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Articles 31 - 60 of 1041
Full-Text Articles in Animal Sciences
Dialect Use Within A Socially Fluid Group Of Southern Resident Killer Whales, Orcinus Orca, Courtney Elizabeth Smith
Dialect Use Within A Socially Fluid Group Of Southern Resident Killer Whales, Orcinus Orca, Courtney Elizabeth Smith
Master's Theses
Resident killer whales, Orcinus orca, of the Northeastern Pacific form stable kinship-based matrifocal associations and communicate with group-specific repertoires of discrete calls (dialects) that reflect these associations. The gradual fission of matrilines is usually consistent with dialect variations among groups that may manifest as differences in call usage at the repertoire level or subtle structural differences of the calls themselves. Therefore, matrilines that are more closely related tend to be more acoustically similar. Within the endangered community of Southern Resident killer whales (SRKWs), recent evidence shows that one particular group (L pod) exhibits the lowest rate of intrapod association …
Holdrege Fall Field Days, Janis Paseka
Holdrege Fall Field Days, Janis Paseka
Nebraska Bird Review
The 2014 NOU Fall Field Days took place in Holdrege at the Super 8 Motel on Sept. 19–21. There were about 35 in attendance at the meeting, which was organized by Dave Heidt and Betty Grenon.
On Friday evening Dave Heidt led a humor-filled discussion on bird songs and calls and ways to remember them, as audience members suggested their own mnemonic devices. On Saturday evening, T. J. Walker gave a presentation on the Nebraska Game and Parks' program of habitat rehabilitation to benefit Ring-necked Pheasants and other species. Field trip destinations on Saturday and on Sunday morning included Harlan …
Summary Of Raptor Banding Efforts At Hitchcock Nature Center, Fall 2014, Jerry Toll
Summary Of Raptor Banding Efforts At Hitchcock Nature Center, Fall 2014, Jerry Toll
Nebraska Bird Review
The 2014 season was by far the most unusual and frustrating season since the inception of raptor banding and hawk watch at the Hitchcock Nature Center in Pottawatomie County, Iowa, just north of Crescent. Banding diurnal raptors commenced on September 7th on schedule. The month of September was typical as far as capture rates and weather systems, although we were stymied somewhat by the lack of availability of European Starlings for lure birds. House Sparrows proved a nearly adequate substitute.
During the peak period of October, weather systems that drive diurnal raptor migration worked against monitoring efforts by forcing migration …
Fall Field Report, August–November 2014, W. Ross Silcock
Fall Field Report, August–November 2014, W. Ross Silcock
Nebraska Bird Review
My first paragraph from last year's fall report read as follows: "In most ways, this was a routine fall season. Most interesting from an ongoing point of view were quite a large number of sightings of passerines at rather late dates, notably among Empidonax flycatchers, wood warblers, native sparrows, tanagers, towhees, grosbeaks, and buntings." This fall was no different; the trend seems to be that final sighting dates for many species are pushing against their latest expected dates.
There were several tidbits of interest, more or less unrelated, but perhaps most notable takeaways for the season were the number of …
First Documented Nest Of Northern Saw-Whet Owl In Nebraska, Wayne J. Mollhoff
First Documented Nest Of Northern Saw-Whet Owl In Nebraska, Wayne J. Mollhoff
Nebraska Bird Review
Northern Saw-whet Owl was added to our avifauna with a list prepared by Taylor and Van Vleet (1889). The first suggestion of breeding was a second-hand report included in a paper presented at the first meeting of the Nebraska Ornithologists' Union in 1899 by M. A. Carriker, Jr., who "was told of a set of five eggs collected near Nebraska City about seven years ago" (1900), but which included no other details. At the next meeting, Bruner (1901) cited that report to include the species on a list of birds that breed in the state. Bruner, Swenk, and Walcott later …
Index To Volume 82
Nebraska Bird Review
Abegglen, Jeff 192
Abraham, K. F. 85
Aderman, T. 80
Addy, Darren 47
Adrian, Brad 47
Ahlschwede, Carla 40
Akers, Danny 100
Alberts: Byron 143; Deb 143
Alisaukis, R. T. 85
Allen: Edward 3, 48, 143, 153; Jan 48
Allison, Scott 49
American Ornithologists' Union 192
. . .
Wright, Rick 49
Wylie, Bruce 41
Yantachka, Jennifer 143
Yellowlegs: Greater 55, 88, 108, 160, 183; Lesser 56, 88, 108, 160, 183
Yellowtbroat, Common 26, 67, 91, 117, 175, 184
Zahurones, Penny 41
Ziolkowski, D. J. 80
In Memory Of Margaret Triplett
In Memory Of Margaret Triplett
Nebraska Bird Review
The NOU lost a longtime member in Kearney on August 7, 2014, when Margaret Sealing Triplett died.
Nebraska Bird Review (December 2014) 82(4), Whole Issue
Nebraska Bird Review (December 2014) 82(4), Whole Issue
Nebraska Bird Review
Fall Field Report, August–November 2014 ... 150
Holdrege Fall Field Days … 181
Summary of Raptor Banding Efforts at Hitchcock Nature Center, Fall 2014 ... 186
First Documented Nest of Northern Saw-whet Owl in Nebraska ... 189
In Memory of Margaret Triplett … 194
Index to Volume 82 … 195
Subscription and Organization Information ... 207
Effects Of Feeding Distillers Grains Throughout A Yearling Heifer Beef Production System On Meat Quality Attributes And Effects Of Omnigen-Af With Ractopamine Hydrochloride On Animal Performance And Beef Carcass Characteristics Of Feedlot Steers, Kelby M. Sudbeck
Department of Animal Science: Dissertations, Theses, and Student Research
Heifers (n = 229; 225 kg ± 2) were used in a 2 x 2 x 2 factorial to determine the effects of longterm exposure to corn distillers grains plus solubles (DGS) on meat quality characteristics. Factors included supplementing 0.91 kg (LW) or 2.3 kg (HW) modified DGS (MDGS) while grazing corn stalks; 0.6% BW dry DGS (SS) daily or none (NS) during summer grazing; and finishing diet containing 40% wet corn gluten feed (CGF) or 40% MDGS (DGF) on a DM basis. An interaction was observed within CGF for discoloration, with SS increasing discoloration 30%; but no differences between …
A Study Of The American Black Bear (Ursus Americanus) In Utah: An Analysis Of The Post-Denning Activities And Bear-Human Conflict, Julie Ann Miller
A Study Of The American Black Bear (Ursus Americanus) In Utah: An Analysis Of The Post-Denning Activities And Bear-Human Conflict, Julie Ann Miller
Theses and Dissertations
This study examined two different aspects of black bear (Ursus americanus) ecology in Utah. First, we determined the post-denning behaviors of female black bears in order to help management agencies protect bears from human disturbances as well as set spring hunts that minimize the taking of females with dependent young. We looked at the timing of den emergence (X = 25 March), the number of days at the den site post emergence (X = 11 days), and departure (X = 8 April) for female black bears in Utah from 2011—2013. We also analyzed the effects of cohort (lone …
Measuring The Role Of Inhibition In Auditory Discrimination In Mice, Tazima Nur
Measuring The Role Of Inhibition In Auditory Discrimination In Mice, Tazima Nur
Graduate Theses and Dissertations
Understanding how inhibitory neurons affect sensory information processing in the cerebral cortex is an ongoing goal of both neuroscience and statistical physics research. In this master's thesis research project, an experimental system has been designed and built for studying how auditory dynamic range depends on inhibitory neurons, based on observations of mouse behavior. In this thesis, firstly, the topic of inhibition and information processing has been introduced. Then two papers related to inhibition and dynamic range has been reviewed in detail. One of the papers is an experimental work that analyzes the affect of inhibition on dynamic range. The other …
Evaluation Of Low-Tannin Grain Sorghum In Broiler Chicken Diets, Samantha Aniecia Shelton
Evaluation Of Low-Tannin Grain Sorghum In Broiler Chicken Diets, Samantha Aniecia Shelton
Graduate Theses and Dissertations
Research trials were conducted to evaluate the effects of different levels of dietary grain sorghum on broiler live performance, carcass yield and shank (leg) and breast skin coloring. Iso-caloric diets were formulated where sorghum replaced corn at rates of 0% (control), 20, 40, 60, 80 and 100% for a total of 6 diets. For each of the two trials, 1500 Cobb 500 male broiler chicks were randomly allocated to 60 pens with 25 birds per pen (10 pens/diet) and grown to 46 days for the first research trial and 41 days for the second. There were no differences (P>.05) …
The Impacts Of Feral Horses On The Use Of Water By Pronghorn On The Sheldon National Wildlife Refuge, Nevada, Amy Marie Gooch
The Impacts Of Feral Horses On The Use Of Water By Pronghorn On The Sheldon National Wildlife Refuge, Nevada, Amy Marie Gooch
Theses and Dissertations
Feral horses occupy 31.6 million acres throughout western North America. Feral horses share similar habitats with a wide range of animal species, including pronghorn. Since horses are larger and often more aggressive than other animals of this region, they are generally socially dominant over all other native ungulate species. Pronghorn share water sources with horses in areas where both occur. In situations where horses exclude pronghorn from water, pronghorn fitness may be impaired, especially during the hottest months of the year when water is limited. The purpose of this study was to investigate interference competition between pronghorn and feral horses …
A Fish Farmer’S Role In Sustainable Aquaculture: An Overview Of Philippine Aquaculture, Janice A. Ragaza, Jonni Fay C. Teves
A Fish Farmer’S Role In Sustainable Aquaculture: An Overview Of Philippine Aquaculture, Janice A. Ragaza, Jonni Fay C. Teves
Biology Faculty Publications
The expansion of Philippine aquaculture is essential to addressing poverty and food insecurity in the midst of a seemingly irreversible decline in national capture fisheries. Concomitant with its rise, however, are compounding environmental problems caused by climate change, lackluster governance and irresponsible farming practices. It is equally important to ensure aquaculture sustainability so that fish demand from a rapidly growing population is continually met and that fish farmers’ socioeconomic well-being is secured. As such, programs have been developed that aim to enjoin the government and private sectors with fishing communities to realize this goal.
Earthworm, Microbial Biomass, And Leaf Litter Decay Responses After Invasive Honeysuckle Shrub Removal From Urban Woodlands., Robert Preston Pipal
Earthworm, Microbial Biomass, And Leaf Litter Decay Responses After Invasive Honeysuckle Shrub Removal From Urban Woodlands., Robert Preston Pipal
Electronic Theses and Dissertations
Invasive species are one of the leading causes of biodiversity loss and their presence can significantly impact the structure and function of native ecosystems. In this dissertation, the impacts of the invasive honeysuckle shrub, Lonicera maackii, on exotic earthworm populations, leaf litter decay responses, and inorganic and organic nutrient pools in an urban woodland park are assessed. Chapter 1 provides a brief review of honeysuckle's effects on forest ecosystems. Chapter 2 describes honeysuckle’s effects on exotic earthworm populations and the seasonal importance of macroinvertebrates to leaf litter decomposition in urban woodlands. This study revealed that L. maackii promoted higher exotic …
Effects Of Step Training On The Kidneys Following Spinal Cord Injury In Rats., Pradeepa Poudyal
Effects Of Step Training On The Kidneys Following Spinal Cord Injury In Rats., Pradeepa Poudyal
Electronic Theses and Dissertations
Detrusor-sphincter dyssynergia after chronic spinal cord injury (SCI) leads to over-distention, high bladder pressures and vesico-ureteric reflux into the kidney, resulting in repeated infections and pyelonephritis. Given the importance/limited knowledge of SCI effects on the upper urinary tract, this study assessed expression of two proteins in the kidneys of contused (T9) male rats relative to controls that are indicative of tissue stress/damage (TGFβ, CD11b). The impact of two therapeutic rehabilitation strategies, 60 minutes of step training and general exercise, was then assessed on these protein levels relative to non-trained SCI controls, plus the effectiveness of a shorter training duration (30 …
Effects Of Stocking Rate On Forage Nutrient Composition Of Nebraska Sandhills Upland Range When Grazed In Early Summer And The Effects Of Grazing On Nebraska Sandhills Meadow Forage Nutrient Composition, Jared Vern Judy
Department of Animal Science: Dissertations, Theses, and Student Research
The objectives of this research were to 1) evaluate the effects of stocking rate on forage nutrient quality 2) quantify the relative proportions of current vs. previous year growth being consumed in early summer upland range pastures and 3) determine how grazing effects forage nutrient quality in subirrigated meadows in the Nebraska Sandhills. Experiment 1 was a two year study conducted on the experimental upland range paddocks at Gudmundsen Sandhills Laboratory. Twelve 2-hectare paddocks were assigned one of three treatments stocked at 0 (control), 0.57 (light), and 0.85 (heavy) AUM/ha. Ten 0.25 m2 quadrats were clipped per paddock during …
Genetic Parameter Estimates And Breed Effects For Calving Difficulty And Birth Weight In A Multi-Breed Population, Cashley Ahlberg
Genetic Parameter Estimates And Breed Effects For Calving Difficulty And Birth Weight In A Multi-Breed Population, Cashley Ahlberg
Department of Animal Science: Dissertations, Theses, and Student Research
There are multiple breeds of beef cattle available to utilize in breeding systems to maximize production and economics. Calving difficulty (dystocia) is a significant cost to beef production and is more prevalent in first-calf heifers. The objectives of this study were to estimate genetic parameters and breed differences for calving difficulty and birth weight as a first step towards the development of across-breed adjustment factors for calving difficulty.
Two models were employed to analyze birth weight (BWT) and calving difficulty (CD) recorded on 4,579 first parity females from the Germplasm Evaluation program at the United States Meat Animal Research Center …
Changes In Corn Residue Quality Throughout The Grazing Period And Effect Of Supplementation Of Calves Grazing Corn Residue, Amanda J. Burken
Changes In Corn Residue Quality Throughout The Grazing Period And Effect Of Supplementation Of Calves Grazing Corn Residue, Amanda J. Burken
Department of Animal Science: Dissertations, Theses, and Student Research
Corn residue is an abundant feed source in Nebraska that can be utilized as an alternative winter feed. Calves were backgrounded on corn residue in order to determine gain and estimate forage intake when supplemented with distillers grains (DGS). Calves grazing the non-irrigated field gained more (1.03 kg/calf daily) when compared to those grazing the irrigated field (0.90 kg/calf daily; P < 0.01). In year 1, a quadratic effect for intake of DGS was present (P < 0.01) while year 2 observed a linear effect for increasing level of DGS (P < 0.01). The nutritional quality of corn residue was evaluated over time in order to determine changes in blade/sheath, cob, husk/shank and stem. Minimal changes in DM of the forage components occurred was grain reached 15.5% moisture. Digestibility of the blade/sheath declined linearly over time (P < 0.01) while the husk remained constant (P = 0.40). Cob digestibility decreased quadratically (P < 0.01) throughout the sampling period with few changes once grain reached 15.5% moisture. Differences observed in the digestibility of the blade/sheath were attributed to the effects of weathering. A third set of trials was conducted to evaluate the effects of by-product supplementation of calves grazing irrigated corn residue and supplemented with DGS or continuous access to lick tubs. The DGS treatment gained more (0.62 kg/calf daily) than the lick tub treatment (0.38 kg/calf daily; P < 0.01). Calves offered DGS consumed more supplement as a percentage of BW (0.52%) when compared with calves offered lick tubs (0.36%; P < 0.01) on a DM basis. Calves supplemented with DGS had a higher supplement efficiency (46.3% to 42.9%, DM basis) although no differences were present between treatments (P = 0.49). When analyzed on an OM basis, however, calves offered lick tubs had a numerically higher supplement efficiency (50.4%) in comparison to calves …
Feedlot Manure Management Considerations Including Anaerobic Digestion Potential And Mineral Retention, Andrea K. Watson
Feedlot Manure Management Considerations Including Anaerobic Digestion Potential And Mineral Retention, Andrea K. Watson
Department of Animal Science: Dissertations, Theses, and Student Research
Three anaerobic digestion trials were conducted to evaluate methane production from feedlot manure. As organic matter content of manure decreased, degradation of manure and methane production also linearly decreased (P ≤ 0.02). Quality, or organic matter content of manure, had a greater impact on anaerobic digestion than diet (with and without distillers grains) fed to cattle. Minerals are concentrated in effluent removed from anaerobic digesters.
Knowing mineral retention within cattle allows for more accurate calculation of mineral excretion in manure. Three serial slaughter trials measured mineral retention in growing and finishing cattle. During a growing phase, beef cattle gaining …
The Effects Of Conjugated Linoleic Acid Supplementation On Lipopolysaccharide-Challenged Synoviocytes, Allison Millican
The Effects Of Conjugated Linoleic Acid Supplementation On Lipopolysaccharide-Challenged Synoviocytes, Allison Millican
All Theses
Osteoarthritis is one of the leading causes of lameness in the athletic horse. Cases of equine osteoarthritis typically present with synovitis, or inflammation of the synovium. This inflammation in the synovium leads to the production of inflammatory cytokines and eicosanoids. Although these downstream products can be beneficial, overproduction can lead to degradation of cartilage and joint disease. Opportunities to help mitigate or decrease inflammation within the joint are of current interest to the equine industry. One method being investigated is nutritional supplementation. The anti-inflammatory effects demonstrated by CLA supplementation make it a potential prospect for utilization in mitigating joint inflammation. …
Protein Turnover In Broiler, Layers, And Broiler Breeders, Karen Vignale Lake
Protein Turnover In Broiler, Layers, And Broiler Breeders, Karen Vignale Lake
Graduate Theses and Dissertations
Protein turnover in skeletal tissue for broiler breeders has shown to increase at sexual maturity and then decline with increased egg production. Regarding broiler chickens the industry is facing two situations that need to be clarified: the appearance of white stripes in birds over 56 days of age and the positive effect in breast meat yield that supplementation of 25OH-D3 is causing. A series of studies were conducted to determine the effect of sexual maturity in protein turnover in broiler breeders and laying hens and to understand the effect of 25OHD3 in breast meat yield in broiler chickens and why …
Evaluation Of Phytate And Phytase Interactions And Phytase Phase-Feeding On Bird Performance, Bone Characteristics And Meat Quality In Young Growing Broilers, Chance L. Williams
Evaluation Of Phytate And Phytase Interactions And Phytase Phase-Feeding On Bird Performance, Bone Characteristics And Meat Quality In Young Growing Broilers, Chance L. Williams
Graduate Theses and Dissertations
Two trials were performed using one-day-old male Cobb x Cobb 500 broilers to determine how dietary phytate and phytase levels as well as phytase phase feeding impacted bird performance parameters, tibia characteristics, and malonaldehyde (MDA) content of the liver, breast and thigh tissues. The first experiment consisted of 1,008 birds randomly placed in 48 floor pens within two commercial broiler houses at the Applied Broiler Research Farm (ABRF; 21 birds per pen; 0.76 ft2 per bird). A 2 X 3 factorial design was used with two levels of dietary phytate (0.21 and 0.31 %) and three levels of phytase supplementation …
Identification And The Significance Of Selective Proteins In Bile And Plasma Of Normal And Health-Compromised Chickens, Balamurugan Packialakshmi
Identification And The Significance Of Selective Proteins In Bile And Plasma Of Normal And Health-Compromised Chickens, Balamurugan Packialakshmi
Graduate Theses and Dissertations
During the last 50 years, animal breeding programs in commercial poultry have made significant progress in the bodyweight gain of broilers but led to several metabolic and skeletal disorders. Lameness associated with proximal femur known as femoral head separation (FHS) or femoral head necrosis (FHN) is one of the major metabolic disorders in poultry industry. In order to select for healthy chickens, markers that can distinguish between healthy and affected birds are required. Biomarkers from blood represent an ideal and rich source of markers which can be obtained using minimally invasive methods. The biomarkers were explored in an experimental model …
Examining Predation As A Possible Means Of Controlling Crown-Of- Thorns Starfish (Acanthaster Planci) Outbreaks On Reefs Around Lizard Island, Australia, Amanda Chan
Independent Study Project (ISP) Collection
Since the world’s coral reefs are currently threatened by a variety of different natural and anthropogenic factors, research on protecting coral reefs is pivotal to protect these diverse ecosystems. However, only Indo-Pacific reefs such as the Great Barrier Reef are dying due to a corallivorous echinoderm threat known as the Crown-of-Thorns Starfish (Acanthaster planci). A. planci is a starfish which feeds on coral tissue and can quickly reduce coral cover on a reef during an outbreak. Although scientists are still unsure as to what causes these outbreaks, one suggestion is the predator-removal theory. The predator removal-theory states that major predators …
Managing Devil Facial Tumour Disease In Tasmanian Devils (Sarcophilus Harrisii): An Investigation Of Heat Shock Proteins As Potential Vaccine Adjuvants, Monika Payerhin
Managing Devil Facial Tumour Disease In Tasmanian Devils (Sarcophilus Harrisii): An Investigation Of Heat Shock Proteins As Potential Vaccine Adjuvants, Monika Payerhin
Independent Study Project (ISP) Collection
The world’s largest carnivorous marsupial, the Tasmanian devil (Sarcophilus harrisii), is facing extinction from a deadly, highly communicable cancer that has already decimated over 85% of devil populations in the wild: devil facial tumour disease (DFTD). DFTD cells effectively evade recognition by the immune system, and every devil that contracts the disease dies from it. Many attempts have been made at developing a vaccine that could help save this now-threatened species. Heat shock proteins have been linked to enhanced immune recognition of pathogens, making them potential candidates for acting as adjuvants to such a vaccine against DFTD. In this study, …
New Study Of Altitude And Butterfly Diversity Evaluation Of Butterfly Diversity In La Hesperia And Influence Of Altitude On Diversity, Savannah Artusi
New Study Of Altitude And Butterfly Diversity Evaluation Of Butterfly Diversity In La Hesperia And Influence Of Altitude On Diversity, Savannah Artusi
Independent Study Project (ISP) Collection
Due to the increasing effects of climate change, studies focusing on bioindicator species are becoming more necessary than ever. Additionally, knowledge about global biodiversity can be very useful to conservation organizations because it helps them determine what areas need to be conserved the most .Butterflies are useful as bioindicators due to their complex life cycles, importance in the food chain, and sensitivity to environmental changes. This project focused on studying the overall biodiversity of butterflies in the La Hesperia reserve. In addition, this study aimed to determine how butterfly diversity varies at different altitudes. To do this, the reserve was …
Methane Production By A Packed-Bed Anaerobic Digester Fed Dairy Barn Flush Water, Sean Richard Thomson
Methane Production By A Packed-Bed Anaerobic Digester Fed Dairy Barn Flush Water, Sean Richard Thomson
Master's Theses
Packed-bed digesters are an alternative to covered lagoon digesters for methane production and anaerobic treatment of dilute wastewaters such as dairy barn flush water. The physical media of packed-beds retain biofilms, often allowing increased treatment rates. Previous studies have evaluated several types of media for digestion of dilute wastewaters, but cost and media fouling have setback commercial development. A major operational cost has been effluent recirculation pumping.
In the present effort, a novel approach to anaerobic digestion of flush dairy water was developed at pilot-scale: broken walnut shells were used as a low-cost packed-bed medium and effluent recirculation was replaced …
The Effects Of Limestone Particle Size On Bone Health And Performance Of Pullets And Hens In Conventional Cage And Alternative Housing Systems, Pamela E. Eusebio Balcazar
The Effects Of Limestone Particle Size On Bone Health And Performance Of Pullets And Hens In Conventional Cage And Alternative Housing Systems, Pamela E. Eusebio Balcazar
Department of Animal Science: Dissertations, Theses, and Student Research
One of the major welfare issues in cage-free housing systems is bone fracture. The goal of this research was to improve bone health in layers by building better pullet skeletons and to investigate limestone particle size (LPS) effects on bone health in conventional cages and cage-free housing systems. Study 1 was a preliminary study that compared conventional cages with litter floor pens in terms of performance, bone health, and eggshell quality from 33 to 47 wk of age. Caged hens had higher egg production and eggshell strength, and improved feed efficiency, but lower tibia bone mineral content compared to floor …
Energy Value Of De-Oiled Distillers’ Grains Plus Solubles In Beef Cattle Diets, Meredith L. Bremer
Energy Value Of De-Oiled Distillers’ Grains Plus Solubles In Beef Cattle Diets, Meredith L. Bremer
Department of Animal Science: Dissertations, Theses, and Student Research
Ethanol plants are centrifuging off oil from the thin stillage stream as it has added market value currently. The impact of oil removal on cattle performance has been minimally researched. Thus four trials, one growing, one digestion, and two feedlot, were designed to determine the energy and feeding value of de-oiled distillers grains in beef cattle diets.
In growing cattle diets, diet concentration of modified distillers grains plus solubles (MDGS) impacted cattle performance more than oil content. Ending BW, ADG, and G:F did not differ between cattle fed de-oiled or full fat MDGS, however cattle fed de-oiled MDGS had lower …