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Articles 1 - 30 of 171
Full-Text Articles in Paleobiology
The Hitchhiker's Guide To The Host: Fleas And The History Of Pikas In North America, Josh Pletcher
The Hitchhiker's Guide To The Host: Fleas And The History Of Pikas In North America, Josh Pletcher
All NMU Master's Theses
Parasite phylogenetics can offer windows into the cryptic evolutionary history of hosts, often highlighting events that are not recorded by the phylogenetics of the host. Pikas (small lagomorphs) host a diverse parasite community comprising endoparasites (e.g., coccidians, nematodes, and cestodes) and ectoparasites (e.g., fleas and mites). Phylogenetic analysis of pika endoparasites has revealed pikas evolved in Asia and colonized North America in two waves, and that contact between these two pika lineages allowed parasites to be transferred from one lineage to another and to survive the extinction of their original host. To uncover these cryptic histories, understanding the biogeography of …
Ontogeny And The Effects Of Perinatal Stress In The First Immature Skeleton Of Teleoceras Aepysoma (Perissodactyla, Rhinocerotidae), Joe Gajewski
Electronic Theses and Dissertations
Recovery of an immature fossil rhinoceros (Teleoceras aepysoma) skeleton (ETMNH 32999) from the Gray Fossil Site in Tennessee offers the first insights into the ontogeny of the species. ETMNH 32999 is interpreted to be a subadult male in an age class equivalent to ~17-20% of potential lifespan. Comparative analyses support the hypothesis that subadults of related taxa are typically ~91% of average adult size in most dimensions. However, certain cranial dimensions are substantially lower than expected in ETMNH 32999, averaging only ~74% of the average conspecific adult. Such departure from typical growth rates is suggested to be attributable …
Overview And Methods Of Molecular Paleontology: A Review, Shelby Touchstone
Overview And Methods Of Molecular Paleontology: A Review, Shelby Touchstone
Williams Honors College, Honors Research Projects
Molecular paleontology—the study of ancient biomolecules preserved in fossils and sediments, including ancient DNA, lipids, and proteins—has motivated researchers since the 1950s, inspiring both scientific inquiry and broader cultural imagination. Think Jurassic Park. This honors thesis presents a comprehensive review of the history, current state, and methods of the field of molecular paleontology. Drawing on peer-reviewed literature and scholarly resources accessed through databases such as EBSCO and Bierce library, this review examines key techniques such as Next Generation Sequencing and amino acid racemization and will go over limitations and controversies in the field. It will also explore potential new …
Ediacaran Paleocommunity Structure, Body Size Evolution, And The Rise Of Complex Animals, Matthew Philip Craffey
Ediacaran Paleocommunity Structure, Body Size Evolution, And The Rise Of Complex Animals, Matthew Philip Craffey
Dissertations and Doctoral Documents, University of Nebraska-Lincoln, 2023–
The Ediacaran period (635-538ma) is a key interval in the evolution of life as it records the first macroscopic animal communities. Major extinction pulses appear to have altered the composition of the global marine biosphere between each sub-interval of the Ediacaran Period: the Avalon, White Sea, and Nama assemblage Zones. While considerable attention has been paid to the causes of these extinction pulses, less has been afforded to the consequences of Ediacaran biotic turnover for community organization and function. Moreover, it is unclear how body size - the defining trait of macrofauna - affected the transition between animal groups across …
Primates From Senèze, Eric Delson
Primates From Senèze, Eric Delson
Publications and Research
Senèze has yielded two primate fossils, neither from the recent field work. A partial ulna can be identified as cf. Macaca sylvanus. A nearly complete female skull is the holotype of Dolichopithecus arvernensis Depéret, 1928, now included in Paradolichopithecus. Emended and differential diagnoses are provided for this genus and species for the first time, and the Senèze skull is described and briefly compared to those of other species of the genus. Dental and cranial measurements are provided, along with comparative data. The macaque and Paradolichopithecus are moderately to highly terrestrial, respectively, and they suggest a woodland to more open …
An Understanding Of The Feeding Ecology Of Teleoceras Fossiger And Teleoceras Aepysoma Using Dental Microwear And Mesowear Analysis, Gabriel S. Goncalves
An Understanding Of The Feeding Ecology Of Teleoceras Fossiger And Teleoceras Aepysoma Using Dental Microwear And Mesowear Analysis, Gabriel S. Goncalves
Master's Theses or Doctor of Nursing Practice
Teleoceras is a genus of rhinocerotids that belongs to the Tribe Teleoceratines and was widespread across North America from the late Early Miocene into the Early Pliocene. This genus was very successful, spreading across the continental United States from Florida and Tennessee to California and Washington state down to central Mexico and Honduras. Teleoceras is characterized by having a large, rounded chest cavity and short limbs with relatively high crowned teeth and lower incisors modified into tusk-like teeth. Initial ecological comparisons were made to the modern hippopotamus based on proportions of the chest cavity and its hypsodont dentition. Stable carbon …
Human Impacts On Mammal Diversity From The Pleistocene To The Present, Carson P. Hedberg
Human Impacts On Mammal Diversity From The Pleistocene To The Present, Carson P. Hedberg
Biology ETDs
Humans have fundamentally altered the planet’s biodiversity. It is critical to characterize these changes, but also to understand how they impact the structure and function of modern ecosystems. This dissertation investigates how anthropogenically mediated species losses and/or introductions influence ecological function and community structure in mammals across multiple temporal and spatial scales. We found that selective loss of large-bodied mammals over the Late Quaternary has led to severe losses in functional diversity in modern ecosystems, suggesting megafaunal mammals contribute distinct ecological functions. These changes to mammal diversity and distribution over millennia have altered both global patterns in functional diversity and …
Sub-Stage Climatic Shifts During Mis 11 Refined From Diatom Assemblage Reconstruction In The Valles Caldera, New Mexico, Savannah Cutler
Sub-Stage Climatic Shifts During Mis 11 Refined From Diatom Assemblage Reconstruction In The Valles Caldera, New Mexico, Savannah Cutler
Earth and Planetary Sciences ETDs
Diatom assemblage reconstruction from the Valles Caldera sediment core (VC-3) was utilized to infer climatic and aquatic changes during a 50-thousand-year [ka] period in the Mid-Pleistocene (435-385 ka). This 50 ka interval spans most of Marine Isotope Stage 11 (MIS 11) – one of the most prominent interglacials of the past 500 ka – and is widely considered an analog for the Holocene and future climate regimes. Nearly all records from MIS 11 are either marine or ice-core based, making VC-3 unique among lacustrine sediment records as it captures insolation variations and climatic changes during the mid-Pleistocene. Analysis of the …
Consequences Of The Megafauna Extinction: Changes In Food Web Networks On The Edwards Plateau Across The Pleistocene–Holocene Transition, Quentin A. Smith Jr.
Consequences Of The Megafauna Extinction: Changes In Food Web Networks On The Edwards Plateau Across The Pleistocene–Holocene Transition, Quentin A. Smith Jr.
School of Biological Sciences: Dissertations, Theses, and Student Research
We are experiencing biodiversity loss due to climate change and human impacts, which is not only harmful to the environment but can also alter the composition of communities and interactions among species. The late Pleistocene experienced a loss of large-bodied mammals which resulted in significant changes in community structure due to changes in body size, diet, and species associations. The impact of these changes on species interactions and community structure across the Pleistocene-Holocene transition remains poorly understood. Using a robust data set of species composition, stable isotopes, body size, and climate variables, we constructed and compared ecological networks of mammal …
New Species Of Dryolestoid From The Late Cretaceous Allen Formation And Implications For South American Faunal Diversity., Brigid Erin Connelly
New Species Of Dryolestoid From The Late Cretaceous Allen Formation And Implications For South American Faunal Diversity., Brigid Erin Connelly
Electronic Theses and Dissertations
Dryolestoids are extinct cladotherians mammals from the Jurassic and Cretaceous periods. I describe a collection of dryolestoid specimens from the Late Cretaceous localities of Cerro Tortuga (Allen Formation), Anfiteatro 1, and Shining (both La Colonia Formation) from Patagonia, Argentina. Using comparative morphology, I identify a new species of meridiolestidan dryolestoid based on eleven specimens across both formations. The new species’ recovery from La Colonia Formation represents the first dryolestoid connection between the two approximately contemporaneous formations. The species’ morphology may represent an ecological shift within Meridiolestida from insectivory to herbivory, showing a transition in characters between the plesiomorphic sharp-toothed meridiolestidans …
Geometric Morphometric Analysis Of Modern Viperid Vertebrae Facilitates Identification Of Fossil Specimens, Lance D. Jessee
Geometric Morphometric Analysis Of Modern Viperid Vertebrae Facilitates Identification Of Fossil Specimens, Lance D. Jessee
Electronic Theses and Dissertations
Snake vertebrae are common in the fossil record, whereas cranial remains are generally fragile and rare. Consequently, vertebrae are the most commonly studied fossil element of snakes. However, identification of snake vertebrae can be problematic due to extensive variation. This study utilizes 2-D geometric morphometrics and canonical variates analysis to 1) reveal variation between genera and species and 2) classify vertebrae of modern and fossil eastern North American Agkistrodon and Crotalus. The results show that vertebrae of Agkistrodon and Crotalus can reliably be classified to genus and species using these methods. Based on the statistical analyses, four of the …
Exploring The Relationships Between Mammalian Functional Trait Distributions And Regional Biomes, With Application To Miocene Paleoecology, Devra Hock-Reid
Exploring The Relationships Between Mammalian Functional Trait Distributions And Regional Biomes, With Application To Miocene Paleoecology, Devra Hock-Reid
Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research
Paleoecology relies on understanding relationships between modern animals and their environment. Animals are adapted to niches in their environments, and those physical adaptations, or functional traits, are utilized as proxies to interpret aspects of paleo-ecosystems. Much is known about individual functional traits in extant mammals and their relationship to the environment. Less is known about how multiple functional traits across a community can be utilized for paleoecological interpretations. I develop models utilizing traits in mammalian communities at the biome level. For Chapter 1, I build a model for North American regional biomes using mammalian trait frequencies. I quantify changes in …
Holocene Rice Rats (Genus Oryzomys) From The Upper Mississippi River Drainage Basin, Hugh H. Genoways
Holocene Rice Rats (Genus Oryzomys) From The Upper Mississippi River Drainage Basin, Hugh H. Genoways
Zea E-Books Collection
The expansion and collapse of the geographic range of the Texas rice rat (Oryzomys texensis) in the upper Mississippi River drainage basin at the end of the Holocene was a unique event in North American mammals. In a period of about 4000 years with a point of origin near the American Bottom in Illinois, these small rodents extended their geographic range in a straight-line distance of over 950 km to the west into Nebraska and the same distance to the east into Pennsylvania. Then in less than 400 years this range expansion collapsed back to a point where …
Reconstructing The Ecological Relationships Of The Latest Cretaceous Antarctic Dinosaurs And How Functional Tooth Morphology Influenced Diet And Ecological Niche Among Basal Ornithopod Dinosaurs, Ian Broxson
EWU Masters Thesis Collection
Of the final three connected Gondwanan landmasses, the dinosaur fossil record of Antarctica in the Cretaceous is the least complete. Most dinosaur faunas in this time period (145 Ma to 66.0 Ma) are widely separated geographically and temporally from one another by million years. However, a group of non-avian dinosaurs from the James Ross Basin (JRB) of the Antarctic Peninsula, composed of two elasmarians, a parankylosaurian ankylosaur, a hadrosaur and a suspected megaraptor, all are represented by fragmentary remains have emerged from the same horizon of the Sandwich Bluff Member (SBM) of the López de Bertodano Formation and were thus, …
The Anatomy And Phylogeny Of A New Large Plioplatecarpine Mosasaur From The Campanian Bearpaw Shale Of Montana (Usa), Richard A. Carr
The Anatomy And Phylogeny Of A New Large Plioplatecarpine Mosasaur From The Campanian Bearpaw Shale Of Montana (Usa), Richard A. Carr
Master's Theses or Doctor of Nursing Practice
In 2018, a large and associated plioplatecarpine mosasaur skull, pectoral girdle, and rib cage, whose total body length may have exceeded five meters, was uncovered in the Late Campanian Bearpaw Shale of Northeast Montana (USA). Phylogenetic analysis of this specimen, MOR 10855, recovers this individual as a basal member of the genus Plioplatecarpus. This specimen, is unique in that it is estimated to be nearly twice the size of any of the other species of Plioplatecarpus found in the Western Interior Seaway during this part of the Cretaceous. While the included phylogenetic study suggests MOR 10855 represents a new …
Investigation Of Dead Ocean Quahogs (Arctica Islandica) Shells On The Mid-Atlantic Bight Continental Shelf, Alyssa Leclaire
Investigation Of Dead Ocean Quahogs (Arctica Islandica) Shells On The Mid-Atlantic Bight Continental Shelf, Alyssa Leclaire
Master's Theses
Ocean quahogs, Arctica islandica, are a long-lived, widely dispersed, biomass dominate in the Mid-Atlantic; therefore, quahog shells are valuable resources for studying climate change over time. Recently, dead ocean quahog shells were discovered south and inshore of the present biogeographic range of this animal. The presence of ocean quahog shells outside the current range is presumably a consequence of past regressions and transgressions of the Cold Pool, the bottom-trapped, cool body of water that allows boreal animals to live at lower latitudes. Dead ocean quahog shells were collected offshore of the DelMarVa Peninsula then radiocarbon-dated, evaluated for taphonomic condition, …
Reconstructing The Ecological Relationships Of Late Cretaceous Antarctic Dinosaurs And How Functional Tooth Morphology Influenced These Relationships, Ian D. Broxson
2022 Symposium
The Sandwich Bluff Formation of the James Ross Basin of Antarctica has recently yielded a group of five late Cretaceous dinosaurs that lived contemporaneously with each other, a first for Antarctica. These five dinosaurs include fragmentary remains of two differently sized elasmarian ornithopods, a possible megaraptor, a hadrosaur, and a nodosaur. In this study we will construct a model of the ecological relationships of late Cretaceous Antarctica. Additionally, we will look at what specific factors allowed this group of four herbivores and a carnivore to coexist in a restricted locality and what niches were filled by each species. Methods to …
Ontogenetic Niche Shift As A Driver Of Community Structure And Diversity In Non-Avian Dinosaurs, Katlin M. Schroeder
Ontogenetic Niche Shift As A Driver Of Community Structure And Diversity In Non-Avian Dinosaurs, Katlin M. Schroeder
Biology ETDs
As some of the most charismatic megafauna to ever walk the earth, the physiology, morphology, growth and evolution of non-avian theropods has been studied exhaustively, yet little is understood about their roles in ecosystems as juveniles. For carnivorous megatheropods, which exceed 1,000kg in mass yet hatched from eggs of limited size, the likelihood of utilizing different prey through ontogeny was high, simply by proxy of the immense difference in size between adults and juveniles. We found these ontogenetic niche shifts, evidenced by significantly different dental microwear in Tyrannosaurids, to have excluded dinosaurian mesocarnivores from Mesozoic communities. The few dinosaurian mesocarnivores …
Rodent Dental Microwear Texture Analysis As A Proxy For Fine-Scale Paleoenvironment Reconstruction, Jenny H. E. Burgman
Rodent Dental Microwear Texture Analysis As A Proxy For Fine-Scale Paleoenvironment Reconstruction, Jenny H. E. Burgman
Graduate Theses and Dissertations
Dental microwear texture analysis (DMTA) of fossil fauna has become a valuable tool for dietary inference and paleoenvironment reconstruction. Most of this work has utilized larger taxa with larger home ranges. These studies may result in broader-scale habitat inferences that could mask the details of complex mosaic habitats. Rodent DMTA offers an opportunity to work at finer spatial scales because most species have smaller home ranges. Rodents are also keystone species within their ecosystems, abundant, ubiquitous, and found in many fossil deposits. These attributes make them excellent proxies for environmental reconstructions. However, the application of DMTA to rodents remains relatively …
Distinguishing Mustela From Neogale (Mustelidae) Through Both A Qualitative And Quantitative Analysis Of Skull And Tooth Morphology, Ronald W. Peery
Distinguishing Mustela From Neogale (Mustelidae) Through Both A Qualitative And Quantitative Analysis Of Skull And Tooth Morphology, Ronald W. Peery
Electronic Theses and Dissertations
Weasels and mink (Mustela and Neogale) can be difficult to distinguish osteologically due to similarities in morphology, thus suggesting the need for an accurate tool in distinguishing among taxa. This study utilized a combination of character state and stepwise discriminant function (DFA) analyses to examine potential distinguishing features of skull and tooth morphology. Measurements and ratios were collected from all 18 extant musteline species, as well as the extinct Neovison macrodon, Mustela rexroadensis, Mustela meltoni, Mustela gazini, and Mustela jacksoni. Unidentified musteline specimens from the Gray Fossil Site were also included. Results of …
Effects Of Environmental Change On Ancestral Pueblo Fishing In The Middle Rio Grande, Jonathan William Dombrosky
Effects Of Environmental Change On Ancestral Pueblo Fishing In The Middle Rio Grande, Jonathan William Dombrosky
Anthropology ETDs
It has long been assumed that fishes were unimportant in the diet of past Pueblo people in the U.S. Southwest. Yet, small numbers of fish remains are consistently recovered from Late pre-Hispanic/Early Historic archaeological sites in the Middle Rio Grande of New Mexico. The end of drought conditions may have impacted food choice and fishing decisions during this time. I use behavioral ecology to understand how fishing could have been an optimal food-getting strategy for Ancestral Pueblo farmers. Stable isotope analysis offers a way to account for environmental change. I provide a refined 13C Suess correction model to support …
Terrestrial Soldier Crab (Coenobita Clypeatus, Fabricius 1787) And Cerion Spp. (Röding 1798) Shell Relationship On San Salvador Island, Bahamas, Harley Hunt
Biology Theses
The Caribbean terrestrial soldier crab, Coenobita clypeatus(Fabricius 1787), coexist and utilize the shells of numerous species of land and marine gastropods. Soldier crabs rely on gastropod shells for protection as the crabs have a soft abdomen, leaving them vulnerable for predation and desiccation, threatening their survival. This creates a strong pressure to obtain well-fitting shells that provide adequate protection against water loss. Cerion of Röding (1798) shells are one of the most commonly used shells among living colonies of C. clypeatuson San Salvador Island. This study is interested in the frequency of shell use by C. clypeatus crabs …
Taphonomy Of Late Jurassic (Tithonian) Morrison Formation Apatosaurus Sp. Vertebrae Found Associated With Teeth From Allosaurus Sp. And Ceratosaurus Sp., And Body Size Extrapolation From The Associated Theropod Teeth., Greg C. Agyan
All Graduate Plan B and other Reports, Spring 1920 to Spring 2023
An Apatosaurus sp. locality from Dinosaur National Monument designated DNM-15 was excavated in 1985, and associated with two Allosaurus teeth and one Ceratosaurus tooth that were near one of the caudal vertebrae. The Ceratosaurus tooth was buried between an overlying rib and that same caudal vertebra. The caudal vertebrae of the DNM-15 Apatosaurus were intact and articulated, but the anterior skeleton was mostly absent, with a row of articulated sacral vertebrae in close association with a femur. Two other Allosaurus teeth were reported near the preserved ilium of the Apatosaurus, but they could not be located in the collections. …
Holocene Records Of Nebraska Mammals, Hugh H. Genoways
Holocene Records Of Nebraska Mammals, Hugh H. Genoways
Zea E-Books Collection
A survey of the archeological and paleontological literature allowed a compilation of Holocene records of mammals in Nebraska. This survey identified Holocene records from 338 sites in 62 of the 93 Nebraska counties. These counties were located throughout state, but there was a concentration of sites in southwestern Nebraska where there were 27 fossil sites in Frontier County and 22 in Harlan County. Fossils sites were underrepresented in the Sand Hills region. Records of fossil mammals covered the entire Holocene period from 13,000 years ago until AD 1850. A minimum of 57 species (with eight additional species potentially present) representing …
Ecological Controls On The Campanian Distribution Of Hesperornis (Aves: Hesperornithiformes) In The Western Interior Seaway, Blake Chapman
Ecological Controls On The Campanian Distribution Of Hesperornis (Aves: Hesperornithiformes) In The Western Interior Seaway, Blake Chapman
Master's Theses or Doctor of Nursing Practice
The epicontinental Western Interior Seaway (WIS) of Late Cretaceous North America provided a unique marine habitat for cephalopods, fish, marine reptiles, and the foot-propelled diving seabird Hesperornis. While several predator-prey relationships among Hesperornis or other hesperornithiforms and other WIS animals have been hypothesized based on gut contents, bite marks, and coprolites/colonites, ecological relationships have not been quantitatively tested. Paleontological species distribution modeling (SDM) studies have focused on extinct non-marine taxa and marine invertebrates, with only two marine vertebrate studies of extant taxa. Here, two SDM methods were used to test the influence of vertebrate faunas, sedimentary rock type, paleogeography, …
African Land Mammal Ages, John Van Couvering, Eric Delson
African Land Mammal Ages, John Van Couvering, Eric Delson
Publications and Research
We define 17 African land mammal ages, or AFLMAs, covering the Cenozoic record of the Afro-arabian continent, the planet’s second largest land mass. While fossiliferous deposits are absent on the eroded plateau of the continent’s interior, almost 800 fossil genera from over 350 locations have now been identified in coastal deposits, karst caves, and in the Neogene rift valleys. Given a well-developed geochronologic framework, together with continuing revision to the fossil record—both stimulated by the story of human evolution in Africa—and also to compensate for the variation in fossil ecosystems across such great distances, the AFLMAs are biochronological units defined …
Faunal Remains From Archaeology Sites In Southwestern New Mexico, Karen Gust Schollmeyer, S. O. Macdonald
Faunal Remains From Archaeology Sites In Southwestern New Mexico, Karen Gust Schollmeyer, S. O. Macdonald
Occasional Papers
This study attempts to create a comprehensive catalog of all known records of faunal remains recovered from 105 archaeological sites located across southwestern New Mexico. These sites range in age from 2000BC to AD 1450. Approximately 250 taxa of mollusks, fishes, amphibians, reptiles, birds and mammals are represented in this compilation. Primary reference materials from published and unpublished sources are included, along with the repositories where relevant collections may be found for potential reexamination by researchers from a variety of fields of interest.
Paleoenvironmental Reconstruction Of Quaternary Saltville, Virginia, Using Ostracode Autecology, Austin Gause
Paleoenvironmental Reconstruction Of Quaternary Saltville, Virginia, Using Ostracode Autecology, Austin Gause
Electronic Theses and Dissertations
The Saltville valley in southwestern Virginia is home to Quaternary localities containing paleontological and archaeological remains. Historically the valley has been mined for salt and the small lakes, ponds and springs along the valley floor have a brackish signature. A preliminary report on the site’s ostracode fauna suggested that the site’s water was not always saline. This study analyzed modern and Quaternary ostracodes to understand the valley’s hydrologic and chemical evolution. Sediments contained primarily freshwater species, including the environmentally sensitive Candona crogmaniana. The presence of Pelocypris tuberculatum and a new Fabaeformiscandona species throughout a vertical section spanning the latest Pleistocene …
Encephalization In Commensal Raccoons: A Unique Test Of The Cognitive Buffer Hypothesis, Peter M. Anderson
Encephalization In Commensal Raccoons: A Unique Test Of The Cognitive Buffer Hypothesis, Peter M. Anderson
Electronic Theses and Dissertations
This study investigated selective pressures associated with encephalization in mammals and discussed broader implications. Relative brain size as measured by EQ (Encephalization Quotient) was compared between ecological categories. Omnivores had higher average EQ than ecological specialists. Since specialists are disproportionately affected by extinction events, selection for ecological generalism is proposed as encephalization mechanism. This mechanism may reinforce the more widely known Cognitive Buffer Hypothesis (CBH)—the idea that possessing relatively large brains has buffered lineages against environmental change. CBH is tested here by comparing EQs in Procyon lotor (raccoon) in urban and rural environments. CBH predicts that raccoons in the most …
A Tale Of Two Species: Black-Tailed And White-Tailed Prairie Dog Biogeography From The Last Interglacial To 2070, April Dawn Bledsoe
A Tale Of Two Species: Black-Tailed And White-Tailed Prairie Dog Biogeography From The Last Interglacial To 2070, April Dawn Bledsoe
Electronic Theses and Dissertations
Ecological niche models (ENMs) were created for White-tailed and Black-tailed prairie dogs and projected into the Last Interglacial (LI), the Last Glacial Maximum (LGM), and mid-Holocene (mid-H) to discern possible past suitable habitat for both species. Additionally, ENMs were projected into the future year 2070 representative concentration pathways (RCPs) 2.6 and 8.5 to discern how climate change may affect future habitat suitability. Kernel density estimations, minimum convex polygons, and median distribution centers of White-tailed and Black-tailed occurrence records were examined between time-periods to discern the effects of anthropogenic westward expansion on both species’ distributions. Current ENMs were constructed from commonly …