Paleobiology Commons^™

A New Kimballian Peccary From Nebraska, C. Bertrand Schultz, Larry D. Martin

Bulletin of the University of Nebraska State Museum

A new species of Pliocene peccary, Prosthennops (Macrogens) graffhami; is described from the Kimball Formation, Ogallala Group, Frontier County, Nebraska. This new species is the latest in geologic age and most advanced in the genus.

The remains of fossil vertebrates are generally rare in the upper part of the Ogallala group, and some have maintained that Hemphillian faunas such as Coffee Ranch and Smith County, Kansas, represent the latest Ogallala faunas. Later faunas than are typically considered as Hemphillian, containing more advanced forms, occur in the Kimball Formation and deposits of equivalent age. Known faunas which may …

Bears (Ursidae) From The Late Cenozoic Of Nebraska, C. Bertrand Schultz, Larry D. Martin

Bulletin of the University of Nebraska State Museum

A ramus and partial premaxilla establish the presence of a new subspecies of Indarctos in the upper Pliocene (Kimball Formation, Ogallala Group) of Frontier County, Nebraska. An extremely large species of Agriotherium is represented by fragmentary remains from the middle Pliocene (middle part of Ash Hollow Formation, Ogallala Group) of Sherman County, Nebraska.

This study is part of a series of papers dealing primarily with the fauna of the Kimball formation in Nebraska (Barbour 1927, 1929; Barbour and Schultz, 1941; Schultz and Stout, 1948, 1961; Kent 1963, 1967; Tanner, 1967; Short, 1969; Martin and Tate, 1970; Schultz, Schultz, and Martin, …

Scimitar-Toothed Cats, Machairodus And Nimravides, From The Pliocene Of Kansas And Nebraska, Larry D. Martin, C. Bertrand Schultz

Bulletin of the University of Nebraska State Museum

"Machairodus catocopis Cope" is shown to be a pseudaelurin cat belonging to the genus Nimravides Kitts. Nimravides thinobates (Macdonald) is a possible synonym of N. catocopis (Cope). Nimravides is compared with the Eurasian Machairodus-like cat, Dinofelis. Machairodus (Heterofelis) coloradensis is reported from the Kimball Formation, upper Pliocene (Kimballian) of Cheyenne County, Nebraska, and from the upper part of the Ash Hollow Formation, Pliocene (Hemphillian) of Sherman County, Nebraska. The Kimballian form is described as a new subspecies, Machairodus coloradensis tanneri.

The genus Machairodus has long been associated with the Hemphillian of North America and the …

Middle And Late Cenozoic Tapirs From Nebraska, C. Bertrand Schultz, Larry D. Martin, R. George Corner

Bulletin of the University of Nebraska State Museum

The distribution and evolution of Late Cenozoic tapirs are discussed and the forms present in Nebraska are reported. Two new species are described from the Ogallala Pliocene of Nebraska, ?Tapirus johnsoni and ?T. simpsoni. Tapirs are known in Nebraska from the Early Oligocene through the Middle Pleistocene. The northern limit of the distribution of the tapirs contracts gradually southward in North America throughout the Tertiary, and even during the Pleistocene interglacials Nebraska must have been near the northern limit of their range.

Stratigraphic Occurrences Of Teleoceras With A New Kimballianspecies From Nebraska, Lloyd G. Tanner

Bulletin of the University of Nebraska State Museum

Study of Teleoceras remains in the University of Nebraska State Museum indicates that this specialized, short-limbed rhinoceros inhabited the Central Great Plains from Early through Late Pliocene. Previously thought to have become extinct at the end of the middle Pliocene, this genus is now known from the very latest Pliocene. A new species, Teleoceras schultzi, is described from the Kimball Formation, Ogallala Group, Frontier County, Nebraska.

A New Species Of Menoceras From The Marsland Formation Of Nebraska, Lloyd G. Tanner

Bulletin of the University of Nebraska State Museum

Further study indicates that the Menoceras remains collected from the "Bridgeport Quarries" (lower port:on or the Marsland Formation, Middle Miocene, of the Hemingford Group) are of a new species.

In 1962, these dicerathere remains were considered to be a geologic variety of Diceratherium niobrarensis (Stecher, Schultz, and Tanner). However, later (Tanner, 1969) a generic distinction was revived, separating the Menoceras Troxell from Diceratherium Marsh. Diceretherium niobrarensis was then placed in synonymy with Menoceras arikarense (Barbour).

The new species is an intermediate between Menoceras arikarense (Barbour) from the Harrison Formation and Menoceras marslandensis Tanner, from the upper portion of the Marsland …

Two Lynx-Like Cats From The Pliocene And Pleistocene, C. Bertrand Schultz, Larry D. Martin

Bulletin of the University of Nebraska State Museum

A new species (Lynx stouti) of small felid is described from the lower Pliocene of Colorado. This form has several characters in common with the modern Lynx and may be ancestral to that genus. A new subspecies of Lynx issiodorensis Croizet and Jobert is described as L. i. kurteni from the Mullen Assemblage, Cherry County, Nebraska. The relationships of this form to other lynxes are discussed along with the paleo-distribution of the genus.

The classification of the felinae has always been somewhat controversial, especially at the generic level. One fairly homogenous group of cats which has been separated …

Phyletic Trends In Certain Lineages Of Quaternary Mammals, C. Bertrand Schultz, Lloyd G. Tanner, Lary D. Martin

Bulletin of the University of Nebraska State Museum

Evolutionary trends in certain carnivores, rodents, bovids, and proboscideans are examined. Chronoclines are demonstrated for muskrats, beavers, mammoths, and bison. The Pliocene-Pleistocene boundary and Quaternary climatic fluctuations are also discussed.

The University of Nebraska State Museum contains a large stratigraphically controlled collection of Pleistocene animals. This collection provides a unique opportunity for the study of phyletic trends in certain lineages of mammals. Although a few phylogenetic lineages have been proposed (Schultz and Frankforter, 1946; Hibbard, et al., 1965), this has not been done for the vast majority of Pleistocene mammals. However, there is much more data available than this would …

The Microtine Rodents Of The Mullen Assemblage From The Pleistocene Of North Central Nebraska, Larry D. Martin

Bulletin of the University of Nebraska State Museum

The University of Nebraska State Museum has had an active collecting program in Hooker and Cherry counties, Nebraska, since the 1930's. The following University of Nebraska State Museum collecting localities have been extensively excavated: Cr-10, Cr-102, Cr-11, Ho-101, Ho-102, and Ho-103. These localities have produced a large vertebrate fauna described by Jakway as the Mullen Local Fauna which he considered for the most part to be Early lllinoian. Further study of this local fauna now demonstrates that assemblage does include Early as well as Middle Pleistocene animals. The purpose of this paper is to report on the evidence regarding the …

A Revision Of The Feather Mite Genus Brephosceles (Proctophyllodidae: Alloptinae), Paul C. Peterson

Bulletin of the University of Nebraska State Museum

A systematic revision is presented for the genus Brephosceles. Data on external morphology, zoogeography, and host-parasite relationships are included. Thirteen named and thirty new species are recognized and described. The new species and the type hosts are: Brephosceles afribycis, from Afribyx lateralis (Charadriidae); B. anhimae, from Anhima cornuta (Anhimidae); B. belonopteri, from Belonopterus chilensis cayennensis (Charadriidae); B. bilobatus, from Puffinus puffinus (Procellariidae); B. chilensis, from Belonopterus chilensis lampronotus (Charadriidae); B. collaracus, from Charadrius collaris (Charadriidae); B. constrictus, from Charadrius bicinctus (Charadriidae); B. diomedei, from Diomedea chrysostoma (Diomedeidae); B. discidicus, from Cygnus bewickii (Anatidae); B. disjunctus, from Pterodroma leucoptera hypoleuca (Procellariidae); …

A Generic Revision Of The Pterodedinae, A New Subfamily Of Feather Mites (Sarcoptiformes: Analgoidea), Chong K. Park, Warren T. Atyeo

Bulletin of the University of Nebraska State Museum

The generic rev1s1on of the Pterodectinae, new subfamily of the Proctophyllodidae, is based on 88 named and over 160 new species. Included are morphology, host-parasite relationships, diagnoses of four named and eight new genera, and illustrations of each type species. The named genera are:

Anisodiscus Gaud and Mouchet, 1957; Montesauria Oudemans, 1905; Proterothrix Gaud, 1968; Pterodectes Robin, 1877. The new genera and their type species are: Dolichodectes, Proctophyllodes (Pterocolus) edwardsi Trouessart, 1885; Megalodectes, Proctophyllodes (Pterodectes) major Trouessart, 1885; Neodectes, Proctophyl/odes (Pterodectes) securiclatus Trouessart and Neumann, 1888; Pedanodectes, Pterodectes hologaster Gaud, 1953; Syntomodectes, Proctophyllodes (Pterodectes) selenurus Trouessart, 1885; Toxerodectes, Pterodectes gladiger …

Machairodont Cats From The Early Pleistocene Broadwater And Lisco Local Faunas, C. Bertrand Schultz, Larry D. Martin

Bulletin of the University of Nebraska State Museum

A new species of Ischyrosmilus (I. crusafonti) is reported from the Early Pleistocene Broadwater Local Fauna of Morrill County, Nebraska. Ischyrosmilus sp. is reported from the Early Pleistocene Lisco Local Fauna. Megantereon is reported from the Broadwater Local Fauna and this is the first record of the genus from North America.

A New Tribe Of Saber-Toothed Cats (Barbourofelini) From The Pliocene Of North America, C. Bertrand Schultz, Marian R. Schultz, Larry D. Martin

Bulletin of the University of Nebraska State Museum

A new genus of Pliocene Saber-toothed felid, Barbourofelis, is proposed and two new species B. fricki and B. morrisi are described. These two forms and other described material represent an unusual lineage of felids with long sabers, shortened crania, and massive postorbital bars. The tribal name Barbourofelini is proposed for this lineage which is presently known in North America from deposits ranging in age from Clarendonian through Kimballian. The Barbourofelini apparently migrated from Eurasia to North America in the Late Miocene or Early Pliocene. Sansanosmilus of the French Vindobonian appears to represent the ancestral stock of these cats. The …

A New Rhinoceros From The Nebraska Miocene, Lloyd G. Tanner

Bulletin of the University of Nebraska State Museum

A new species of rhinoceros, Menoceras marslandensis, is here proposed. It is based upon a nearly complete skull from the upper part of the Marsland Formation, Hemingford Group (Miocene), of Box Butte County, Nebraska. The new species was probably derived from Menoceras arikarense (Barbour), collected from the Agate Springs Quarries (Arikaree Group, Harrison Formation) of Sioux County, Nebraska, with which both Diceratherium niobrarensis Peterson and D. cooki Peterson are likely synonymous. From Menoceras arikarense, the presumed ancestral stock, Menoceras marslandensis differs in possessing the following characters: (1) very long, fused but slightly cleft, nasals, with flattened, rugose areas …

Revisions Of Two Subgenera Of Andrena: Micrandrena Ashmead And Derandrena, New Subgenus (Hymenoptera: Apoidea), David O. Ribble

Bulletin of the University of Nebraska State Museum

This paper is a monographic revision of the bee subgenera Micrandrena Ashmead and Derandrena, new subgenus, of the genus Andrena in North America. Both subgenera occur in the New World and Micrandrena also occurs in the Old World. Twenty-eight species are treated, of which twelve are new: Andrena (Micrandrena) annectens, ishii, labergei, lamelliterga, lepidii, micheneri, Andrena (Derandrena) arctostaphylae, californiensis, hermosa, murietae, penutiani, viridissima. Thirteen names are placed in synonymy: Andrena semotula Cockerell with candidiformis Viereck and Cockerell; subtilicornis Viereck, nitidicornis Cockerell, chlorogaster nesiotes Timberlake, chlorogaster gavilanica Timberlake, radialis Timberlake, catalinica Cockerell with chlorogaster Viereck; vegana Viereck and Cockerell with illinoiensis …

Two Feather Mite Genera (Analgoidea, Proctophyllodidae) From Birds Of The Families Oxyruncidae And Pipridae (Passeriformes, Tyranni), Warren T. Atyeo, J. Gaud

Bulletin of the University of Nebraska State Museum

The genus Hemipterodectes Berla is redefined and the type species, H. squalocauda is figured. A new genus, Diproctophyllodes (type species, Proctophyllodes (Alloptes) dielytra Trouessert, 1885) is described; D. exquisita (Berla), 1959 is synomized with D. dielytra; and D. oxyrunci, new species, is described.

Biometrical Study Of Morphology And Development Of The Pennsylvanian Trilobite Ameura Sangamonensis (Meek And Worthen), Roger K. Pabian, J. A. Fagerstrom

Bulletin of the University of Nebraska State Museum

Biometrical study of morphology and development in a sample of 100 cranidia, 94 pygidia, 26 free cheeks, and 9 complete specimens of the trilobite Ameura sangamonensis (Meek and Worthen) from the Bonner Springs Shale (Pennsylvanian; Missouri Series) in eastern Nebraska indicates that the dominant growth pattern was isometric. Evidence supporting this conclusion consists of rectilinear size relations among eight pairs of cranidial dimensions, three pairs of pygidial dimensions, and three pairs of dimensions of the free cheeks. Qualitative changes in pygidial morphology during development include a progressive decrease in prominence of the border, increasing width of the posterior border relative …

New Genera Related To The Genus Brephosceles Hull, 1934 (Acarina: Proctophyllodidae), Paul C. Peterson, Warren T. Atyeo

Bulletin of the University of Nebraska State Museum

Five new genera are described; these and the included species are (type species listed first): Dichobrephosceles with Dermaleichus actitidis Canestrini, 1878 (=Brephosceles dolichocaulus Gaud and Mouchet, 1957, new synonymy), D. eroliae, new species (ex: Erolia alpina, England); Homeobrephosceles with Proctophyllodes (A lloptes) discosurus Trouessart, 1886, Brephosceles orthothrix Gaud and Mouchet, 1957; Onychalloptes with Proctophyllodes (Alloptes) microphaeton Trouessart, 1885, Alloptes minutus Trouessart, 1899; Aramolichus with Aramolichus foliatus, new species (ex: Aramus guarauna, Bolivia); Psilobrephosceles with Dermalichus ortygometrae Canestrini, 1878. Species included in the genus Brephosceles (s.s.) are listed; species transferred to Brephosceles are: Proctophyllodes (Pterocolus) fiagellifer discursus Trouessart, 1885; Pterocolus lambda …

The Genera And Species Of The Feather Mite Subfamily Trouessartinae Except Trouessartia (Acarina: Proctophyllodidae), Kenneth R. Orwig

Bulletin of the University of Nebraska State Museum

A systematic study of the acarine subfamily Trouessartinae (except Trouessartia) of the family Proctophyllodidae is presented. Diagnoses are given for the subfamily, five named genera and four new genera. The named genera and their type species are: Allanalges Trouessart, 1886, A. podagricus (Trouessart), 1886; Calcealges Gaud, 1952, C. cyathoplax Gaud, 1952; Hemicalcealges Gaud and Mouchet, 1957, H. margaropygus (Gaud and Mouchet), 1957; Pseudalges Radford, 1950, Proctophyllodes (Pterocolus) analgoides Trouessart, 1885; and Trouessartia Canestrini, 1899, Dermaleichus corvinus Koch, 1840. New genera and their type species are: Arthrogynalges, A. biovoidatus, new species; Bicentralges, B. orientalis, new species; Neocalcealges, N. inattditus, new species; …

Citellus Kimballensis, A New Late Pliocene Ground Squirrel, Douglas C. Kent

Bulletin of the University of Nebraska State Museum

A new fossil sciurid, Citellus kimballensis, is described. This new species was found in the Kimball Formation, uppermost Ogallala (very late Pliocene) at the University of Nebraska State Museum Collecting Locality Cn-1O1, northeast of Sidney, Cheyenne County, Nebraska. Characters of the dentition and skull of C. kimballensis are compared with those of other species of the genus, and with those of Cynomys.

The Monotypic North American Subgenus Larandrena Of Andrena (Hymenoptera: Apoidea), D. W. Ribble

Bulletin of the University of Nebraska State Museum

Larandrena contains a single species, A. miserabilis Cresson. This small, common Andrena is found throughout most of the United States and southern Canada in the spring. The subgeneric position, nomenclature, redescription. variation, notes on the biology, parasites, range, seasonal activity and plant host records are included. A. rniserabilis is thought to be an important pollinator of fruit trees.

A New Species Of Rhinoceros, Aphelops Kimballensis, From The Latest Pliocene Of Nebraska, Lloyd G. Tanner

Bulletin of the University of Nebraska State Museum

An uncrushed skull and associated skeletal elements of a very large rhinoceros are the basis for the description of a new species of Aphelops from Frontier County, Nebraska. The major differences are: the skull is much larger in most dimensions than other Aphelops; it has an extremely elevated occipital region in comparison to Aphelops mutilus; the narial notch is retracted to a point perpendicular to the center of molar one; and the teeth are more hypsodont than any other species of this genus. The sediments which yielded the skull and skeletal parts are considered to be Kimballian in …

A Revision Of The Bees Of The Genus Andrena Of The Western Hemisphere. Part I. Callandrena. (Hymenoptera: Andrenidae), Wallace E. Laberge

Bulletin of the University of Nebraska State Museum

This paper is the first part of a monograph of the bee genus Andrena in the western hemisphere and treats the subgenus Callandrena. Available data regarding phylogeny, distribution, biology, and flower preferences are presented together with keys to separate the species, diagnoses and descriptions of the species and discussions of geographic variation when applicable. Seventy-nine species and one subspecies are recognized. Sixteen names are relegated to synonymy, one to homonymy and eight are removed from the subgenus Callandrena. The thirty-nine species new to science are: aerifera, aeripes, afimbriata, ardis, auripes, balsamorhizae, beameri, bilimeki, bullata, calvata, dreisbachorum, fulminea, fulminoides, …

The Brule-Gering (Oligocene-Miocene) Contact In The Wildcat Ridge Area Of Western Nebraska, C. Bertrand Schultz, Charles H. Falkenbach, Carl F. Vondra

Bulletin of the University of Nebraska State Museum

The contact between the Brule Formation (Oligocene) and the Gering Formation (Miocene) can be readily distinguished in the Wildcat Ridge area, as elsewhere in western Nebraska. At the critical fossiliferous exposures at Castle Rock in Scotts Bluff County, the contact on the south face between the two formations is defined as 129 feet above the base of the "Upper Ash" bed, which corresponds to the upper portion of Darton's (1899, PI. C, Fig. D, following p. 754) "sandy phase" in the upper part of the Brule. Certain key beds in the Gering Formation can be traced laterally from a channel …

The Feather Mite Genus Proctophyllodes (Sarcoptiformes: Proctopbyllodidae), Warren T. Atyeo, Norman L. Braasch

Bulletin of the University of Nebraska State Museum

A systematic revision is presented for the genus Proctophyllodes. Data on the bionomics, morphology, and host-parasite relationships are included.

Prodromus Of American Bees Of The Genus Andrena (Hymenoptera, Apoidea), Wallace E. Laberge

Bulletin of the University of Nebraska State Museum

Keys to the females and males and diagnoses of the American subgenera of the genus Andrena are presented in this paper. Previously described subgenera recognized in this work are as follows: Callandrena Cockerell, Scaphandrena Lanham, Plastandrena Hedicke, Parandrena Robertson, Dactylandrena Viereck, Andrena Fabricius, Iomelissa Robertson, Oligandrena Lanham, Hesperandrena Timberlake, Micrandrena Ashmead, Gonandrena Viereck, Opandrena Robertson, Cnemidandrena Hedicke, Simandrena Perez, Thysandrena Lanham, Taeniandrena Hedicke, Ptilandrena Robertson, Trachandrena Robertson, Xanthandrena Lanham, Diandrena Cockerell, Leucandrena Hedicke, Melandrena Perez, Gymnandrena Hedicke, Scoliandrena Lanham, Onagrandrena Linsley and MacSwain. The following five new subgenera are described: Eremandrena, Larandrena, Tylandrena, Geandrena, Chaulandrena.

New Species Of Proctophyllodes From Bulgaria (Sarcoptiformes, Analgoidea), Warren T. Atyeo, I. D. Vassilev

Bulletin of the University of Nebraska State Museum

Two new species of feather mites are described: Proctophyllodes tenericaulus from Turdus viscivorus L. (Turdidae) and Galerida cristata (L.) (Alaudidae) and Proctophyllodes emberizae from Emberiza melanocephala Scopoli and E. hortulana (L.) (Fringillidae).

A Review Of The Mites Of The Family Pseudocheylidae Oudemans, 1909 (Acarina, Prostigmata), Edward W. Baker, Warren T. Atyeo

Bulletin of the University of Nebraska State Museum

Four genera and ten species are included; these are: Pseudocheylus biscalattts Ber1ese, 1888, P. americanus (Ewing), 1909, Stigmocheylus brevisetus Berlese, 1910, Anoplocheylus europaeus (Berlese), 1910, A. clavatus, n. sp., A. aegypticus, n. sp., A. protea (Womersley), 1935, Neocheylus natalensis Tragardh, 1906, N. nidcolus Lawrence, 1954, and N. collis, n. sp.

Tarsocheylidae, A New Family Of Prostigmatic Mites (Acarina), Warren T. Atyeo, Edward W. Baker

Bulletin of the University of Nebraska State Museum

The new family is based on two genera, Tarsocheylus Berlese, 1904 and Hoplocheylus, new genus. The species included are: Tarsocheylus paradoxus Berlese, 1904, Hoplocheylus atomarius (Berlese), 1913, H. johnstoni, n. sp., H. aethiopicus (Cooreman), 1951, H. longispinus, n. sp., and H. cliscalis, n. sp.

The Bdellidae (Acarina) Of The Australian Realm, Warren T. Atyeo

Bulletin of the University of Nebraska State Museum

Although primarily a systematic study, variations in the integumental striae and chaetotaxy of the legs are discussed. The genus Octobdellodes Atyeo, 1960 is synonymized with Bdellodes Oudemans, 1937 and the sugenus Cytobdella Mihelcic, 1958 is synonymized with Cyta von Heyden, 1826. The genus Bdellodes is divided into the subgenera Bdellodes and Hoploscirus Thor, 1937 (=Thoribdella Grandjean, 1938, new synonymy). Twenty-two species are now known to occur in New Zealand and the outlying islands. These include four known species: Bdella iconica, Bdellodes (Roploscirus) lapidaria, B. (R.) reticulata, and Cyta latirostris, and eighteen new species: Bdellodes (Bdellodes) harpax, B. (B.) oraria, …