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Biological and Physical Anthropology Commons™

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2017

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Articles 31 - 58 of 58

Full-Text Articles in Biological and Physical Anthropology

Claiming The Indomitable Wave: Masculinities, Sexualities, And The Realm Of Surfing In Costa Rica, Joseph C. Recupero Apr 2017

Claiming The Indomitable Wave: Masculinities, Sexualities, And The Realm Of Surfing In Costa Rica, Joseph C. Recupero

Student Publications

Examining the relationship between masculinity, sexuality, and the sport of surfing in the context of Costa Rica. Questions the nature of emergent counter identities in the hyper-masculine realm of the surfing subculture and the ways in which the emergence of counter identities changes the nature of the subculture. Focuses on the anthropology of sport, the anthropology of sexuality, and theories of territoriality.


Horticultural Activity Predicts Later Localized Limb Status In A Contemporary Pre-Industrial Population, Jonathan Stieglitz, Benjamin C. Trumble, Hillard Kaplan, Michael Gurven Mar 2017

Horticultural Activity Predicts Later Localized Limb Status In A Contemporary Pre-Industrial Population, Jonathan Stieglitz, Benjamin C. Trumble, Hillard Kaplan, Michael Gurven

ESI Publications

Objectives—Modern humans may have gracile skeletons due to low physical activity levels and mechanical loading. Tests using prehistoric skeletons are limited by the inability to assess behaviour directly, while modern industrialized societies possess few socio-ecological features typical of human evolutionary history. Among Tsimane forager-horticulturalists, we test whether greater activity levels and, thus, increased loading earlier in life are associated with greater later-life bone status and diminished age-related bone loss.

Materials and Methods—We used quantitative ultrasonography to assess radial and tibial status among adults aged 20+ years (mean±SD age=49±15; 52% female). We conducted systematic behavioural observations to assess earlier-life …


Human Grooming In Comparative Perspective: People In Six Small-Scale Societies Groom Less But Socialize Just As Much As Expected For A Typical Primate, Adrian V. Jaeggi, Karen Kramer, Raymond Hames, Evan J. Kiely, Cristina Gomes, Hillard Kaplan, Michael Gurven Feb 2017

Human Grooming In Comparative Perspective: People In Six Small-Scale Societies Groom Less But Socialize Just As Much As Expected For A Typical Primate, Adrian V. Jaeggi, Karen Kramer, Raymond Hames, Evan J. Kiely, Cristina Gomes, Hillard Kaplan, Michael Gurven

ESI Publications

Objectives—Grooming has important utilitarian and social functions in primates but little is known about grooming and its functional analogues in traditional human societies. We compare human grooming to typical primate patterns to test its hygienic and social functions.

Materials and Methods—Bayesian phylogenetic analyses were used to derive expected human grooming time given the potential associations between grooming, group size, body size, terrestriality, and several climatic variables across 69 primate species. This was compared against observed times dedicated to grooming, other hygienic behavior and conversation among the Maya, Pumé, Sanöma, Tsimane’, Yanomamö, and Ye’kwana (mean number of behavioral scans …


Zooarchaeological And Palaeoenvironmental Reconstruction Of Newly Excavated Middle Pleistocene Deposits From Elandsfontein, South Africa, Frances L. Forrest Feb 2017

Zooarchaeological And Palaeoenvironmental Reconstruction Of Newly Excavated Middle Pleistocene Deposits From Elandsfontein, South Africa, Frances L. Forrest

Dissertations, Theses, and Capstone Projects

Increased consumption of animal tissue is arguably one of the most important adaptive transitions in early hominin behavior. A dietary shift toward regular tool-assisted meat consumption and increased competition with the carnivore paleoguild likely helped shape many important hominin adaptations such as foraging patterns, habitat preferences, and social behaviors. Yet, the ecological and behavioral implications for increased hominin carnivory remain poorly understood. This dissertation examines the zooarchaeological and paleoenvironmental history of an important Acheulean hominin locality, Elandsfontein, South Africa (ca. 1.0 – 0.6 Ma). The goal is to begin addressing under-investigated aspects of Acheulean hominin behavioral ecology and place Acheulean …


The Index Of Texas Archaeology: Open Access Gray Literature From The Lone Star State, Robert Z. Selden Jr., C. Britt Bousman Jan 2017

The Index Of Texas Archaeology: Open Access Gray Literature From The Lone Star State, Robert Z. Selden Jr., C. Britt Bousman

CRHR: Archaeology

Cultural resources management (CRM) reports represent a rapidly growing proportion of our knowledge associated with archaeological undertakings in the United States. Historically, these reports were printed in limited numbers and distributed to a few libraries and individuals, and few were distributed beyond the political boundaries of any given state. Libraries on the distribution list are reticent to allow patrons to check out these reports due to the fact that they have—and will only ever have—a single copy. Late in 2009, the Texas Historical Commission (THC) permitting guidelines for CRM reports were updated, requiring CRM contractors to submit a digital copy …


Mitochondrial-Dna Phylogenetic Information And The Reconstruction Of Human Population History: The South American Case, María Bárbara Postillone, S. Ivan Perez Jan 2017

Mitochondrial-Dna Phylogenetic Information And The Reconstruction Of Human Population History: The South American Case, María Bárbara Postillone, S. Ivan Perez

Human Biology Open Access Pre-Prints

Objectives: Mitochondrial DNA (mtDNA) sequences are becoming increasingly important in the study of human population history. Here, we explore the differences in the amount of information of different mtDNA regions and their utility for the reconstruction of South American population history.

Material and methods: We analyzed six datasets comprising 259 mtDNA sequences from South America: Complete mtDNA, Coding, Control, hypervariable region I (HVRI), cytochrome b (cytb) plus Control, and cytb plus 12S plus 16S. The amount of information in each dataset was estimated employing several site-by-site and haplotype based statistics, distances among sequences, Neighbor-joining trees, distances among the estimated trees, …


Use Of Vertical Enclosure Space And Species-Typical Locomotion By A Rehabilitating Spider Monkey (Ateles Fusciceps), Jake Funkhouser Jan 2017

Use Of Vertical Enclosure Space And Species-Typical Locomotion By A Rehabilitating Spider Monkey (Ateles Fusciceps), Jake Funkhouser

Undergraduate Honors Theses

With wild spider monkey populations in decline, investigations contributing to captive welfare, and successful rehabilitation and reintroduction knowledge is increasingly pressing. Quantifying and analyzing the appropriateness of naturalistic enclosure designs to foster species-typical behaviors is an effective way to address both of these needs. This study investigates enclosure space use, vertical space preference, substrate use, positional/postural modes, and interactions with human caregivers of a wild-caught, pet-trade rehabilitant Columbian black spider monkey (Ateles fusciceps rufiventris, N = 1). Video data collected daily from August to October 2015 via focal animal sampling (from 08:00 to 10:00) at Alouatta Sanctuary, Panama …


Talking With The Dead From Southern Mexico: Tracing Bioarchaeological Foundations And New Perspectives In Oaxaca, Ricardo Higelin Ponce De León, Guy David Hepp Jan 2017

Talking With The Dead From Southern Mexico: Tracing Bioarchaeological Foundations And New Perspectives In Oaxaca, Ricardo Higelin Ponce De León, Guy David Hepp

Anthropology Faculty Publications

In recent years, bioarchaeological research has emerged as one of the principal tools for studying diverse aspects of ancient society through the analysis of human remains. Oaxaca, Mexico is a culturally diverse region of Mesoamerica that today boasts at least sixteen discrete ethnolinguistic groups. Bioarchaeology in Oaxaca has traditionally focused on the Late Formative and Classic period (500 BCE–800 CE) Zapotec city of Monte Albán, and on its hinterland. Largely descriptive, these studies provided basic discussions of health, nutrition, and funerary offerings. Beyond the Valley of Oaxaca, much less comprehensive bioarchaeological research has been undertaken to date. Never before have …


The Effect Of Polycystic Ovarian Syndrome (Pcos) On The Human Skeleton, Sarah Bruckler Jan 2017

The Effect Of Polycystic Ovarian Syndrome (Pcos) On The Human Skeleton, Sarah Bruckler

Senior Honors Theses and Projects

The purpose of this pilot study is to determine the systemic effects of Polycystic Ovary Syndrome (PCOS) on bone mineral density (BMD). Excessive male sex honnones, excessive insulin levels, and weight gain characterize PCOS, and are correlated to increased BMD. PCOS is also associated with symptoms and comorbid conditions, like chronic vitamin D deficiency, menstrual dysfunction, and hypothyroidism, which are all correlated with decreased BMD. Existing research on this topic reports conflicting results; some studies show a significant correlation between PCOS and increased BMD, while others suggest that no significant correlation exists. These previous studies focused on load-bearing areas of …


Healthy Aging In The North: Sociocultural Influences On Diet And Physical Activity Among Older Adults In Anchorage, Alaska, Britteny M. Howell Jan 2017

Healthy Aging In The North: Sociocultural Influences On Diet And Physical Activity Among Older Adults In Anchorage, Alaska, Britteny M. Howell

Theses and Dissertations--Anthropology

Increasing rates of overweight, obesity, and related cardiovascular diseases among older adults in the United States present unique public health challenges. Cross-cultural research has shown marked variation in health across the world’s elder populations because aging is a biological process rooted in sociocultural context. The sociocultural environment contributes to complex negotiations of food and physical activity patterns for older adults. It is well established in the literature that urban residents report low levels of physical activity and have easy access to fast food outlets, which tend to be concentrated in lower-income neighborhoods. I utilize a biocultural framework, integrating nutritional anthropology …


Prevalence Of Dental Pathology In A Juvenile Population From The Ancient Maya Site Of Altun Ha, Lindsey D. Lefebvre Jan 2017

Prevalence Of Dental Pathology In A Juvenile Population From The Ancient Maya Site Of Altun Ha, Lindsey D. Lefebvre

Honors Undergraduate Theses

The present research seeks to assesses the presence and prevalence of two distinct dental pathologies: linear enamel hypoplasia and caries in an ancient Maya juvenile subsample from Altun Ha, Belize spanning the Preclassic (ca. 600 B.C.) through the Terminal Classic (ca. 900 A.D.) periods. Teeth offer a remarkable wealth of information about the human experience in the past. Developmental and post-eruption pathology can provide insight into cultural and evolutionary processes by illuminating social and biological factors such as diet, weaning, illness, and overall health that manifest in observable changes to the composition of teeth. In addition, growth and developmental stages …


Juvenile Remains: Predicting Body Mass And Stature In Modern American Populations, Erin F E Pinkston Jan 2017

Juvenile Remains: Predicting Body Mass And Stature In Modern American Populations, Erin F E Pinkston

Cal Poly Humboldt theses and projects

There are increasing numbers of unidentified persons in the U.S. and abroad. To generate positive identifications, forensic anthropologists and others working in the medicolegal field employ a variety of methods to produce biological profiles to match to case files and missing persons databases. Body mass, and stature are two important components of a biological profile, and both can be estimated using regression formulae derived from skeletal metrics. In cases of unidentified juvenile remains, these are particularly important metrics, as it is difficult or impossible to determine sex in prepubescent remains, and the quality of ancestry estimation is currently under debate …


Fertility And Reproduction's Niche: Human Sexual Diversity, Samuel W. Austin Jan 2017

Fertility And Reproduction's Niche: Human Sexual Diversity, Samuel W. Austin

Graduate Student Theses, Dissertations, & Professional Papers

Abstract: Biologically exploring the origins and forms of human sexuality is of paramount importance. Scientific research has indicated that homosexuality was linked to reproduction, fertility, and adaptive child caring strategies, traits that seem to display cross-cultural similarities. This suggests that sexual diversity may be one of human’s earliest adaptations. While most of the previous research has been on individuals of European descent, little research on Native American populations has been completed to test whether these patterns continue in their population.

The research presented here tests the Sexually Antagonistic Hypothesis for Male Homosexuality, Fraternal Birth Order Effect, and childhood atypical gender …


Evolution Of Lemur Pelage Color Variation, Amanda Nicole Spriggs Jan 2017

Evolution Of Lemur Pelage Color Variation, Amanda Nicole Spriggs

Legacy Theses & Dissertations (2009 - 2024)

This dissertation investigates the selective pressures that may be driving variation in pelage color patterns in Eulemur, Varecia, and Propithecus. This is addressed by exploring the relationship between the lemur pelage colors and ecological variables, such as habitat type, predation pressures, visibility variables, and level of sympatry. The following model is presented here to describe the evolutionary mechanisms that are thought to be maintaining pelage coloration in lemurs: Pelage color and pattern are potentially used to (1) signal to conspecifics information such as mate quality, (2) signal to congenerics species identity, and (3) avoid signaling the individual’s presence to predators. …


A Stylistic Analysis Using Multivariate Statistics Of Oneota Pottery From The Upper Mississippi, Blue Earth, And St. Croix River Valleys, Michelle Neumann Jan 2017

A Stylistic Analysis Using Multivariate Statistics Of Oneota Pottery From The Upper Mississippi, Blue Earth, And St. Croix River Valleys, Michelle Neumann

All Graduate Theses, Dissertations, and Other Capstone Projects

Pottery is one of the most abundant artifact types recovered from late pre-contact habitations sites in the upper Midwest. As a material with inherent plasticity, pottery reflects changes in people's preferences and traditions in aspects of its form and design quickly through time and space. Analyzing different facets of pottery has the ability to provide extensive information about people in the past: their resource utilization, technology, traditions, economic exchange, regional interaction, ideology, and or group identity. Yet, a significant challenge in pottery analysis is deriving comprehensive and testable conclusions in terms of types and styles that reflect patterned cultural behavior …


A Comprehensive Case Report Of The University Of Montana Case 37, Cody M. Lawson Jan 2017

A Comprehensive Case Report Of The University Of Montana Case 37, Cody M. Lawson

Graduate Student Theses, Dissertations, & Professional Papers

In this professional paper I examine the human skeletal remains of one individual. The remains were analyzed to gain insight into the age, sex, ancestry, stature, weight, pathology, and trauma of the individual. Forensic anthropological methods were applied to UMFC 37. The remains of UMFC 37 represent a male, between the age of 40 and 60. He is likely a Caucasian. UMFC 37 is between 5 feet 6 inches and 5 feet 10 inches tall and weighs between 148 and 167 pounds.


An Assessment Of Trace Elements Distribution In Teeth Utilizing A Sample Group From Postclassic Lamanai: The Application Of La-Icp-Ms In Bioarchaeology And Forensics, Michelle Hawkins Jan 2017

An Assessment Of Trace Elements Distribution In Teeth Utilizing A Sample Group From Postclassic Lamanai: The Application Of La-Icp-Ms In Bioarchaeology And Forensics, Michelle Hawkins

Electronic Theses and Dissertations

Trace element analysis of skeletal remains and teeth is a common research technique in biological and forensic anthropology. In particular, LA-ICP-MS has become a widely-accepted tool for analyzing and mapping the distribution of trace elements in teeth. Investigation into the relative spectral intensities and spatial distribution of thirteen trace isotopes (13C, 24Mg, 27Al, 31P, 44Ca, 47Ti, 52Cr, 55Mn, 56Fe, 66Zn, 88Sr, 138Ba, 208Pb) within teeth was undertaken using LA-ICP-MS. The total archaeological sample of teeth (N=26) was comprised of four tooth types (UCI, ULI, UPM1, and UPM2) and 18 individuals from a Postclassic Lamanai site. In preparation for analysis, teeth …


Ancient Dna Extraction From Stone Tools, Clare Super Jan 2017

Ancient Dna Extraction From Stone Tools, Clare Super

Undergraduate Theses, Professional Papers, and Capstone Artifacts

Proteins and DNA can be trapped in the microcracks on the surface of stone tools, which can then be extracted and analyzed to aid in inferring the use of the tool (Shanks et al. 2001; 2005). This nondestructive method involves the use of sonication to release DNA from the microcracks, then amplification of regions of mitochondrial DNA that are species specific. This technique was applied to stone tools from the Bridge River site in British Columbia by researchers at UMT’s Ancient and Modern Molecular Anthropology labs. Bridge River archaeologists have designated the tools as used in “food processing or tool …


Cognitive Performance Across The Life Course Of Bolivian Forager-Farmers With Limited Schooling, Michael Gurven, Eric Fuerstenberg, Benjamin C. Trumble, Jonathan Stieglitz, Bret Beheim, Helen Davis, Hillard Kaplan Jan 2017

Cognitive Performance Across The Life Course Of Bolivian Forager-Farmers With Limited Schooling, Michael Gurven, Eric Fuerstenberg, Benjamin C. Trumble, Jonathan Stieglitz, Bret Beheim, Helen Davis, Hillard Kaplan

ESI Publications

Cognitive performance is characterized by at least two distinct life course trajectories. Many cognitive abilities (e.g. “effortful processing” abilities including fluid reasoning, and processing speed) improve throughout early adolescence and start declining in early adulthood, while other abilities (e.g. “crystallized” abilities like vocabulary breadth) improve throughout adult life, remaining robust even at late ages. Although schooling may impact performance and cognitive “reserve”, it has been argued that these age patterns of cognitive performance are human universals. Here we examine age patterns of cognitive performance among Tsimane forager-horticulturalists of Bolivia, and test whether schooling is related to differences in cognitive performance …


Nine Gal Tavern Faunal Analysis, Zachary T. Boyer Jan 2017

Nine Gal Tavern Faunal Analysis, Zachary T. Boyer

A with Honors Projects

Over 400 pieces of bone and eggshell were collected during excavation at the Nine Gal Tavern site (11CH541) located in western Champaign County, Illinois in 1987 and 1991 by a team led by archaeologist Lenville Stelle (Stelle 2006). The majority of the remains analyzed were recovered within feature context in the immediate vicinity of the established Nine Gal Tavern structure. The purpose of this paper is to describe the identification of these faunal remains which are housed at the Anthropology Program at Parkland College. The identification of these remains was carried out as part of an Honors Project under the …


Anthropometric Measures Of Hadza Forager Children And Juveniles: Implications For Anthropology And Human Biology, Elle Ford, Alyssa N. Crittenden Jan 2017

Anthropometric Measures Of Hadza Forager Children And Juveniles: Implications For Anthropology And Human Biology, Elle Ford, Alyssa N. Crittenden

McNair Poster Presentations

Understanding growth trajectories during child development is important for the fields of human biology, public health, medicine, and evolutionary medicine. To date, the majority of current references for childhood development and health are primarily derived from the World Health Organization (WHO) and the Centers for Disease Control and Prevention (CDC). To ensure a child is developing properly, is equipped with proper nutrition, and will sustain a healthy life through adolescence and adulthood, these measurements and standards are necessary. The current study aims to test whether or not these standards are applicable in non-industrial small-scale societies where differences from western populations …


Managing Sociality Of A Captive Female Bornean Orangutan From Breeding To Post-Partum At The Smithsonian's National Zoo, Marie Vergamini Jan 2017

Managing Sociality Of A Captive Female Bornean Orangutan From Breeding To Post-Partum At The Smithsonian's National Zoo, Marie Vergamini

Theses and Dissertations

The Association of Zoos and Aquariums’ Orangutan Species Survival Plan® aims to maintain 100 Bornean orangutans (Pongo pygmaeus) in captivity. Because investment in breeding these lineages is high, properly managing sociality of potential mothers is essential. This study assessed how behaviors of a captive breeding female at the Smithsonian’s National Zoo changed from pre-gestation through the offspring’s sixth month of age to improve breeding recommendations.

The infant Bornean orangutan was born September 2016. Results indicate that during breeding, the mother socialized most with two adult females. During pregnancy, the pregnant female socialized in less energy-consuming ways, i.e. grooming. …


Traditional Medicine Use In Chamorro Cué, Gral. E. Aquino, San Pedro, Paraguay, Noah Goyke Jan 2017

Traditional Medicine Use In Chamorro Cué, Gral. E. Aquino, San Pedro, Paraguay, Noah Goyke

Dissertations, Master's Theses and Master's Reports

Rural Paraguay presents great opportunities for investigation into the subtle differences in the use of medicinal plants across seasons and the urban versus rural dichotomy in a rapidly globalizing society. One of the most interesting opportunities is comparing a rural population to the population of an urban center that has be the destination of local migration from the rural communities surveyed. This study investigates the seasonal differences and difference between urban and rural residents in the type of medication used to treat thirteen different common ailments and the source of medicinal plants used to treat those ailments. Interviews performed in …


Chimpanzee (Pan Troglodytes) Space Use In A Sanctuary Setting, Holly Soubiea Jan 2017

Chimpanzee (Pan Troglodytes) Space Use In A Sanctuary Setting, Holly Soubiea

All Master's Theses

Studies of chimpanzee space use can provide information about chimpanzees’ preferences and welfare. This study examined the space use of 12 chimpanzees at Fauna Foundation, a chimpanzee sanctuary located in Carignan, Québec, Canada. Observers recorded the chimpanzees’ location, elevation, perimeter, and substrate using a scan sampling procedure. In total, there were 93 hours of data and 3,190 scans. The author used Geographical Information System (GIS) to plot the chimpanzees’ location for each scan. The chimpanzees used the largest rooms most often, were inside 61% (n = 1,942) of the time, an arm’s reach away from the perimeter 90% (n = …


The Social Behavior Of Rehabilitated Spider Monkeys (Ateles Geoffroyi), Anthony Denice Jan 2017

The Social Behavior Of Rehabilitated Spider Monkeys (Ateles Geoffroyi), Anthony Denice

All Master's Theses

The welfare of captive primates in laboratories, sanctuaries, and zoos is affected by various aspects of their environments. Although space restrictions increase aggression and stress-related behaviors in most captive animals, primates show diverse mechanisms for displacing stress and mitigating conflict. Many primates, including wild spider monkeys (genus Ateles), use these mechanisms flexibly to cope with social and environmental stressors. I investigated whether or not captive black-handed spider monkeys (Ateles geoffroyi) use behavioral strategies to cope with potential stressors in captivity. In particular, I tested whether an affiliative or avoidant strategy was used in response to changes in …


Collective Decision Making In Tibetan Macaques: How Followers Affect The Rules And Efficiency Of Group Movement, Amanda Rowe Jan 2017

Collective Decision Making In Tibetan Macaques: How Followers Affect The Rules And Efficiency Of Group Movement, Amanda Rowe

All Master's Theses

Primate societies must undergo successful collective decision making during group movement to stay cohesive and provide the ecological and evolutionary benefits of sociality. This study investigates how a fan structure facilitates successful group movement in the YA1 group of Tibetan macaques (Macaca thibetana) at Mt. Huangshan National Reserve in Anhui, China. We used structural equation modeling (SEM) to determine the relative influences of sex, age, maternal kinship, dominance, and social network centrality on the number of fans (consistent followers) an individual had, and the number of group members an individual was a fan of (fandom). SEM revealed that …


Family Dynamics In Eastern Hoolock (Hoolock Leuconedys), Javan (Hylobates Moloch), Pileated (H. Pileatus) And Northern White-Cheeked Gibbons (Nomascus Leucogenys) At The Gibbon Conservation Center, Alexandra Sheldon Jan 2017

Family Dynamics In Eastern Hoolock (Hoolock Leuconedys), Javan (Hylobates Moloch), Pileated (H. Pileatus) And Northern White-Cheeked Gibbons (Nomascus Leucogenys) At The Gibbon Conservation Center, Alexandra Sheldon

All Master's Theses

Few studies have explored the continuing relationship and behaviors within primate groups as the offspring mature and continue to depend on parental care. Between June 7 and July 7, 2016, I studied the family dynamics of 5 gibbon groups from 4 species: hoolock (Hoolock leuconedys), Javan (Hylobates moloch), pileated (H. pileatus) and northern white-cheeked (Nomascus leucogenys) gibbons housed at the Gibbon Conservation Center (GCC) in Santa Clarita, CA. I recorded all occurrences of approach, retreat, groom and play behaviors as well as the proximity individuals maintain between each other. I predicted: 1. …


Enrichment Use & Social Interactions In A Mixed-Species Enclosure Of Sumatran (Pongo Abelij) & Bornean Orangutans (P. Pygmaeus) & Northern White-Cheeked Gibbons (Nomascus Leucogenys), Emily Veitia Jan 2017

Enrichment Use & Social Interactions In A Mixed-Species Enclosure Of Sumatran (Pongo Abelij) & Bornean Orangutans (P. Pygmaeus) & Northern White-Cheeked Gibbons (Nomascus Leucogenys), Emily Veitia

All Master's Theses

Enrichment is an aspect of captive husbandry that has been shown to positively impact animals’ well-being and can be designed to encourage species-typical behaviors, such as foraging and arboreality. Enrichment can include housing together multiple compatible species. Orangutans are sympatric with siamangs and agile and Bornean gibbons, so several zoos house these four Asian ape species in the same enclosure, in part, as social enrichment. In my study, I observed enrichment use and social interactions in a mixed-species enclosure at the Oregon Zoo, which housed two Sumatran (Pongo abelii) and two Bornean (P. pygmaeus) orangutans with …