim …

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Scorpions Of The Horn Of Africa (Arachnida, Scorpiones). Part Xvi. Compsobuthus Maidensis Sp. N. (Buthidae) From Somaliland, František Kovařík Apr 2018

Scorpions Of The Horn Of Africa (Arachnida, Scorpiones). Part Xvi. Compsobuthus Maidensis Sp. N. (Buthidae) From Somaliland, František Kovařík

Euscorpius

Compsobuthus maidensis sp. n. from Somaliland is described and fully complemented with color photos of specimens, as well as its habitat. Data on the occurrence of the genus Compsobuthus Vachon, 1949 in the Horn of Africa is summarized.

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Scorpions Of The Horn Of Africa (Arachnida: Scorpiones). Part Xv. Review Of The Genus Gint Kovařík Et Al., 2013, With Description Of Three New Species From Somaliland (Scorpiones, Buthidae), František Kovařík, Graeme Lowe, Pavel Just, Ahmed Ibrahim Awale, Hassan Sh Abdirahman Elmi, František Šťáhlavský Apr 2018

Scorpions Of The Horn Of Africa (Arachnida: Scorpiones). Part Xv. Review Of The Genus Gint Kovařík Et Al., 2013, With Description Of Three New Species From Somaliland (Scorpiones, Buthidae), František Kovařík, Graeme Lowe, Pavel Just, Ahmed Ibrahim Awale, Hassan Sh Abdirahman Elmi, František Šťáhlavský

Euscorpius

We describe herein three new species of Buthidae: Gint amoudensis sp. n., G. gubanensis sp. n., and G. maidensis sp. n. from Somaliland. Additional information is provided on the taxonomy and distribution of other species of the genus Gint, fully complemented with color photos of live and preserved specimens, as well as of their habitat. The hemispermatophores of most Gint species are illustrated and described for the first time. In addition to the analyses of external morphology and hemispermatophores, we also describe the karyotype of four Gint species. The number of chromosomes is different for every one …

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Scorpions Of Sri Lanka (Arachnida, Scorpiones). Part Ii. Family Hormuridae, František Kovařík, Kithsiri B. Ranawana, V. A. Sanjeewa Jayarathne, Sanjaya Karunarathna, Alexander Ullrich Feb 2018

Scorpions Of Sri Lanka (Arachnida, Scorpiones). Part Ii. Family Hormuridae, František Kovařík, Kithsiri B. Ranawana, V. A. Sanjeewa Jayarathne, Sanjaya Karunarathna, Alexander Ullrich

Euscorpius

Liocheles australasiae (Fabricius, 1775) is reported for the first time from Sri Lanka. Sri Lankan population is fully complemented with color photos of live and preserved females, as well as its habitat.

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Two New Tityus C. L. Koch, 1836 (Scorpiones: Buthidae) From Hispaniola, Greater Antilles, Rolando Teruel, Gabriel De Los Santos Feb 2018

Two New Tityus C. L. Koch, 1836 (Scorpiones: Buthidae) From Hispaniola, Greater Antilles, Rolando Teruel, Gabriel De Los Santos

Euscorpius

Two new species of Buthidae scorpions of the genus Tityus C. L. Koch, 1836 are herein described from the Greater Antillean island of Hispaniola. One of them belongs to the "crassimanus" species-group and is known from an adult pair collected at Massif de la Hotte, in southwestern Haiti. The other belongs to the "quisqueyanus" species-group and is known from a single adult female from a high peak in the Central Range (= Cordillera Central), in northwestern Dominican Republic. Moreover, two fossil taxa from this island are retained as junior synonyms of Tityus geratus Santiago-Blay, 1988†.

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Scorpions Of The Horn Of Africa (Arachnida, Scorpiones). Part Xiv. Hottentotta Somalicus Sp. N. (Buthidae) From Somalia, František Kovařík Feb 2018

Scorpions Of The Horn Of Africa (Arachnida, Scorpiones). Part Xiv. Hottentotta Somalicus Sp. N. (Buthidae) From Somalia, František Kovařík

Euscorpius

Hottentotta somalicus sp. n. from Somalia is described and fully complemented with color photos. Morphologically it is similar to H. polystictus (Pocock, 1896). These two species have very narrow metasomal segments (1.63–1.73 in both sexes versus 1.31–1.61 in both sexes of other Hottentotta species from the Horn of Africa). H. polystictus and H. somalicus sp. n. occur in separate areas (Somaliland versus Somalia) and can be differentiated by color.

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Advancing Behavioural Genomics By Considering Timescale, Clare C. Rittschof, Kimberly A. Hughes Feb 2018

Advancing Behavioural Genomics By Considering Timescale, Clare C. Rittschof, Kimberly A. Hughes

Entomology Faculty Publications

Animal behavioural traits often covary with gene expression, pointing towards a genomic constraint on organismal responses to environmental cues. This pattern highlights a gap in our understanding of the time course of environmentally responsive gene expression, and moreover, how these dynamics are regulated. Advances in behavioural genomics explore how gene expression dynamics are correlated with behavioural traits that range from stable to highly labile. We consider the idea that certain genomic regulatory mechanisms may predict the timescale of an environmental effect on behaviour. This temporally minded approach could inform both organismal and evolutionary questions ranging from the remediation of early …

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Revision Of The Mesobuthus Caucasicus Complex From Central Asia, With Descriptions Of Six New Species (Scorpiones: Buthidae), Victor Fet, František Kovařík, Benjamin Gantenbein, Ronald C. Kaiser, Alexander K. Stewart, Matthew R. Graham Feb 2018

Revision Of The Mesobuthus Caucasicus Complex From Central Asia, With Descriptions Of Six New Species (Scorpiones: Buthidae), Victor Fet, František Kovařík, Benjamin Gantenbein, Ronald C. Kaiser, Alexander K. Stewart, Matthew R. Graham

Euscorpius

A widespread Mesobuthus caucasicus complex, which includes some of the most common scorpions found from the Caucasus to China, is revised for the first time based on new extensive collections from Central Asia, using both morphological and DNA marker data. Mesobuthus caucasicus (Nordmann, 1840), s.str. is restricted to the Caucasus Mts. Four taxa are elevated to species rank: M. fuscus (Birula, 1897) (Tajikistan), M. intermedius (Birula, 1897) (Tajikistan), M. kaznakovi (Birula, 1904) (Tajikistan, Uzbekistan), and M. parthorum (Pocock, 1889) (Afghanistan, Iran, Turkmenistan). Six new species are described: M. brutus sp. n. (Iran), M. elenae sp. n. (Tajikistan, Uzbekistan), M. gorelovi …

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Is There A Link Between Aging And Microbiome Diversity In Exceptional Mammalian Longevity?, Graham M. Hughes, John Leech, Sebastien J. Puechmaille, Jose V. Lopez, Emma C. Teeling Jan 2018

Is There A Link Between Aging And Microbiome Diversity In Exceptional Mammalian Longevity?, Graham M. Hughes, John Leech, Sebastien J. Puechmaille, Jose V. Lopez, Emma C. Teeling

Biology Faculty Articles

A changing microbiome has been linked to biological aging in mice and humans, suggesting a possible role of gut flora in pathogenic aging phenotypes. Many bat species have exceptional longevity given their body size and some can live up to ten times longer than expected with little signs of aging. This study explores the anal microbiome of the exceptionally long-lived Myotis myotis bat, investigating bacterial composition in both adult and juvenile bats to determine if the microbiome changes with age in a wild, long-lived non-model organism, using non-lethal sampling. The anal microbiome was sequenced using metabarcoding in more than 50 …

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Chickens Play To The Crowd, Cinzia Chiandetti Jan 2018

Chickens Play To The Crowd, Cinzia Chiandetti

Animal Sentience

The time was ripe for Marino’s review of chickens’ cognitive capacities. The research community, apart from expressing gratitude for Marino’s work, should now use it to increase public awareness of chickens’ abilities. People’s views on many animals are ill-informed. Scientists need to communicate and engage with the public about the relevance and societal implications of their findings.

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Post-Darwin Skepticism And Run-Of-The-Mill Suicide, John Hadley Jan 2018

Post-Darwin Skepticism And Run-Of-The-Mill Suicide, John Hadley

Animal Sentience

Peña-Guzmán’s depiction of the opponent of animal suicide as a conservative is a straw man. It is possible to accept that animals are self-conscious and reflexive yet still reject the view that they have the mental wherewithal to commit run-of-the-mill suicide. That animal behaviour can be positioned on a continuum of self-destructive behaviour does not establish that animals can intentionally kill themselves.

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An Adaptationist Perspective On Animal Suicide, Timothy P. Racine Jan 2018

An Adaptationist Perspective On Animal Suicide, Timothy P. Racine

Animal Sentience

Peña-Guzmán’s discussion of suicide in nonhuman animals has broad implications. In this commentary, I focus on the logical relation between suicide and intention. Proximate cause must be distinguished from ultimate function in explanations of suicide. I briefly discuss two adaptationist accounts of suicidal behavior.

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Lessons From Chimpanzee Sign Language Studies, Mary Lee Jensvold Jan 2018

Lessons From Chimpanzee Sign Language Studies, Mary Lee Jensvold

Animal Sentience

Claims are often made about behaviors being unique to humans; the evidence usually shows they are not. Sign language studies on chimpanzees may provide a useful model for comparative studies of suicide. A productive approach to comparative studies is to focus on observable behaviors rather than getting lost in the pitfalls of vague definitions and changing measures.

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Animal Suicide: An Account Worth Giving?, Irina Mikhalevich Jan 2018

Animal Suicide: An Account Worth Giving?, Irina Mikhalevich

Animal Sentience

Peña-Guzmán (2017) argues that empirical evidence and evolutionary theory compel us to treat the phenomenon of suicide as continuous in the animal kingdom. He defends a “continuist” account in which suicide is a multiply-realizable phenomenon characterized by self-injurious and self-annihilative behaviors. This view is problematic for several reasons. First, it appears to mischaracterize the Darwinian view that mind is continuous in nature. Second, by focusing only on surface-level features of behavior, it groups causally and etiologically disparate phenomena under a single conceptual umbrella, thereby reducing the account’s explanatory power. Third, it obscures existing analyses of suicide in biomedical ethics and …

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Can Nondolphins Commit Suicide?, David M. Peña-Guzmán Jan 2018

Can Nondolphins Commit Suicide?, David M. Peña-Guzmán

Animal Sentience

This Response addresses the scientific and philosophical criticisms of my 2017 target article “Can nonhuman animals commit suicide?” It defends my key claims and explores topics (such as animal judgment, animal theory of mind, and the evolution of suicide) that did not appear in the original article. It also points out areas in which further research is needed and concludes that we should be wary of accusations of “anthropomorphism” in debates about animal suicide.

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Animal Suicide: Evolutionary Continuity Or Anthropomorphism?, Antonio Preti Jan 2018

Animal Suicide: Evolutionary Continuity Or Anthropomorphism?, Antonio Preti

Animal Sentience

Evolutionary processes are characterized by both continuity and discontinuity. Evidence on suicide in nonhuman animals is faint and often rests on the metaphorical or anthropomorphic use of the term. Suicidal behavior might be an evolutionary jump relatively recent in our species: a byproduct of living in groups of people who are not as closely related genetically as in social groups of nonhuman mammals.

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Roots Of Self-Preservation Failure In Animal Behavior, Denys Decatanzaro Jan 2018

Roots Of Self-Preservation Failure In Animal Behavior, Denys Decatanzaro

Animal Sentience

Affective variation from euphoria to dysphoria, best understood in humans, is also found in the behavior and neurochemistry of many other mammals. Suicide in humans typically occurs in highly dysphoric and despondent individuals. Self-injurious behavior has been observed in dysphoric and despondent nonhuman primates. In humans, suicide is facilitated by a highly-evolved neocortex giving rise to behavioral flexibility and culture. As Peña-Guzmán indicates, some other mammals also have elaborate neocortices and the capacity for cognitive insight, particularly apes, delphinids, and whales. Suicide is most likely to occur in species where individuals live in stable groups of highly interdependent kin.

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Animal Suicide And "Anthropodenial", Ryan Hediger Jan 2018

Animal Suicide And "Anthropodenial", Ryan Hediger

Animal Sentience

Increasing understanding of the impressive cognitive and social capacities of nonhuman animals suggests the possibility that they may sometimes commit suicide. Such notions tend to be dismissed as “anthropomorphism.” That interpretive hazard, I argue, must be weighed against the opposite hazard of “anthropodenial” — “the a priori rejection of shared characteristics between humans and animals” (de Waal 2006). If animals do commit suicide, how often is it motivated precisely by the impact of humans on animal life?

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