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Articles 31 - 60 of 1624

Full-Text Articles in Paleontology

Latah Formation: History And Case Studies, Kylee M. Woodworth May 2025

Latah Formation: History And Case Studies, Kylee M. Woodworth

2025 Symposium

The Latah Formation is a series of discontinuous sedimentary interbeds between the flows of the Columbia River Flood Basalts outcropping across northern Idaho and eastern Washington. Ages ranging between the Middle to Upper Miocene, the sediments represent varied fluvio-lacustrine environments. The deposits are predominantly composed of claystones and shales as alternating, laminated couplets, many containing a variety of fossils. Though the sediments may appear similar across localities, they come from different source rocks, ranging from ashes from Yellowstone hotspot to metamorphic rock from nearby hills. These ancient lacustrine and fluvial environments allowed for great preservation of fossils, often in great …


The Demise Of Fossil Cycad National Monument As A Consequence Of Paleontological Resource Mismanagement, Nathaniel Valentine, Vincent Santucci, Justin Tweet, Aubrey Bond May 2025

The Demise Of Fossil Cycad National Monument As A Consequence Of Paleontological Resource Mismanagement, Nathaniel Valentine, Vincent Santucci, Justin Tweet, Aubrey Bond

Undergraduate Research Symposium Posters

Located in the Black Hills of South Dakota (Figure 1), Fossil Cycad National Monument boasted well-preserved specimens of cycadeoid fossils from the Cretaceous and concentrations of these fossil plants were greatest at the site’s location (Santucci and Hughes, 1998). I studied historical records to estimate the number of fossils removed by paleontologists at the site between 1890-1957.


Inferring Feeding Biology In Extinct Shrews (Soricidae: Mammalia): Application Of A Linear Morphometric Approach, Derek J. Den Ouden May 2025

Inferring Feeding Biology In Extinct Shrews (Soricidae: Mammalia): Application Of A Linear Morphometric Approach, Derek J. Den Ouden

Electronic Theses and Dissertations

Linear morphometrics have been used extensively to infer the ecology of fossil organisms, but shrews have not yet been thoroughly explored using this technique. Often considered to have a homogenous insectivorous diet, direct dietary observation suggests shrew dietary diversity is substantial. Diversity of this nature is well established to have influenced skull morphology in other mammal clades, so similar patterns are hypothesized to be detectable in shrews. To test this, I examined the link between known diet and morphological disparity in extant shrews using linear morphometrics and then utilized that framework to infer the dietary ecology of several fossil taxa. …


The Petm: A Window Into Earth's Future Climate, Jacalyn M. Wittmer Malinowski May 2025

The Petm: A Window Into Earth's Future Climate, Jacalyn M. Wittmer Malinowski

Climate Change and the Individual, 2025-26

The Paleocene-Eocene Thermal Maximum (PETM) is not just an event that occurred in Earth’s distant past (~55 million years ago) but is a crucial lesson for today’s meteoric climate change. The PETM offers a historical parallel of rapid flux of greenhouse gases into the atmosphere that caused global warming, ocean acidification, dramatic changes in weather, and extinctions. This global warming event was the result of 20,000 years of “rapid” warming (in geological terms) however, we are experiencing rates of warming that are 5-10x higher than PETM levels. We are not experiencing rapid warming today, in comparison to the past, we …


The Human Legacy: Agriculture, Civilizations, Strife, And Pandemics Changing Our Climate, Jacalyn M. Wittmer Malinowski May 2025

The Human Legacy: Agriculture, Civilizations, Strife, And Pandemics Changing Our Climate, Jacalyn M. Wittmer Malinowski

Climate Change and the Individual, 2025-26

For most of human history, climate has shaped our lives, while our influence on it remained minimal. However, the notion that early humans had no measurable impact on the climate before industrialization is inaccurate. Early in human history, activities – such as the rise of agriculture, the growth of civilizations, periods of conflict, and pandemics – altered greenhouse gas levels in the atmosphere. These changes contributed to significant periods of climate change long before the burning of fossil fuels set us on the path to today’s global climate crisis.


Cranial Ontogeny Of The Extinct Dwarf Tapir Tapirus Polkensis; With Comparison To That Of The Extant Tapirus Bairdii (Tapiridae, Perissodactyla), Johannah Orendorff May 2025

Cranial Ontogeny Of The Extinct Dwarf Tapir Tapirus Polkensis; With Comparison To That Of The Extant Tapirus Bairdii (Tapiridae, Perissodactyla), Johannah Orendorff

Electronic Theses and Dissertations

Tapirus polkensis is the smallest known species of tapir to ever live, and the largest deposit of their fossils is currently located at the GFS in Gray, TN. Phylogenetics suggest that the extant T. bairdii may be the sister taxon to T. polkensis, although they differ greatly in their morphology. Variation exists among adult T. polkensis specimens, with patterns not present in T. bairdii. For this study, as series of 2D GMAs, with TPSs, were rendered on 10 skulls of T. polkensis and 48 skulls of T. bairdii, across both species’ ontogenetic series. Analyses confirmed that growth …


A System And Method For Measuring Spatially Varying Surface Appearances With A Study Of Feathers, Jessica Baron-Lis May 2025

A System And Method For Measuring Spatially Varying Surface Appearances With A Study Of Feathers, Jessica Baron-Lis

All Dissertations

Real-world materials, particularly biological structures such as feathers exhibit complex appearances that vary spatially across their surfaces. The field of computer graphics provides a means of understanding such surfaces through material modeling which uses both analytical models and data acquired from light-surface interactions. There are many efforts within the past decade in measuring materials for graphics, but common limitations in these works include not accounting for spatially varying properties and reliance on neural networks and synthetic datasets.

Feathers from modern birds present diverse appearances due to how light interacts with their unique hierarchical microstructures. Variations in those structures lead to …


Older Groundwater Reservoirs In Nebraska, Marvin P. Carlson, Steven S. Sibray Apr 2025

Older Groundwater Reservoirs In Nebraska, Marvin P. Carlson, Steven S. Sibray

Conservation and Survey Division: Faculty and Staff Publications

Sedimentary rocks below Nebraska's regional water table are saturated down to the crystalline igneous and metamorphic rocks of Precambrian age. The total thickness of these sedimentary rocks ranges from about 600 feet to over 10,000 feet. Nearly all current wells in Nebraska produce water from the near-surface, unconsolidated or partly consolidated rocks. Most studies to date -- interpretive reports, test drilling, inventory of wells, monitoring water levels, and water quality determinations -- have concentrated on these near-surface sources of groundwater.


Palynological Analysis To Interpret Inputs And Paleoenvironmental Conditions Of Tidal Flat Sediments In The Wouri Estuary, Rahmatoulaye Thialaw Diop, Fatou Penda Cisse, Maguette Ka, Onema Adojoh, Henry Agbogun Apr 2025

Palynological Analysis To Interpret Inputs And Paleoenvironmental Conditions Of Tidal Flat Sediments In The Wouri Estuary, Rahmatoulaye Thialaw Diop, Fatou Penda Cisse, Maguette Ka, Onema Adojoh, Henry Agbogun

SACAD: Scholarly Activities

This study examines the environmental significance of microfossils in specifically pollens, spore and cuticles in understanding paleoenvironment and sediments inputs in Tidal flats in the tropical Wouri estuary. Palynological slides of sediments from different depths with two sediment cores from the east and Western flanks of the Wouri estuary were examined using a trinocular microscope. Results showed the predominance of pollens families of Rhizophoraceae and Poaceae at different depths. Cuticles also varied with depth in both the West and east Wouri sediments cores. The Rhizophoraceae family are interpreted to indicate coastal and mangrove environment inputs into the sediments of the …


A Thorn In The Side: Space-Efficient Storage Of Large Ribs​, Kale Link, Carson Cope, Paige Black, Israel Rivera-Molina, Amanda Peng Apr 2025

A Thorn In The Side: Space-Efficient Storage Of Large Ribs​, Kale Link, Carson Cope, Paige Black, Israel Rivera-Molina, Amanda Peng

SACAD: Scholarly Activities

In 2022, Fort Hays State University’s Sternberg Museum of Natural History (FHSM) began a project to address the long-term preservation of a Late Miocene mammal collection. Many of the fossils from the collection were poorly consolidated, uncurated, and improperly stored, leaving them unstable and prone to significant degradation. As part of our housing overhaul, we worked to develop new methods of housing rib bones, which are often too large to fit in traditional archival boxes. Two versions of rib mounts were developed using wedges of Ethafoam (closed-cell polyethylene foam). This housing method provides a cost effective way to safely store …


Primates From Senèze, Eric Delson Apr 2025

Primates From Senèze, Eric Delson

Publications and Research

Senèze has yielded two primate fossils, neither from the recent field work. A partial ulna can be identified as cf. Macaca sylvanus. A nearly complete female skull is the holotype of Dolichopithecus arvernensis Depéret, 1928, now included in Paradolichopithecus. Emended and differential diagnoses are provided for this genus and species for the first time, and the Senèze skull is described and briefly compared to those of other species of the genus. Dental and cranial measurements are provided, along with comparative data. The macaque and Paradolichopithecus are moderately to highly terrestrial, respectively, and they suggest a woodland to more open …


Biogenic Modification Of Tidally Modified Substrates From The Micro- To Macroscopic Scale, Rogers Clark Caldwell Buntin Ii Apr 2025

Biogenic Modification Of Tidally Modified Substrates From The Micro- To Macroscopic Scale, Rogers Clark Caldwell Buntin Ii

OES Theses and Dissertations

Digital outcrop models generated by high-resolution photogrammetric imagery permit investigations into organism-substrate interactions at the centimeter to sub-millimeter scale. This dissertation uses this three-dimensional technology to study two sedimentary successions— the Archaean Dresser Formation and Cretaceous Dakota Sandstone— by capturing images via unmanned aircraft systems and single lens reflex cameras. The ~3.48 Ga Dresser Formation provides evidence for some of the earliest life on Earth in the form of stromatolites and microbially induced sedimentary structures. Aerial imagery of the 3.5 km2 Buick Geoheritage reserve capture the site’s geologic structures and close-range imagery documents stromatolites and several microbially induced sedimentary structures …


Sedimentary Archives Of Late Quaternary Limnological, Ecological, And Environmental Change In The Eastern Sierra Nevada (California, Usa), Bailee Hodelka Jan 2025

Sedimentary Archives Of Late Quaternary Limnological, Ecological, And Environmental Change In The Eastern Sierra Nevada (California, Usa), Bailee Hodelka

Theses and Dissertations--Earth and Environmental Sciences

The Sierra Nevada essentially serve as the headwaters of California, providing crucial freshwater to the state through seasonal snowmelt. This water supply is threatened by climate change in complex ways that are not fully understood. Geological records provide a valuable source of information on how the hydrology, limnology, and ecology of the Sierra Nevada region has responded to climate changes of the past. This dissertation presents new sedimentary and paleoecological research from Mono Lake and Convict Lake (Mono County, California) and aims to shed light on environmental changes in the eastern Sierra Nevada from ~16 ka to present.

At Mono …


Stratigraphy And Osl Chronology Of Lunette Deposits Indicates That Nebraska’S Rainwater Basins Formed By Aeolian Deflation Between 39 And 25 Ka, P.R. Hanson, Robert Matthew Joeckel Jan 2025

Stratigraphy And Osl Chronology Of Lunette Deposits Indicates That Nebraska’S Rainwater Basins Formed By Aeolian Deflation Between 39 And 25 Ka, P.R. Hanson, Robert Matthew Joeckel

Conservation and Survey Division: Faculty and Staff Publications

No abstract provided.


Microbial Mats In Dinosaur Ichnocoenoses, Nora Noffke, Paul C. Murphey, Roger C. Buntin, Martin G. Lockley Jan 2025

Microbial Mats In Dinosaur Ichnocoenoses, Nora Noffke, Paul C. Murphey, Roger C. Buntin, Martin G. Lockley

OES Faculty Publications

Until now, the significance of microbial mats in preservation of dinosaur tracks and in reconstructing the palaeoenvironment in which dinosaurs roamed was rarely studied. Dinosaur tracks are commonly found close to ancient aquatic bodies where moist sediment had once allowed footstep registration. At such sites, epibenthic or endobenthic microbial mats may have developed as well and may have led to exquisite track preservation. Microbial mats in coastal, lacustrine or flood plain environments cause ‘microbially induced sedimentary structures—MISS’, which are micobialites found in sandy or evaporitic settings. The microbial structures record environmental parameters including meteorological conditions that existed at the time …


Early Holocene Interaction Of Aeolian, Alluvial, And Lacustrine Processes In A Dune-Dammed Valley In The Central Nebraska Sand Hills, Gpsia Mahoney, R. D. Mandel, P.R. Hanson, Sherilyn Fritz Jan 2025

Early Holocene Interaction Of Aeolian, Alluvial, And Lacustrine Processes In A Dune-Dammed Valley In The Central Nebraska Sand Hills, Gpsia Mahoney, R. D. Mandel, P.R. Hanson, Sherilyn Fritz

Conservation and Survey Division: Faculty and Staff Publications

No abstract provided.


An Understanding Of The Feeding Ecology Of Teleoceras Fossiger And Teleoceras Aepysoma Using Dental Microwear And Mesowear Analysis, Gabriel S. Goncalves Jan 2025

An Understanding Of The Feeding Ecology Of Teleoceras Fossiger And Teleoceras Aepysoma Using Dental Microwear And Mesowear Analysis, Gabriel S. Goncalves

Master's Theses or Doctor of Nursing Practice

Teleoceras is a genus of rhinocerotids that belongs to the Tribe Teleoceratines and was widespread across North America from the late Early Miocene into the Early Pliocene. This genus was very successful, spreading across the continental United States from Florida and Tennessee to California and Washington state down to central Mexico and Honduras. Teleoceras is characterized by having a large, rounded chest cavity and short limbs with relatively high crowned teeth and lower incisors modified into tusk-like teeth. Initial ecological comparisons were made to the modern hippopotamus based on proportions of the chest cavity and its hypsodont dentition. Stable carbon …


Distribution And Tooth Morphoguilds Of Mosasaurids (Squamata) Of The Campanian Western Interior Seaway With Implications For Controls On Mosasaur Paleobiogeography, Alec Zaborniak Jan 2025

Distribution And Tooth Morphoguilds Of Mosasaurids (Squamata) Of The Campanian Western Interior Seaway With Implications For Controls On Mosasaur Paleobiogeography, Alec Zaborniak

Master's Theses or Doctor of Nursing Practice

The Western Interior Seaway (WIS) of North America is well known for its mosasaurid squamate diversity, particularly during the Campanian of the Late Cretaceous. Mosasaurid diversity has historically been examined at a fine scale, with many studies investigating the faunal composition of specific assemblages; however, studies focusing on mosasaur communities within the Western Interior Seaway during specific Ages are currently lacking. Mosasaur tooth morphology has also been extensively studied, with tooth characteristics commonly being used as phylogenetic characters. However, many studies examining mosasaur communities and tooth morphology have been focused on broader spatiotemporal or phylogenetic scales. This study investigates the …


A Histological Survey Of A Large, Sub-Adult Tylosaurus Nepaeolicus With Implications On The Validity Of Tylosaurus Kansasensis, Carson Cope Jan 2025

A Histological Survey Of A Large, Sub-Adult Tylosaurus Nepaeolicus With Implications On The Validity Of Tylosaurus Kansasensis, Carson Cope

Master's Theses or Doctor of Nursing Practice

Tylosaurus nepaeolicus is a basal, medium-sized tylosaurine mosasaur that lived during the late Coniacian and early Santonian in the Western Interior Seaway. Despite being a well-studied species, there is debate over whether Tylosaurus kansasensis represents the juvenile form of T. nepaeolicus, or if the two species are distinct. Osteohistology can determine organism age at time of death and can be used to help resolve this debate and provide a better understanding of the growth of T. nepaeolicus. Thin-sections were taken from a humerus, rib, and vertebra of a single individual (FHSM VP-2209). All thin-sections show primarily parallel-fibered cortical …


Insights On The Cranial Anatomy And Osteohistology Of Cyonosaurus, Alexander Acker Jan 2025

Insights On The Cranial Anatomy And Osteohistology Of Cyonosaurus, Alexander Acker

Dissertations

This study investigates the cranial anatomy and life history of the small-bodied gorgonopsian Cyonosaurus. Previous research on gorgonopsians has disproportionately focused on larger members of the clade, leaving smaller forms relatively understudied. This gap has led to several unresolved questions, the most pressing of which concerns gorgonopsian ontogeny. Specifically, it remains unclear whether small-bodied gorgonopsians represent genuinely diminutive adult individuals or juveniles of larger species. The specimen NHCC LB1087 presents an ideal opportunity to address this issue and gain new insights into the biology of small-bodied gorgonopsians. The first chapter focuses on cranial anatomy. Due to the specimen’s preservation, many …


The Tyrant King Of The Auction House: The Disastrous “Fine Artification” Of The Dinosaur, Amelia C. Kolstad Jan 2025

The Tyrant King Of The Auction House: The Disastrous “Fine Artification” Of The Dinosaur, Amelia C. Kolstad

Anthós

This article examines how fine art auctioneers market dinosaur fossils as art pieces to wealthy collectors, depriving the paleontological field of specimens for scientific research. Drawing from recent examples of fossils sold at fine art auctions, this article notes how rising auction prices for specimens as ‘art’ prevent museums and research institutions from acquiring specimens for research and restoration. Kolstad concludes that the art auction houses’ marketing of fossils as art is detrimental to paleontological research. To protect fossils and their scientific value, Kolstad proposes alternatives, such as selling replicas to collectors and highlighting the importance of original specimens for …


Conservation And Survey Division 2024 Annual Report, R.M. Joeckel, Michele Waszgis Jan 2025

Conservation And Survey Division 2024 Annual Report, R.M. Joeckel, Michele Waszgis

Conservation and Survey Division: Faculty and Staff Publications

The Conservation and Survey Division of the University of Nebraska - Lincoln is the agency designated by statute to investigate and interpret the geologically related natural resources of the State, to make available to the public the results of these investigations, and to assist in the development and conservation of these resources. It consists of program areas in geology, water, soils and remote sensing-geographic information systems.


Geology, Paleoecology And Vertebrate Biostratigraphy Of The Hell Creek Formation, Carter County, Montana, Usa, Joshua Collin Mathews Jan 2025

Geology, Paleoecology And Vertebrate Biostratigraphy Of The Hell Creek Formation, Carter County, Montana, Usa, Joshua Collin Mathews

Graduate Research Theses & Dissertations

The Hell Creek Formation of the Northern Great Plains records one of the greatest mass extinction events in Earth’s history, bringing to a close the reign of the dinosaurs at the end of the Cretaceous period. This rock unit comprises one of the most studied geological formations in the world and is one of the few locations on the planet that preserve terrestrial deposits containing ejecta from the bolide impact that slammed into the Yucatan Peninsula 66 million years ago. This profound event in the geologic record has led to numerous studies of paleobiotas, climate change, ecosystem recovery dynamics, and …


A Paleobiological And Paleoecological Analysis Of Mississippian Trilobites With Segmentation Abnormalities, April Catherine Cerami Jan 2025

A Paleobiological And Paleoecological Analysis Of Mississippian Trilobites With Segmentation Abnormalities, April Catherine Cerami

Graduate Theses, Dissertations, and Problem Reports (ETD)

Trilobites are a well-preserved group of arthropods, with a fossil record extending from the Cambrian to the end of the Permian. Abnormalities, such as injuries or teratological defects, have been observed and described in multiple individuals, although investigating how these abnormalities occur in populations has been largely overlooked.

One possible cause of abnormalities in marine arthropods is heavy metal pollution. Such metals can leave individuals more susceptible to teratological deformities and injuries, interfering with the molting and healing process. Here, methods for analyzing elemental composition and preservation of the trilobite cuticle were refined, finding that 30 μm thin sections are …


Late Ordovician Evolution Of Sea Scorpions: Eurypterids Of The Big Hill Lagerstätte, Joseph Donaldson Bingham Jr. Jan 2025

Late Ordovician Evolution Of Sea Scorpions: Eurypterids Of The Big Hill Lagerstätte, Joseph Donaldson Bingham Jr.

Graduate Theses, Dissertations, and Problem Reports (ETD)

Eurypterids are a group of diverse chelicerates that first appeared during the Middle Ordovician (467.3 Ma) and went extinct at the end of the Permian (251.9 Ma). The fossil record of eurypterids during the Ordovician is relatively sparce, comprising only 12 species representing the Megalograptidae, a ‘waeringopterid’ clade, Rhenopteridae, Onychopterellidae, Adelophthalmoidea, and Eurypteroidea clades. This study describes early eurypterids from the Katian aged (~452 Ma) Big Hill Lagerstätte of Michigan. The described material represents two new species, herein named Machipterus margaritae sp. nov. (based on 21 specimens preserving the ventral prosoma and prosomal appendages, and opisthosoma) and Dikranopterus lysinae sp. …


Visualization Of Paleocurrents On A Web Application Using Gplates, Anjan Sapkota Dec 2024

Visualization Of Paleocurrents On A Web Application Using Gplates, Anjan Sapkota

MS in Computer Science Theses

Paleocurrents are flow directions derived from features of sedimentary rocks that reveal the direction of the current of wind or water that deposited the sediment. In 2015, Brand et al. created a global database of paleocurrents, which contains over 1,000,000 measurements worldwide: North America, South America, Australia, Great Britain, parts of Western Europe, China, Africa are fairly well represented; Antarctica, Eastern Europe, and Asia are modestly represented and Russia is poorly represented. The contribution of this thesis is a web application that uses the GPlates’ Application Programming Interface (API) to visualize global paleocurrents through time in an interactive way based …


A Day In The Badlands: A 52 Million-Year-Old Safari Adventure, Devra Hock-Reid, Judy Diamond, Jenn Paul Dec 2024

A Day In The Badlands: A 52 Million-Year-Old Safari Adventure, Devra Hock-Reid, Judy Diamond, Jenn Paul

Zea E-Books Collection

with Amy Chew, Christopher Gilbert, Ross Secord, and Stephen Chester

Illustrations by Jenn Paul • Book design by Aaron Sutherlen

About the Badlands Fossils -- This story tells about the fossils of ancient animals found in the Wind River and Bighorn basins of Wyoming. Wyoming’s rich fossil record tell us about amazing animals from millions of years ago. About 66 million years ago, an asteroid slammed into the Earth, causing devastating changes that led to the extinction of dinosaurs. As the planet began to recover, it was much warmer than today, allowing palm trees and other tropical plants to grow …


Traittrainr: Accelerating Large-Scale Simulation Under Models Of Continuous Trait Evolution, Jenniffer Roa Lozano, Mataya Duncan, Duane D. Mckenna, Todd A. Castoe, Michael Degiorgio, Richard Adams Dec 2024

Traittrainr: Accelerating Large-Scale Simulation Under Models Of Continuous Trait Evolution, Jenniffer Roa Lozano, Mataya Duncan, Duane D. Mckenna, Todd A. Castoe, Michael Degiorgio, Richard Adams

Entomology and Plant Pathology Faculty Publications and Presentations

Abstract

Motivation

The scale and scope of comparative trait data are expanding at unprecedented rates, and recent advances in evolutionary modeling and simulation sometimes struggle to match this pace. Well-organized and flexible applications for conducting large-scale simulations of evolution hold promise in this context for understanding models and more so our ability to confidently estimate them with real trait data sampled from nature. Results

We introduce TraitTrainR, an R package designed to facilitate efficient, large-scale simulations under complex models of continuous trait evolution. TraitTrainR employs several output formats, supports popular trait data transformations, accommodates multi-trait evolution, and exhibits flexibility …


Invertebrate Functional Diversity Of Fossils From Arrow Canyon, Nevada, Nathaniel Valentine Nov 2024

Invertebrate Functional Diversity Of Fossils From Arrow Canyon, Nevada, Nathaniel Valentine

Undergraduate Research Symposium Podium Presentations

Paleontology can be used to study how ecosystems evolve and respond to changes in the environment. Here we examine invertebrate fossils from the Pennsylvanian, a time period that ranged from 323.2 to 298.9 million years ago, which are preserved along the walls of Arrow Canyon (Nevada) in the Bird Spring Formation. We surveyed three sites within the canyon, and used data from the Paleobiology Database to create a list of taxa found on the canyon from this time interval. Although ecosystems have traditionally been studied using the composition of the ecosystem (the number of species), recent work has shown that …


Tooth Type And Position In A Hypothetical Pectinodon..., Keith Snyder, Jonathon Campos, Neal Doran, Charity Simmons Sep 2024

Tooth Type And Position In A Hypothetical Pectinodon..., Keith Snyder, Jonathon Campos, Neal Doran, Charity Simmons

Achieve

In 1856, Leidy rst identied Troodontidae as a taxonomic unit. It was not until 1945 that C.M. Sternberg assigned it to a family status. There has been much debate on taxonomic naming in this family, but in 1987, Currie conated all previous names into a single species, Troodon formosus. On the basis of isolated teeth, Carpenter (1982) suggested a second species, Pectinodon bakkeri from the Lance Formations. With limited specimens (24), Longrich (2008) proposed tooth positioning for the 4 tooth types he dened. This was especially challenging since no cranial or post-cranial skeletal structures have yet been identied.

Over the …