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Ecology

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Articles 61 - 90 of 541

Full-Text Articles in Ecology and Evolutionary Biology

Relationship Between Trunk Cross-Sectional Area Growth And Water Stress In Garry Oaks (Q. Garryana): A Species Of Conservation Concern, John Cochrane May 2024

Relationship Between Trunk Cross-Sectional Area Growth And Water Stress In Garry Oaks (Q. Garryana): A Species Of Conservation Concern, John Cochrane

Student Research Symposium

In the Pacific Northwest, Quercus garryana (Garry oak trees) support over 627 native species. Garry oak natural habitat was originally maintained by indigenous practices but has been reduced to 5-10% of its range due to change in land management strategies. To support oak conservation, we need to understand heat and water affect the growth and physiology of this species. In this study, we created a fixed linear model of trunk cross-sectional growth with water stress (Ψ), photosynthetic water-use efficiency (δ13C), and leaf Carbon-Nitrogen ratio. We collected branch samples from the canopies of 47-64 mature Garry oaks in the …


Causes And Consequences Of Space-Use Behavior Under Predation Risk In A Free-Living System, Brian J. Smith May 2024

Causes And Consequences Of Space-Use Behavior Under Predation Risk In A Free-Living System, Brian J. Smith

All Graduate Theses and Dissertations, Fall 2023 to Present

Predators can have important ecological effects through killing and eating their prey, the so-called consumptive effect, but predators can also have a nonconsumptive effect (NCE) on their prey – this happens when the risk of predation itself causes prey to alter their behaviors or other traits and these alterations ultimately reduce prey survival, reproduction, or population size. While scientists understand the consumptive effects of predators well, we are still unsure whether NCEs are important in free-living systems. In this dissertation, I sought to better understand the potential NCEs of predators (wolves and cougars) on elk in northern Yellowstone National Park …


Ecological Dynamics Of Cenozoic North American Mammals, Alexandria Brandis Shupinski May 2024

Ecological Dynamics Of Cenozoic North American Mammals, Alexandria Brandis Shupinski

Dissertations and Doctoral Documents, University of Nebraska-Lincoln, 2023–

Understanding how community structure is affected by various ecological, climatic, and environmental changes is a long-standing goal in ecology. However, disentangling human impacts from these other variables is difficult when using short modern timescales and anthropogenically altered communities. The fossil record provides a means to investigate community structure dynamics across critical intervals, excluding humans as a variable. Here, in three studies, I assess changes in North American mammal paleocommunity structure across the last 66 million years (Ma) at multiple temporal and spatial scales. In Chapter 1, I identify changes in mammal functional diversity (FD) locally and continentally, using three independent …


Threads Of Connection: An Offering To Re-Tangle Humanity And Nature With The Patterns Of Our World, Emily Shelton May 2024

Threads Of Connection: An Offering To Re-Tangle Humanity And Nature With The Patterns Of Our World, Emily Shelton

Graduate Theses

In our world there are patterns of self-similarity that serve as evidence of the interconnectedness between humankind and the rest of the natural world. They are reflected in our bodies, behaviors, and environments, both natural and manmade, and can be found throughout systems at every scale, micro through macro. These organic, linear motifs branch into smaller iterations that seem to shape our existence on this planet as we gravitate towards experiences that echo these patterns. During everyday acts like shopping in a grocery store or a crowd at a concert, we unconsciously participate in self-similar collective movements as we navigate …


Rescape: Transforming Coral-Reefscape Images For Quantitative Analysis, Zachary Ferris, Eraldo Ribeiro, Tomofumi Nagata, Robert Van Woesik Apr 2024

Rescape: Transforming Coral-Reefscape Images For Quantitative Analysis, Zachary Ferris, Eraldo Ribeiro, Tomofumi Nagata, Robert Van Woesik

Ocean Engineering and Marine Sciences Faculty Publications

Ever since the first image of a coral reef was captured in 1885, people worldwide have been accumulating images of coral reefscapes that document the historic conditions of reefs. However, these innumerable reefscape images suffer from perspective distortion, which reduces the apparent size of distant taxa, rendering the images unusable for quantitative analysis of reef conditions. Here we solve this century-long distortion problem by developing a novel computer-vision algorithm, ReScape, which removes the perspective distortion from reefscape images by transforming them into top-down views, making them usable for quantitative analysis of reef conditions. In doing so, we demonstrate the …


The Descent Of Anolis: Assemblages, Convergence, And Ecomorphological Evolution, Christopher G. Anderson Apr 2024

The Descent Of Anolis: Assemblages, Convergence, And Ecomorphological Evolution, Christopher G. Anderson

Biology ETDs

The evolutionary outcomes we observe in modern organisms, particularly associations between phenotypes and environments, have been and remain an invaluable tool in interpreting the biological phenomena that shape our world. In the following chapters, I leverage a comprehensive phylogenetic and morphological dataset for 351 species comprising a diverse group of arboreal lizards, the Anolis clade, to explore how communities and morphologies evolve. First, I characterized patterns among coexisting lineages of Anolis to reveal a general pattern of phylogenetic relatedness among lineages comprising assemblages and identify differences attributable to a biogeographic variable. Then I examined the morphological consequences associated with the …


A Novel Study Of Cliff Swallow (Petrochelidon Pyrrhonota) Feather Coloration In Relation To Habitat Characteristics, Colony Size, And Body Condition, Sonja Brandt, Medhavi Ambardar Apr 2024

A Novel Study Of Cliff Swallow (Petrochelidon Pyrrhonota) Feather Coloration In Relation To Habitat Characteristics, Colony Size, And Body Condition, Sonja Brandt, Medhavi Ambardar

SACAD: Scholarly Activities

Feather coloration is used for social signaling in many avian species, and can be associated with their ability to live and breed in habitats with high quality resources (Jenkins et al. 2013, Saino et al. 2013). It can signify individual quality, (Saino et al. 2013) and influence mate choice (Bennet et al. 1996). We analyzed different aspects of Cliff Swallow (Petrochelidon pyrrhonota) feather coloration in relation to morphology and habitat characteristics. We measured luminance, hue (theta and phi), and saturation for four different color patches on the swallows. We predicted that individuals in brighter coloration would be able to settle …


The Virginia Academy Of Science: A Scholarly Forum Over Decades For Students Of A Plant Ecologist, Stewart Ware Apr 2024

The Virginia Academy Of Science: A Scholarly Forum Over Decades For Students Of A Plant Ecologist, Stewart Ware

Virginia Journal of Science

As a newcomer to Virginia in 1968, I learned that annual meetings of the Virginia Academy of Science (VAS) were regarded as an excellent place for students to have their first experience giving a talk about their research to fellow scientists, and a place for faculty to get to know other Virginia scientists in their fields. Over the next four decades William and Mary students (undergraduate and graduate) and I were presenters or co-authors of 42 talks or posters at VAS annual meetings, 29 of them based on studies of plant ecology across Virginia. Two were talks by colleagues with …


An Analysis On The Capacity Of Visual Art To Promote Conservation Efforts, Devi Bell Apr 2024

An Analysis On The Capacity Of Visual Art To Promote Conservation Efforts, Devi Bell

Honors Projects

In recent decades, we have become more aware of the ways in which anthropogenic actions are harming Earth’s climate, ecosystems, and overall stability. Scientists generally agree on the urgency of our situation, yet the public may find difficulty in comprehending the multitude of information on it. This study aims to educate the public on biodiversity loss and related ecological issues in Ohio by discussing these topics through the medium of visual art. Five pieces involving different scientific subject matter and art mediums were created and displayed for participants to view. Participants completed two surveys, one before viewing the art and …


Checking For Ticks: Analyzing The Efficiency Of Different Tick Collection Methods, Peter Briggs, Lawson Trimmell, Javier Monzón Mar 2024

Checking For Ticks: Analyzing The Efficiency Of Different Tick Collection Methods, Peter Briggs, Lawson Trimmell, Javier Monzón

Seaver College Research And Scholarly Achievement Symposium

Ticks are the most important arthropod disease vectors in both North America and Europe. Field collection of wild ticks is vital for research on the ecology of vector-borne diseases. Dragging and trapping are the two most common methods for collecting wild ticks. Dragging involves a researcher pulling a canvas through a field to collect ticks searching for a host, while trapping exploits ticks’ natural attraction to carbon dioxide to lure them onto a canvas where they get caught on tape. Our study aimed to evaluate which of these two methods is more effective. We chose six sites across three states, …


Synopsis Of The Genus Cerabilia, André Larochelle, Marie-Claude Larivière Mar 2024

Synopsis Of The Genus Cerabilia, André Larochelle, Marie-Claude Larivière

Insecta Mundi

The genus Cerabilia Laporte de Castelnau (Carabidae: Abacetini) is revised for New Zealand. Thirteen species are recognized.

Seven species are described as new: Cerabilia (Cerabilia) cordata Larochelle and Larivière new species, Cerabilia (Cerabilia) kaihoka Larochelle and Larivière new species, Cerabilia (Cerabilia) laevis Larochelle and Larivière new species, Cerabilia (Cerabilia) motunau Larochelle and Larivière new species, Cerabilia (Cerabilia) rugosa Larochelle and Larivière new species, Cerabilia (Cerabilia) takaka Larochelle and Larivière new species, Cerabilia (Cerabilia) willi Larochelle and Larivière new species. …


Literature Review Nature-Based Art Therapy Exploring Connections And Relationships, Janell Lopez-Curtis Jan 2024

Literature Review Nature-Based Art Therapy Exploring Connections And Relationships, Janell Lopez-Curtis

Expressive Therapies Capstone Theses

Art therapy is a modality used in clinical psychotherapy. It is supported through both quantitative and qualitative research. Branching out from art therapy is nature-based art therapy. This branch of expressive therapies holds the potential to be beneficial as art therapy due to the interconnected access to the scientific fields of ecology, ecopsychology, art therapy, expressive therapies, and other nature-based therapeutic activities; this includes intersectionality in ecological theories such as ecofeminism and deep ecology as well. Through an exploration of literature, this paper will provide definitions and theory-based support through reviewing clinical psychotherapy, evidence-based practices, and art therapy theories. The …


Factors Associated With Acoustic Bat Presence During Spring Emergence In The Appalachian Mountains Of Western Virginia, Emily Kirk Pody Jan 2024

Factors Associated With Acoustic Bat Presence During Spring Emergence In The Appalachian Mountains Of Western Virginia, Emily Kirk Pody

Theses, Dissertations and Capstones

Studies of threats that bats face during hibernation have increased in response to white-nose syndrome (WNS), a fungal disease that has ravaged North American bat populations. However, impacts of WNS on bat ecology during spring emergence, when bats are potentially recovering from infection and allocating resources for reproduction, is relatively understudied. As more bat species become listed at the federal and state level, the need to understand the factors associated with spring emergence is critical for improving conservation guidelines and habitat management practices. Acoustic monitoring is an efficient method for monitoring bat presence for prolonged periods of time, giving biologists …


Environmental Biology Masters Capstone, Antonio Gonzalez-Pita Jan 2024

Environmental Biology Masters Capstone, Antonio Gonzalez-Pita

Regis University Student Publications (comprehensive collection)

Human wildlife interactions (HWI) pose a complex challenge for wildlife managers. Human encroachment into wildlife habitat and the growing number of outdoor recreationists are increasing the frequency of contact and conflict, especially in regions such as the Front Range of Colorado. Geographic information systems (GIS), which use a combination of remote sensing and environmental survey data, allow for predictive spatial analyses of where human wildlife interactions are likely to occur. I used publicly reported observations of moose to create spatial predictive maps in a species distribution model framework. Slope and elevation were shown to be the strongest predictors of HWI, …


Impacts Of Anthropogenic Sedimentation On Shell-Bed Habitats In Lake Tanganyika, Africa, Michael Soreghan, Andrew Cohen, Michael M. Mcglue, Kevin M. Yeager, Emily Ryan, Alison Johns, Ishmael Kimirei Jan 2024

Impacts Of Anthropogenic Sedimentation On Shell-Bed Habitats In Lake Tanganyika, Africa, Michael Soreghan, Andrew Cohen, Michael M. Mcglue, Kevin M. Yeager, Emily Ryan, Alison Johns, Ishmael Kimirei

Earth and Environmental Sciences Faculty Publications

Lake Tanganyika, in central Africa, contains a diverse and endemic fauna under threat from global climate change, overfishing, and nearshore sediment pollution. Previous studies of sediment pollution focused justifiably on impacts along rocky shorelines where diversity is high, but Lake Tanganyika also contains widespread shelly accumulations (shell beds) unprecedented in the modern East African lakes, but where impacts are less con- strained. Here we integrate multiple datasets from three sites along the Tanzanian shoreline to explore how variation in sedimentation rates and sediment quality impacts shell-bed substrate and diversity and abundance of ostracodes and sponges across sites that exhibit varying …


Nekton Community Responses To Living Shoreline Restorations In Alabama, Shelby Kuck, Christopher Grant, Matheus De Barros, Alexandra Rodriguez, Ronald Baker Jan 2024

Nekton Community Responses To Living Shoreline Restorations In Alabama, Shelby Kuck, Christopher Grant, Matheus De Barros, Alexandra Rodriguez, Ronald Baker

Gulf and Caribbean Research

No abstract provided.


Interdisciplinary Learning: How Different Ways Of Thinking Can Fuel Passion And Expand Understanding., Allison Amy Peschek Jan 2024

Interdisciplinary Learning: How Different Ways Of Thinking Can Fuel Passion And Expand Understanding., Allison Amy Peschek

Undergraduate Theses, Professional Papers, and Capstone Artifacts

This Capstone project was created to show the profound experience resulting from an extended range of interdisciplinary learning, and how different methods, when paired together, can enhance a learning experience. This was a personal project of exploration into those different angles, and primarily focused on avenues of logical and creating learning, respectively. These methods have been implemented by this author often in the past, but this capstone will explore this concept further, as there is a lot of potential in combining these different methods and biases in learning, to both further passion and understanding as a learner, but to also …


Are Elaeagnus Umbellata Invasions Limited By Microbial Interactions?, Cody Anderson Leverette Jan 2024

Are Elaeagnus Umbellata Invasions Limited By Microbial Interactions?, Cody Anderson Leverette

Masters Theses

Mutualistic interactions with soil microbes are a key component of plant invasions. Mutualists such as ectomycorrhizal fungi and Rhizobia bacteria often limit invasive plant establishment and performance, but the role of nitrogen-fixing Frankia bacteria in invasions remains largely unexplored. In this study, I quantified the net effects of positive and negative plant-soil feedbacks, including Frankia nodulation, on invasive Elaeagnus umbellata (autumn olive; hereafter AO) seedling performance and traits at varying distances from focal shrubs using live soil and sterilized soil. The expectation was that seedling performance would decrease with further distance from shrubs as Frankia abundance decreased. Contrary to expectations, …


Plasticity In Response To Temperature Magnitude And Variation---From Life History To Thermal Tolerance And Coloration, Austin Rex Hoffman Jan 2024

Plasticity In Response To Temperature Magnitude And Variation---From Life History To Thermal Tolerance And Coloration, Austin Rex Hoffman

University of the Pacific Theses and Dissertations

Warming associated with climate change has many direct and downstream effects on animals. For example, warmer developmental temperatures can improve heat tolerance and reduce insect melanization, which is related to coloration, thermoregulation, immunity, desiccation resistance, and potentially life-history traits, such as investment into reproduction and dispersal. Increased temperature variation is also a feature of climate change, and it may have a larger impact on animals than warming. However, I lack insight into the combined effects of temperature magnitude and variability on important traits of life history, thermal tolerance, and coloration. Thus, I integrated temperature variation into thermal performance using a …


Habitat And Demography Of The Ozark Chinquapin (Castanea Ozarkensis) At Roaring River State Park In Barry County, Missouri, Danielle Evilsizor Dec 2023

Habitat And Demography Of The Ozark Chinquapin (Castanea Ozarkensis) At Roaring River State Park In Barry County, Missouri, Danielle Evilsizor

Electronic Theses & Dissertations

The Ozark chinquapin, Castanea ozarkensis Ashe, is a chestnut tree with a range concentrated in the Interior Highlands of North America. Like other North American members of Castanea, it was reduced from an overstory tree to an understory shrub by the invasive chestnut blight fungus (Cryphonectria parasitica [Murrill] M.E. Barr) during the early 20th century. However, relatively little is known about the habitat of this species or its health and reproductive capability post chestnut blight. Chapter one of this study analyzed the habitat of this species through a random forest species distribution model (SDM) to predict where …


Addressing Water Hyacinth (Pontederia Crassipes) Impacts On Aquatic Biota In Lake Okeechobee, Joseph Salerno Dec 2023

Addressing Water Hyacinth (Pontederia Crassipes) Impacts On Aquatic Biota In Lake Okeechobee, Joseph Salerno

All HCAS Student Capstones, Theses, and Dissertations

The incursion of water hyacinth, Pontederia crassipes in Lake Okeechobee has resulted in management systems to be implemented to reduce the coverage of the invasive macrophyte. Its residence in the Lake Okeechobee ecosystem and the effects it has on organisms in the lake, whether it be positive or harmful is unknown. This study attempted to assess the potential effects that water hyacinth has on aquatic biota in Lake Okeechobee. Biotic data were collected on open water, water hyacinth covered, and native vegetation covered habitats via hook-and-line fishing, electrofishing, baited minnow traps, and the sampling of plant roots over a thirteen-month …


Developing Next-Generation Ecoinformatics Tools For Advancing Global Change Science, Ifeanyi H. Nwigboji Dec 2023

Developing Next-Generation Ecoinformatics Tools For Advancing Global Change Science, Ifeanyi H. Nwigboji

Open Access Theses & Dissertations

Ecosystems are responding to a variety of human-induced, interlinked stressors that have emerged from changing climate, alteration to the global water cycle, sea-level rise, and land use and land cover change, among others. Quantifying these changes and their associated impacts on ecosystems requires a huge amount of long-term data. Due to advances in data collection techniques, such as remote sensing platforms, environmental sensors, synthesized datasets, and various software technologies, the volume and variety of long-term ecological data being collected has tremendously increased. Although there are several complex models and analyses that are increasingly parametrized with data from such sensors, there …


Restoration Strategies For Wetlands In The Arid West: Seeding And Planting Approaches For Lakeshore Ecosystems, Jes Braun Dec 2023

Restoration Strategies For Wetlands In The Arid West: Seeding And Planting Approaches For Lakeshore Ecosystems, Jes Braun

All Graduate Theses and Dissertations, Fall 2023 to Present

Wetlands are widely recognized for their valuable benefits such as providing habitat, improving water quality, and reducing the impacts of flooding. However, wetlands face threats from development, drought, and invasive species. This is particularly apparent in the arid west, where upstream water use and drought make water scarcer and contribute to dramatically changing water levels. Here, I investigated revegetation techniques for lakeshore wetlands, using Utah Lake as a case study. Although recent management efforts have minimized invasive Phragmites cover, the desired plant communities are not returning as quickly as needed, highlighting the need to research restoration techniques. Through my research, …


Species Richness Of Moths In Parks Surrounded By Varying Levels Of ​Urbanization Around Nashville, Tennessee, Maxwell Stone, Allie Bennett Nov 2023

Species Richness Of Moths In Parks Surrounded By Varying Levels Of ​Urbanization Around Nashville, Tennessee, Maxwell Stone, Allie Bennett

Science University Research Symposium (SURS)

Moths can act as indicators of environmental wellness due to their pollution sensitivity and the complexity of biodiversity required to support their life cycles. Urbanization can impact the occurrence of moths in protected green spaces. Higher moth species richness was hypothesized to occur in parks surrounded by more rural areas. Three metropolitan parks were chosen in the Nashville area: downtown, within a suburban neighborhood, and in a rural area. Tree canopy cover and degree of impervious surfaces were assessed for each park and surrounding area using iTreeCanopy. Moths were attracted to a white sheet using a mercury vapor bulb, UV …


Needham’S Girls: A Historical Case Study Of Gender And Mentoring In Entomology, Pat Munday Oct 2023

Needham’S Girls: A Historical Case Study Of Gender And Mentoring In Entomology, Pat Munday

Interdisciplinary Arts and Sciences Articles

James G. Needham (1868-1958) was a professor of entomology at Cornell University from 1906 to 1936, and an active emeritus for about ten years thereafter. As a professor, mentored many women graduate students at Cornell, a group that included twenty-nine who took doctorates. As a scientist, he was a member of an extensive network that included many more women entomologists. These women were located throughout the United States and Canada. Some had been his students at Cornell, some were colleagues with whom he did fieldwork, and others were young women who, even though students at colleges other than Cornell, worked …


The Deep Ecology Movement Sep 2023

The Deep Ecology Movement

The International Journal of Ecopsychology (IJE)

There are two great streams of environmentalism in the latter half of the twentieth century. One stream is reformist, attempting to control some of the worst of the air and water pollution and inefficient land use practices in industrialized nations and to save a few of the remaining pieces of wildlands as "designated wilderness areas." The other stream supports many of the reformist goals but is revolutionary, seeking a new metaphysics, epistemology, cosmology, and environmental ethics of person/planet. This paper is an intellectual archeology of the second of these streams of environmentalism, which I will call deep ecology.


Co-Flowering Community Effects On The Relative Contribution Of Pollen Quantity And Quality Limitation To The Reproductive Success Of Four Clarkia Species, Emma Moore Aug 2023

Co-Flowering Community Effects On The Relative Contribution Of Pollen Quantity And Quality Limitation To The Reproductive Success Of Four Clarkia Species, Emma Moore

Electronic Theses and Dissertations

More than 60% of plant populations experience low pollen availability which limits seed production (pollen limitation). Pollen limitation occurs due to low quantity or quality of pollen delivered to stigmas. Despite its ubiquity in nature, to date we still have limited understanding of the drivers of pollen limitation in plant communities. Plants typically occur in diverse communities where pollinator sharing is common and can influence quality and quantity aspects of pollen limitation. Co-flowering species can attract larger numbers of pollinators or they can compete for pollinators affecting pollen loads on stigmas. Pollinator competition may also lead to higher rates of …


An Ecological Survey Of East Texas Salamanders Across The Camp Tyler Outdoor Field School In Smith County, Texas, Justin C. Hunt Jul 2023

An Ecological Survey Of East Texas Salamanders Across The Camp Tyler Outdoor Field School In Smith County, Texas, Justin C. Hunt

Biology Theses

Amphibians are a unique class of organisms with a very long and storied evolutionary history of survival. Many modern amphibian clades occupy several vital ecological roles within their native freshwater environments. One of these roles, typically includes functioning as an ecological indicator species, whereby the presence of stable and diverse populations of many amphibian species, including salamanders, within a freshwater ecosystem have long been considered ecological indicators of good habitat quality and stable ecosystem health. Similarly, salamanders also function as important members of their local food webs and act as valuable mediators of complex trophic hierarchies to facilitate nutrient cycling …


Monitoring The Effects Of Poultry Waste On Fishes And Macroinvertebrates In The Sabine River, Karley R. Parker Jul 2023

Monitoring The Effects Of Poultry Waste On Fishes And Macroinvertebrates In The Sabine River, Karley R. Parker

Biology Theses

Freshwater is a vital resource that provides life and sustainability for almost all organisms on Earth. It is important to maintain its health and protect it from emerging pollutants that pose a threat to the organisms that use it. Pollution continues to threaten the well-being of the environment’s freshwater sources all around the world that could lead to damaging effects in the future. The Sabine River is a major freshwater resource in the east Texas and western Louisiana areas that provides a habitat for thousands of organisms as well as other domestic uses for humans. In 2019, a waste discharge …


A Foundational Population Genetics Investigation Of The Sexual Systems Of Solanum (Solanaceae) In The Australian Monsoon Tropics Suggests Dioecious Taxa May Benefit From Increased Genetic Admixture Via Obligate Outcrossing, Jason T. Cantley, Ingrid E. Jordon-Thaden, Morgan Roche, Daniel S. Hayes, Stephamie Kate, Christopher T. Martine May 2023

A Foundational Population Genetics Investigation Of The Sexual Systems Of Solanum (Solanaceae) In The Australian Monsoon Tropics Suggests Dioecious Taxa May Benefit From Increased Genetic Admixture Via Obligate Outcrossing, Jason T. Cantley, Ingrid E. Jordon-Thaden, Morgan Roche, Daniel S. Hayes, Stephamie Kate, Christopher T. Martine

Faculty Journal Articles

Solanum section Leptostemonum is an ideal lineage to test the theoretical framework regarding proposed evolutionary benefits of outcrossing sexual systems in comparison to cosexuality. Theoretically, non-cosexual taxa should support more genetic diversity within populations, experience less inbreeding, and have less genetic structure due to a restricted ability to self-fertilize. However, many confounding factors present challenges for a confident inference that inherent differences in sexual systems influence observed genetic patterns among populations. This study provides a foundational baseline of the population genetics of several species of different sexual systems with the aim of generating hypotheses of any factor—including sexual system—that influences …