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Articles 31 - 60 of 1031
Full-Text Articles in Ecology and Evolutionary Biology
Implications Of Spatially Variable Costs And Habitat Conversion Risk In Landscape-Scale Conservation Planning, Max Post Van Der Burg, Neil Chartier, Ryan Drum
Implications Of Spatially Variable Costs And Habitat Conversion Risk In Landscape-Scale Conservation Planning, Max Post Van Der Burg, Neil Chartier, Ryan Drum
United States Geological Survey, Northern Prairie Wildlife Research Center: Publications
‘‘Strategic habitat conservation’’ refers to a process used by the U.S. Fish and Wildlife Service to develop cost-efficient strategies for conserving wildlife populations and their habitats. Strategic habitat conservation focuses on resolving uncertainties surrounding habitat conservation to meet specific wildlife population objectives (i.e., targets) and developing tools to guide where conservation actions should be focused on the landscape. Although there are examples of using optimization models to highlight where conservation should be delivered, such methods often do not explicitly account for spatial variation in the costs of conservation actions. Furthermore, many planning approaches assume that habitat protection is a preferred …
Sociodemographic And Climatic Factors Shaping The Development Of Drought Policies In Major U.S. Cities, Riana S. Gayle
Sociodemographic And Climatic Factors Shaping The Development Of Drought Policies In Major U.S. Cities, Riana S. Gayle
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
In most parts of the world, drought is an inevitable and natural occurrence. However, as the climate continues to warm, and populations grow and expand, the negative impacts of this extreme weather event are predicted to become more pronounced. This leads many communities and stakeholders to question what is being done to prepare society for widespread drought? The following research determines different social and atmospheric characteristics that affect a city’s likelihood of having a drought policy in place.To do this, a thorough search was conducted at the city level to determine where drought policies are currently located in theU.S. The …
An Evaluation Of Bull Trout Movement Dynamics In The Walla Walla River, Courtney Newlon
An Evaluation Of Bull Trout Movement Dynamics In The Walla Walla River, Courtney Newlon
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Bull trout are a fish species listed as threatened under the Endangered Species Act. Historically, they ranged from Northern California at the southernmost extent, into Canada at the northern most extent, and east into Nevada and Montana. Bull trout are highly migratory and require large, unfragmented habitats to persist and are thus highly susceptible to human induced land-use practices. The goal of my thesis was to obtain a better understanding of bull trout movement patterns in the Walla Walla River, Washington using complimentary techniques; Passive Integrated Transponder (PIT) technology and otolith microchemistry. PIT tags can be injected into a fish …
Organic Matter Sources, Composition, And Quality In Rivers And Experimental Streams, Julia E. Kelso
Organic Matter Sources, Composition, And Quality In Rivers And Experimental Streams, Julia E. Kelso
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Organic matter (OM) is often considered the “currency” for ecosystem processes, such as respiration and primary production. OM in aquatic ecosystems is derived from multiple sources, and is a complex mixture of thousands of different chemical constituents. Therefore, it is difficult to identify all the sources of OM that enter and exit aquatic ecosystems. As humans develop undisturbed land, the rate at which terrestrial OM (e.g. soil and plants) and associated nutrients (e.g. nitrogen) enters rivers has increased. Increased nutrients may lead to increased primary production from aquatic plants and algae, potentially causing eutrophication and harmful algal blooms. …
Plant Evolutionary Response To Climate Change: Detecting Adaptation Across Experimental And Natural Precipitation Gradients, Jacqueline J. Peña
Plant Evolutionary Response To Climate Change: Detecting Adaptation Across Experimental And Natural Precipitation Gradients, Jacqueline J. Peña
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Global climate change is a real-time problem that presents threats to many species. Climate change can alter ecosystems and may lead to species extinction. Species can respond to climate change by moving to a better environment or adapting. Therefore, it is necessary to rely on several approaches and perspectives to anticipate ecological impacts of climate change. A common strategy uses models to understand how populations respond to different climate scenarios. Ecological models have helped us understand population persistence, but they often ignore how populations adapt to environmental stress. Adaptive evolution has been ignored because it was assumed that evolution was …
Assessing The Ecological Implications Of The Altered Flow And Sediment Regimes Of The Rio Grande Along The West Texas-Mexico Border, Demitra E. Blythe
Assessing The Ecological Implications Of The Altered Flow And Sediment Regimes Of The Rio Grande Along The West Texas-Mexico Border, Demitra E. Blythe
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Large, exotic (those whose headwaters are in distant places) rivers are some of the most unique and diverse ecosystems on earth. Because they often flow through a multitude of biomes and climates, their waters are a vital resource not only for the organisms that inhabit these rivers, but for human societies as well. Thus, large rivers, like the Rio Grande, that flow through arid and agricultural regions are highly regulated and diverted. Regulation and dewatering upset a river’s natural flow regime (e.g., magnitude, duration, timing of large flood events), subsequently impacting the river’s ability to transport its sediment supply, and …
Geospatial Modeling Of Land Cover Change In The Chocó-Darien Global Ecoregion Of South America: Assessing Proximate Causes And Underlying Drivers Of Deforestation And Reforestation, José Camilo Fagua
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
The Chocó-Darien Global Ecoregion (CGE) in South America is one of 25 global biodiversity hotspots prioritized for conservation. I performed the first land-use and land-cover (LULC) change analysis for the entire CGE in this dissertation. There were three main objectives: 1) Select the best available imagery to build annual land-use and land-cover maps from 2001 to 2015 across the CGE. 2) Model LULC across the CGE to assess forest change trends from 2002 to 2015 and identify the effect of proximate causes of deforestation and reforestation. 3) Estimate the effects of underlying drivers on deforestation and reforestation across the CGE …
The Macroinvertebrate And Fish Communities Of In-Stream Beaver Ponds In Northeastern Utah, Susan Washko
The Macroinvertebrate And Fish Communities Of In-Stream Beaver Ponds In Northeastern Utah, Susan Washko
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Beavers were virtually extirpated from North America during the fur trade, but populations have since recovered. Dams built by recolonizing beaver alter stream habitat by forming deep, slow ponds within the streams. Such changes to the habitat is likely to have consequences for organisms such as macroinvertebrates and fish. The objective of this study was to identify the differences in the macroinvertebrate and trout community in beaver ponds and lotic (e.g. flowing reaches of a stream) reaches in tributaries to the Logan River in northeastern Utah. The macroinvertebrate community of beaver ponds had fewer species, fewer numbers, and lower biomass …
The Nesting Ecology Of Woodpeckers In The Eastern Cascades And Their Interactions With Nest Competitors And Predators, Samuel D. Cowell
The Nesting Ecology Of Woodpeckers In The Eastern Cascades And Their Interactions With Nest Competitors And Predators, Samuel D. Cowell
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Woodpeckers create nesting cavities for other birds and animals in forests. This creates dynamic interactions between both woodpeckers and these other animals. Using video cameras, we documented direct behavioral interactions between nesting woodpeckers and other animals in the Eastern Washington Cascades during the 2015 and 2016 breeding seasons. Additionally, we offered 937 students in a General Biology laboratory course to participate in this original research opportunity and described and the impact the experience had on the participants as well as the researchers.
In 2015, Western Blubebirds took over two active Black-backed Woodpecker nests by physically attacking the woodpeckers. In 2016, …
Efficacy Of Translocation As A Management Tool For Urban Mule Deer In Utah, Channing R. Howard
Efficacy Of Translocation As A Management Tool For Urban Mule Deer In Utah, Channing R. Howard
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
An increase in urbanization in the United States has led to an increase in human-wildlife interactions with deer (Odocoileus spp.) which have been able to adapt and thrive in these urban environments. In Utah, urbanization has occurred along the Wasatch Front which was once traditional mule deer (Odocoileus hemionus) winter range habitat. This urban expansion coupled with an increasing use of these urban areas by mule deer, have led to increasing conflicts with deer. Overabundant urban deer have led to increased concerns over safety from deer-vehicle collisions, and damage to personal property including gardens and landscaping. …
Influence Of Environmental Variables On Survival Rates Of Pronghorn (Antilocapra Americana) Neonates Across Idaho, Brett R. Panting
Influence Of Environmental Variables On Survival Rates Of Pronghorn (Antilocapra Americana) Neonates Across Idaho, Brett R. Panting
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
This study was completed to better understand pronghorn antelope (Antilocapra americana) populations found throughout Idaho. Antelope were studied in three separate and distinct study areas. The Big Desert, Camas Prairie, and Little Lost and Pahsimeroi valleys were all selected as study sites. Idaho Department of Fish and Game (IDFG) is concerned with current pronghorn populations found throughout Idaho. Pronghorn are a valued big game species found in Idaho. Increasing pronghorn populations in Idaho is a focus of IDFG.
We captured and VHF-collared pronghorn fawns found in our three study areas. Fawns were monitored daily with telemetry equipment for …
Landscape Planning For Climate Change Resilience In The Southern Rockies, Jeffrey D. Haight
Landscape Planning For Climate Change Resilience In The Southern Rockies, Jeffrey D. Haight
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
The unique species, ecosystems and landscapes of the Western United States are experiencing unprecedented pressures from climate change, creating new challenges for conservation. As temperatures rise and patterns of precipitation shift, plant and wildlife species have been shifting their ranges to new areas in search of more suitable climates, building groupings of species that are historically unfamiliar. These climate -driven migrations place an additional burden on species that are already threatened from habitat loss and other human-related activities. The impacts of climate change are of particular concern in landscapes that have long been conserved and managed based on the ecological …
Interactions Between Fire Severity And Forest Biota In The Central Sierra Nevada: Formation And Impact Of Small-Scale Fire Refugia And The Effect Of Fire On Forest Structure Predictive Of Fisher (Pekania Pennanti) Den Habitat, Erika M. Blomdahl
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Fire is a natural and essential component of forests in western North America. Fire maintains biodiversity through the creation of different habitat types, and regular fire rotations reduce the accumulation of woody fuels and thick understory plant densities that give rise to catastrophic fire. The practice of fire exclusion has altered western forests and increased the risk of widespread change under rising temperatures projected for the 21st century. To manage for the reintroduction of fire it is critical that we understand the interactions between fire and forest biota in recently fire-suppressed forests.
In Chapter 2, I studied the formation …
Impacts Of Ammonia And Temperature On Freshwater Snail Behavior And Physiology., Megan Christine Dewhatley
Impacts Of Ammonia And Temperature On Freshwater Snail Behavior And Physiology., Megan Christine Dewhatley
Electronic Theses and Dissertations
Gastropods are one of the most imperiled groups of animals in North America, yet there are major gaps in the literature pertaining to pollutants and climate change, and especially sublethal impacts. This dissertation assesses the effects of climate warming and unionized ammonia (NH3), one of the most abundant water pollutants, on the behavior and physiology of two caenogastropod snails: fine-ridged elimia (Elimia semicarinata) and Shawnee rocksnails (Lithasia obovata) (Gastropoda: Neotaenioglossa: Pleuroceridae). Righting behavior, or the movement used by snails to turn themselves right-side-up, was used as the main endpoint; delays in this behavior compromise …
Sandhill Crane Activity In The Central Platte River Valley In Late May And Early June, Jenna M. Malzahn, Andrew J. Caven, Marin Dettweiler, Joshua D. Wiese
Sandhill Crane Activity In The Central Platte River Valley In Late May And Early June, Jenna M. Malzahn, Andrew J. Caven, Marin Dettweiler, Joshua D. Wiese
Nebraska Bird Review
In this report we discuss observations of Sandhill Cranes remaining in the Central Platte River Valley until early June 2018 and discuss potential explanations for this extended stay into the breeding season.
We detected two pairs of adult Sandhill Cranes in two different locations on 15 May 2018 (Table 1). We then detected three injured adult Sandhill Cranes in a third location on 16 May 2018 (Table 1). Two of the three Sandhill Cranes each had a missing leg and the third crane’s leg was broken above the tibiotarsal joint; however, all were still capable of foraging and flight. Later, …
Fall Field Report, August - November 2018, W. Ross Silcock
Fall Field Report, August - November 2018, W. Ross Silcock
Nebraska Bird Review
As far as birds go, this was a relatively quiet season, but, as usual, there was much of interest to chew on. Most important in any report are range changes, major changes in expected numbers or early and late dates, all of which relate directly to the status, positive or negative, of a given species.
Waterfowl in general were only average in numbers; new high counts are rare these days compared with numbers 20 or so years ago. Two winter finches, Common Redpoll and Red Crossbill, that normally arrive in late fall, were virtually unreported. Notable range expansions are under …
Review: Canids Of The World: Wolves, Wild Dogs, Foxes, Jackals, Coyotes, And Their Relatives. José R. Castelló., Jonathan (Jon) Way
Review: Canids Of The World: Wolves, Wild Dogs, Foxes, Jackals, Coyotes, And Their Relatives. José R. Castelló., Jonathan (Jon) Way
The Prairie Naturalist
Canids of the World is one of those reads where just when you think that something cannot be beat, this book comes out. What I mean by that is that I have read many works on wolves and other canids, most recently Nate Blakeslee’s American Wolf (Way 2017), and I have enjoyed and been enthralled with many of them. But Canids of the World may take the cake for being the most impressive book available on canids. For a modest price of just under $30.00, you can own a book that has over 600 amazing high-resolution glossy photographs of every …
New Breeding Record And Location For Wilson’S Phalarope (Phalaropus Tricolor) In The Nebraska Great Plains, Usa, Madison O. Sutton, Nico Arcilla
New Breeding Record And Location For Wilson’S Phalarope (Phalaropus Tricolor) In The Nebraska Great Plains, Usa, Madison O. Sutton, Nico Arcilla
The Prairie Naturalist
Wilson’s phalarope (Phalaropus tricolor; Scolopacidae) is a migratory shorebird that relies on interior wetlands for foraging and breeding (Colwell and Jehl 1994, van Gils et al. 2018). Its global population status is unclear (Colwell and Jehl 1994, Lesterhuis and Clay 2010), and is variously listed as declining (Morrison et al. 2006, van Gils et al. 2018), increasing (Andres 2009, BirdLife International 2018), and exhibiting a long-term decline but recent stability (Sauer et al. 2011, Andres et al. 2012). Its global population estimate of 1.5 million birds has not been updated for 30 years, since 1988 (Colwell and Jehl 1994, Lesterhuis …
Review: Behavior Of The Golden Eagle: An Illustrated Ethogram. David H. Ellis (Illustrated By N. John Schmitt)., Jeremy E. Guinn
Review: Behavior Of The Golden Eagle: An Illustrated Ethogram. David H. Ellis (Illustrated By N. John Schmitt)., Jeremy E. Guinn
The Prairie Naturalist
Author David H. Ellis and illustrator N. John Schmitt deliver precisely what is promised in Behavior of the Golden Eagle: An Illustrated Ethogram. This “little volume,” as the author coins it, represents a single source for defining, identifying, and describing behaviors of the Golden Eagle (Aquila chrysaetos). With the inclusion of Schmitt’s exceptional drawings, the book is a piece of art, as well as the most useful manual describing Golden Eagle behaviors. Nearly four decades earlier, Ellis (1979) authored the very first Golden Eagle ethogram—the set of repeated standard behaviors for a species called action patterns—and in his new book, …
Review: Edges And Rushes Of Minnesota: The Complete Guide To Species Identification, Welby R. Smith; Photography By Richard Haug, Edward S. Dekeyser
Review: Edges And Rushes Of Minnesota: The Complete Guide To Species Identification, Welby R. Smith; Photography By Richard Haug, Edward S. Dekeyser
The Prairie Naturalist
Even the most seasoned individual with a plant identification background can relate to the difficulty of identifying sedges and rushes to the species level. Historically, one has had to rely on dichotomous keys to identify a sedge or rush species in the field. After hours of frustration, a person ends up collecting the plant and, if lucky, bringing the collection back to a herbarium where it can be compared to known specimens. I have been collecting and identifying sedge and rush species for over 25 years, and author Welby Smith along with photographer Richard Haug have published what I believe …
Red-Bellied Snake (Storeria Occipitomaculata) Copulation In South Dakota, Drew R. Davis
Red-Bellied Snake (Storeria Occipitomaculata) Copulation In South Dakota, Drew R. Davis
The Prairie Naturalist
Storeria occipitomaculata is a small, terrestrial species of snake that occurs across much of eastern North America (Ernst and Barbour 1989, Ernst 2002, Ernst and Ernst 2003). Due to this widespread distribution, S. occipitomaculata faces varied climates that likely result in regional differences in reproductive phenology. Much of what is known about the reproductive ecology of S. occipitomaculata has been documented from the southeastern portion of its range in North America (South Carolina: Semlitsch and Moran 1984, North Carolina: Willson and Dorcas 2004), though Blanchard (1937) studied a population from northern Michigan. The exact reproductive timing in this …
The Nebraska Bird Review, Volume 86 December 2018 Number 4
The Nebraska Bird Review, Volume 86 December 2018 Number 4
Nebraska Bird Review
Fall Field Report, August - November 2018, by W. Ross Silcock …146-167
Northern Saw-whet Owl (Aegolius acadius) Nest Box Project: The First Seven Years, by Wayne J. Mollhoff …168-174
Sandhill Crane Activity in the Central Platte River Valley in Late May and Early June, by Jenna Malzahn, Andrew J. Caven, Marin Dettweiler, Joshua D. Wiese .... 175-180
2018 Fall Banding Migration Summary, Chadron State Park, by Chris Murray …181-185
Ainsworth Fall Field Days, Sept. 14-16, 2018, by Janis Paseka …186-190
Index to Volume 86 …191-206
Subscription and Organization Information … 207
The Thermal Ecology Of Sceloporus Occidentalis, Luis Patricio Burgos
The Thermal Ecology Of Sceloporus Occidentalis, Luis Patricio Burgos
Master's Theses
With temperatures rising globally, assessing the possible impacts of the changing climate becomes more and more urgent. Ectotherms are excellent indicators of potential climatic ramifications on biodiversity because of their heavy reliance on the environment for their thermoregulation. Studies have historically looked at thermal tolerance values to establish predictive models for population and species extinctions.
In chapter 1, we looked at recent studies that suggest that thermal tolerance may be a plastic trait and test the effects empirically. Most studies are based on captive lizards acclimated to laboratory conditions that do not necessarily reflect natural environments, and if thermal tolerance …
Review: Using Physiologically Based Models To Predict Population Responses To Phytochemicals By Wild Vertebrate Herbivores, J. S. Forbey, T. T. Caughlin
Review: Using Physiologically Based Models To Predict Population Responses To Phytochemicals By Wild Vertebrate Herbivores, J. S. Forbey, T. T. Caughlin
Biology Faculty Publications and Presentations
To understand how foraging decisions impact individual fitness of herbivores, nutritional ecologists must consider the complex in vivo dynamics of nutrient–nutrient interactions and nutrient–toxin interactions associated with foraging. Mathematical modeling has long been used to make foraging predictions (e.g. optimal foraging theory) but has largely been restricted to a single currency (e.g. energy) or using simple indices of nutrition (e.g. fecal nitrogen) without full consideration of physiologically based interactions among numerous co-ingested phytochemicals. Here, we describe a physiologically based model (PBM) that provides a mechanistic link between foraging decisions and demographic consequences. Including physiological mechanisms of absorption, digestion and metabolism …
Relationships Among Biodiversity Dimensions Of Birds In Nebraska, Nadejda Mirochnitchenko
Relationships Among Biodiversity Dimensions Of Birds In Nebraska, Nadejda Mirochnitchenko
School of Natural Resources: Dissertations, Theses, and Student Research
Biological diversity, or biodiversity, is a multi-dimensional concept that can be decomposed to measure information about taxonomic, phylogenetic, and functional variation within communities. Although the dimensions of biodiversity are interrelated, the assumption that measuring one dimension of diversity can inform about patterns in another dimension does not necessarily follow from theory or empirical study. The relationships among biodiversity dimensions is not well understood, nor how differences among dimensions could influence conservation decision making. Using the avian community as a study system, we explored the relationships of breadth metrics from the taxonomic, phylogenetic, and functional dimensions among each other and across …
The Biological Significance And Utility Of Feeding By Dermestes Maculatus, Braymond Adams
The Biological Significance And Utility Of Feeding By Dermestes Maculatus, Braymond Adams
School of Natural Resources: Dissertations, Theses, and Student Research
With their efficient feeding habits and tolerance to very low moisture and humidity, beetles in the Family Dermestidae are especially adapted to variable environments and habitats. Dermestid cultures have been in use since 1922 in cleaning tissue and flesh from bones, and proven benefit in multiple fields, including zoology, ornithology, and forensics. Dermestid feeding behaviors when coupled with known life stage and insect succession information aids in providing significant entomological evidence. However, the feeding activities of insects, like those of vertebrate scavengers and predators, change remains and may leave artifacts that can be sometimes be difficult to assign to a …
The Effects Of Diet And Mating System On Reproductive (And Post‐Reproductive) Life Span In A Freshwater Snail, Josh R. Auld
The Effects Of Diet And Mating System On Reproductive (And Post‐Reproductive) Life Span In A Freshwater Snail, Josh R. Auld
Biology Faculty Publications
The length of the reproductive life span, along with the number/frequency/magnitude of reproductive events, quantifies an individual’s potential contribution to the next generation. By examining reproductive life span, and distinguishing it from somatic life span, we gain insight into critical aspects of an individual’s potential fitness as well as reproductive and somatic senescence. Additionally, differentiating somatic and reproductive life spans can provide insight into the existence of a post‐reproductive period and factors that shape its duration. Given the known importance of diet and mating system on resource allocation, I reared individual freshwater snails (Physa acuta) from 22 full‐sib families under …
Patterns Of Genetic Structure In The American Kestrel (Falco Sparverius): Influence Of Distance And Migration And Implications For Monitoring And Management, Michaela Brinkmeyer
Patterns Of Genetic Structure In The American Kestrel (Falco Sparverius): Influence Of Distance And Migration And Implications For Monitoring And Management, Michaela Brinkmeyer
Boise State University Theses and Dissertations
Genetic structure is useful for inferring patterns of selection, gene flow and connectivity, and can define management units that aid in interpretation of spatially-specific trends and species management. American kestrels (Falco sparverius) are a widespread, generalist species with fully migratory, partial migrant, and resident populations. In many parts of their range, kestrels show evidence of declining population trends; however, it has been difficult to identify threats to kestrels because of differences in regional trends. We used a genome-wide sequencing approach to investigate the genetic structure of American kestrels, test hypotheses about the processes that influence genetic structuring of …
Safety, Stephen M. Vantassel, Brenda K. Osthus
Safety, Stephen M. Vantassel, Brenda K. Osthus
Wildlife Damage Management Technical Series
Wildlife damage management (WDM) is an exciting field with many opportunities to provide solutions to the complex issues involved in human-wildlife interactions. In addition, WDM wildlife control operators (WCO) face a variety of threats to their physical well-being. Injuries can result from misused (Figure 1), faulty, or poorly maintained equipment, inexperience, mishandled wildlife, harsh weather, and dangerous situations, such as electrical lines. The goals of this publication are to: * Develop an awareness of safety issues and adopt a mindset of “Safety First”, * Review the major safety threats that WCOs face, * Provide basic information for WCOs to protect …
Patterns Of Authorship In Ecology And Evolution: First, Last, And Corresponding Authorship Vary With Gender And Geography, Charles W. Fox, Josiah P. Ritchey, C. E. Timothy Paine
Patterns Of Authorship In Ecology And Evolution: First, Last, And Corresponding Authorship Vary With Gender And Geography, Charles W. Fox, Josiah P. Ritchey, C. E. Timothy Paine
Entomology Faculty Publications
The position of an author on the byline of a paper affects the inferences readers make about their contributions to the research. We examine gender differences in authorship in the ecology literature using two datasets: submissions to six journals between 2010 and 2015 (regardless of whether they were accepted), and manuscripts published by 151 journals between 2009 and 2015. Women were less likely to be last (i.e., “senior”) authors (averaging ~23% across journals, years, and datasets) and sole authors (~24%), but more likely to be first author (~38%), relative to their overall frequency of authorship (~31%). However, the proportion of …