Paleontology Commons^™

Exploring The Relationships Between Mammalian Functional Trait Distributions And Regional Biomes, With Application To Miocene Paleoecology, Devra Hock-Reid

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Paleoecology relies on understanding relationships between modern animals and their environment. Animals are adapted to niches in their environments, and those physical adaptations, or functional traits, are utilized as proxies to interpret aspects of paleo-ecosystems. Much is known about individual functional traits in extant mammals and their relationship to the environment. Less is known about how multiple functional traits across a community can be utilized for paleoecological interpretations. I develop models utilizing traits in mammalian communities at the biome level. For Chapter 1, I build a model for North American regional biomes using mammalian trait frequencies. I quantify changes in …

The Diatom Dark Ages: Identification Of Mid-Cretaceous Arctic Platform Diatoms From The Basal Transgression Of The Kanguk Formation, Devon Island, Nunavut, Canada, Megan Heins

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

The lower part of the mid-Cretaceous Kanguk Formation (Lower Turonian interval) contains an important paleontological record crucial to the characterization of a poorly known interval of fossil marine diatoms history. Kanguk Formation mudstones are exposed in a ~200 m-thick section on Devon Island, Nunavut, Canadian High Arctic. Diatoms at this location are well-preserved due to shallow burial on this Arctic Platform site. The rock sequence was protected from glacial erosion that removed much of the Cretaceous record by being down-faulted in a linear graben. Study of these well-preserved fossil diatoms allows for a documentation of the assemblage, identification of potentially …

Changes In Mammalian Abundance Through The Eocene-Oligocene Climate Transition In The White River Group Of Nebraska, Usa, Robert Gillham

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Marine records show major cooling during the Eocene-Oligocene Climate Transition (EOCT). Most proxy studies in the White River Group suggest drying across the EOCT, and some suggest cooling. The lower resolution continental record has hindered a direct correlation of the marine climate record to Nebraska. I explore various correlation schemes and what they imply for faunal changes. This study compiles and analyzes data from 4,875 specimens in the University of Nebraska State Museum (UNSM) collection to test the hypothesis that climate change across the Eocene-Oligocene (E-O) boundary caused significant abundance changes in mammals. A series of binning schemes was created. …

A Quantitative Analysis Of Calcareous Nannofossils Across A Late Oligocene Paleolatitude Transect Of The North Atlantic Ocean, William Barrett Clark

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Samples from ODP Sites 926, 628, 563, U1406, 647, and 918, were analyzed quantitatively across a paleolatitude transect of the North Atlantic Ocean to determine the paleolatitudinal distribution of calcareous nannofossils in the Late Oligocene and the effects of that distribution on biostratigraphic resolution. Detrended Correspondence Analysis (DCA), a Temperature index (TI), and the Shannon Diversity Index (H), were used to examine the paleoenvironmental gradients which exerted the most control over the distribution of species and their abundances. The temperature index correlates significantly to the first axis of the DCA, suggesting that thermal controls were the most important factor in …

A Taxon-Free, Multi-Proxy Model For Making Paleoecological Interpretations Of Neogene North American Faunas, Devra G. Hock

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Proxies used for interpreting the paleoecology of extinct vertebrate communities are usually based on modern ecosystems, with many developed from Old World ecosystems. However, because no model is completely taxon-free and phylogenetic influences cannot be entirely discounted, these proxies may not be appropriate for paleoecological interpretations of North American ecosystems. Additionally, many proxies based on modern vertebrate communities exclude small-bodied mammals. Here I explore several new paleoecological models based on the frequency of mammalian traits within three ecological categories: locomotion, diet, and body mass. Since these models are intended for interpreting paleoenvironments occupied by Neogene North American mammals, the data …

Middle Miocene Paleoenvironmental Reconstruction Of The Central Great Plains From Stable Carbon Isotopes In Large Mammals, Willow H. Nguy

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Middle Miocene (18-12 Mya) mammalian faunas of the North American Great Plains contained a much higher diversity of apparent browsers than any modern biome. This has been attributed to greater primary productivity, which may have supported greater browser diversity that commonly corresponds with densely vegetated habitats. However, several lines of proxy evidence suggest that open woodlands or savannas dominated middle Miocene biomes; neither of which support many browsers today. Stable carbon isotopes in mammalian herbivore tooth enamel were used to reconstruct vegetation structure of middle Miocene biomes.

Stable carbon isotopes in C₃ dominated environments reflect vegetation density and herbivores …

Early Miocene Quantitative Calcareous Nannofossil Biostratigraphy From The Tropical Atlantic, Waheed A. Albasrawi

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Quantitative analysis for the Lower Miocene of Ocean Drilling Program Hole 959A from the West African margin was performed to document all the calcareous nannofossil biostratigraphic events present. Combined with data from previous investigations of the Lower Miocene from the tropical Atlantic, this research identifies and tests the viability of markers used in current zonation scheme, identifies alternative markers for age boundaries, and examine statistically the most probable order of event in the Lower Miocene using the Ranking and Scaling method (RASC).

The examination of Hole 959A was performed on a 112 samples. Seven additional sites that collectively span the …

Reconstructing The Paleoecology And Biogeography Of Rhinoceroses (Mammalia: Rhinocerotidae) In The Great Plains Of North America, Leading Up To Their Extinction In The Early Pliocene, Bian Wang

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Members of the family Rhinocerotidae first appeared in the middle Eocene and were one of most successful mammal groups of the Oligocene and Miocene in North America. Their extinction in the early Pliocene has been attributed to several causes, including cooling climate, an expansion of C₄ grasslands, and faunal turnover favoring high-crowned, open habitat-adapted mammalian taxa. This study tests whether the extinction of North American rhinoceroses in the Great Plains was abrupt or gradual by examining changes in their paleogeographic distribution in a series of time-slices through the Barstovian, Clarendonian, and Hemphillian North American land-mammal ages. It further examines …

Upper Albian To Lower Cenomanian Calcareous Nannofossil Biostratigraphy Of The Proto-North Atlantic, Shamar Chin

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Lower Cenomanian calcareous nannofossil assemblages from Deep Sea Drilling Project (DSDP) Sites 137 and 547 in the proto-North Atlantic Ocean were analyzed quantitatively to examine the fidelity of the widely used CC and UC calcareous nannofossil Zonal schemes. Datasets from Ocean Drilling Program (ODP) holes 1050C and 1052E (Blake Nose) and Tanzania Drilling Project (TDP) Site 24 were integrated into this dataset. Four biozones spanning the upper Albian through middle Cenomanian were determined using the method of unitary associations (UA). Data were also used from these sequences to generate a ranking and scaling (RASC) optimum sequence. A new reliability index …

Constraining Neogene Temperature And Precipitation Histories In The Central Great Plains Using The Fossil Record Of Alligator, Evan Whiting

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Most amphibians and reptiles (excluding birds) are poikilothermic; their internal body temperature varies with that of their external environment. This makes them useful as climate proxies, especially when linked to geographic distributions of ambient climate. I evaluate the utility of the extant crocodylian genus Alligator as a paleoclimate proxy for the Central Great Plains (CGP) using species distribution modeling. Alligator is a readily identifiable taxon with a good CGP fossil record during the Neogene (~23–2.6 Ma). Alligator first appeared in the CGP in the late Eocene (~37 Ma), was absent during most of the Oligocene, reappeared in the early Miocene …

Ecology And Morphology Of The Late Miocene Musk Deer, Longirostromeryx Wellsi (Artiodactyla: Moschidae: Blastomerycinae), Katheryn Y. C. Chen

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Longirostromeryx wellsi, one of the latest surviving members of the extinct clade Blastomerycinae (Artiodactyla: Moschidae), possesses highly derived craniodental morphology that deviates from typical musk deer form. Previous work suggests that the unique anatomy of L. wellsi represents adaptations for occupying open savannas. To test this hypothesis I conduct principal components analysis on five postcranial bones of L. wellsi, comparing them to that of several extant ruminant artiodactyls, which are divided among seven habitat categories. These elements are also compared with the postcrania of other blastomerycines. These analyses indicate that L. wellsi anatomy is most similar to that of other …

Climate Effects On Mammalian Body Size And Grassland Composition Inferred From Late Quaternary Grazers In The Great Plains Of North America, Tom Baldvins

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Several mammalian taxa exhibited a diminution during the Pleistocene-Holocene transition. Two hypotheses have been proposed to explain this phenomenon: (1) human hunting and (2) climate change along with its effects on grasslands. I use stable isotopes from a variety of Quaternary grazing mammals to measure grassland changes following the Last Glacial Maximum (LGM) in Nebraska and to develop a climate proxy to compare with mammalian body size. Results indicate that latest Pleistocene equids and mammoths have δ¹³C values indicative of a C₃ or mixed C₃/C₄ diet. However, Holocene bison have greatly elevated δ^{13 …}

Fossil Crocodilians From The High Guajira Peninsula Of Colombia, And The History Of Neogene Crocodilian Diversity In Tropical South America, Jorge W. Moreno-Bernal

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

The greatest diversity of Cenozoic crocodilians occurred during the Miocene in equatorial South America. However, the origin of this high diversity and its relationship to environmental factors are poorly understood. Most described species come from localities assigned to Laventan (13.8-11.8 Ma) and Huayquerian (9.0-6.8 Ma) South American land mammal ages (SALMAS), whereas the record is sparse in the early to middle Miocene and after the latest Miocene and Pliocene. Field research in the Castilletes (early Miocene-Pliocene) Formation in the High Guajira Peninsula of Colombia provides new fossil data on the origin of Neotropical crocodylian diversity. The Castilletes Formation crops out …

Body Size And Species Richness Changes In Glyptosaurinae (Squamata: Anguidae) Through Climatic Transitions Of The North American Cenozoic, Sara Elshafie

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Poikilothermic vertebrates offer excellent climate proxies based on relationships between environment and measurable variables such as body size and species richness. Relationships of these variables in lizards to environmental transitions over long time scales are poorly understood. Here I show that patterns of body size and species richness in a lizard clade, Glyptosaurinae (Squamata: Anguidae), correspond to known histories of paleotemperatures through the Cenozoic of North America. Glyptosaurines have the richest fossil record among North American Cenozoic lizards and exhibit a wide range of skull sizes. In order to estimate body size for glyptosaurines and other fossil anguids, I collected …

Marine Diatom Assemblage Variation Across Pleistocene Glacial-Interglacial Transitions And Neogene Diatom Biostratigraphy Of Site C9001, Nw Pacific Ocean, Marcella K. Purkey

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

In 2006, D/V-Chikyu cruise CK06-06 drilled Hole C9001C at Site C9001 in the Northwest Pacific Ocean, 80 km east of the Shimokita Peninsula, Japan. An existing chronostratigraphic framework provides a continuous glacial-interglacial (GI) climate record from which a diatom record of paleoenvironmental changes was developed across several GI cycles. Species counts, diatom temperature values, calculated sea-surface temperatures (SST) and factor analysis were produced for each sample and calibrated to prior diatom studies in this region. These features were used to characterize and compare interglacial maxima of Marine Isotope Stages (MIS) 1, 5e, 9 and 11 and transitions from the preceding …

Paleoecology Of Nebraska’S Ungulates During The Eocene-Oligocene Climate Transition, Grant S. Boardman

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

The White River Group (WRG) preserves the Eocene-Oligocene climate transition (EOCT), an interval of global cooling and drying during the onset of Antarctic glaciation. In the Great Plains, a shift from forested conditions to drier woodland-savanna biomes is hypothesized to have occurred at this time. I test this hypothesis through the analyses of several paleoenvironmental proxies on the teeth of 12 WRG ungulate species: stable carbon and oxygen isotopes from tooth enamel, and mesowear and microwear texture. The EOCT shift toward more open habitats and lower vegetation density under drying climates should have resulted in an increase in mean carbon …

Plankton Evolution Driven By Paleoceanographic Change: Prediscosphaera From The Mid-Cretaceous In The Western North Atlantic, Kristen L. Mitchell

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Well-preserved nannofossil assemblages in late Albian and Cenomanian hemipelagic sections from Ocean Drilling Program (ODP) Leg 171B preserve a record of the early history and adaptive radiation of the calcareous nannofossil genus Prediscosphaera. There was a significant rise in the abundance of the genus (from an average of 2% in the early late Albian to about 6% in the latest Albian and Cenomanian. This rise, and other shorter term pulses of increased generic abundance, corresponds to black shale deposition episodes. Twelve distinct morphotypes that were derived from the long-ranging Prediscosphera columnata and Prediscosphaera spinosa, are recognized and differentiated …

The Application Of Biostratigraphy And Paleoecology At Southern Ocean Drill Sites To Resolve Early To Middle Miocene Paleoclimatic Events, Ryan Farmer

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

The diatom biostratigraphy and paleoceanography of Ocean Drilling Program (ODP) Site 744 on the Southern Kerguelen Plateau, southern Indian Ocean are documented for the early to middle Miocene to improve chronostratigraphic age control for the Southern Ocean and Antarctic region. Paleoenvironmental fluctuations in the Southern Ocean are inferred from changes in fossil diatom abundance, preservation, and assemblage composition. A robust, new age model for Holes 744A and 744B is constructed using Constrained Optimization (CONOP) model ages for diatom biostratigraphic datum levels and new magnetic polarity data, which enables assessment of a nearly continuous record of paleoenvironmental change from ~20.25 to …

New Stable Isotope Record Of Paleoecological Change In The Late Neogene Of The Western Great Plains From Enamel In Large Mammals, Zachary Kita

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

An expansion of C₄ grasslands occurred between 6 and 8 million years ago in the Great Plains of North America, as evidenced by a marked shift to more ¹³C-enriched carbon isotope compositions from large fossil mammal tooth enamel and paleosol carbonates. Prior to this expansion, habitats were comprised of exclusively C₃ vegetation. To explore this problem I present a compilation of bulk stable carbon (δ¹³C) and oxygen (δ¹⁸O) isotope values from a variety of large mammals from 6 localities that span from the late Clarendonian to the late Blancan in Nebraska. As expected, …

An Analysis Of Anchitherine Equids Across The Eocene–Oligocene Boundary In The White River Group Of The Western Great Plains, David M. Masciale

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Anchitherine horses are a subfamily of equids that are abundantly represented in the late Eocene and early Oligocene of North America. This group has been heavily studied in the past, but important questions still remain. Some studies have focused on the Eocene-Oligocene boundary and have used these equids along with other taxa to study mammalian diet and climate change through this interval. I reexamine two anchitherine genera, Mesohippus and Miohippus, from stratigraphic sequences of the White River Group in western Nebraska and southwestern South Dakota. These sequences span the Chadronian (late Eocene), Orellan (early Oligocene), and Whitneyan (early Oligocene) …