Marine Biology Commons^™

Variations In Otolith-Zinc Concentrations Of Brown Surgeonfish (Acanthurus Nigrofuscus) Between Marine Protected Areas And Fished Reefs, Matthew Austin Nichols

Theses and Dissertations

As human populations continue to increase, the growing reliance on exploited fish stocks has furthered patterns of declining body size and biomass worldwide. To mitigate these impacts, fisheries managers have implemented a variety of conservation strategies, including marine protected areas (MPA), as these serve as a refuge for fish, resulting in increases in numerous health metrics (fish density, biomass, body-size, and species diversity). By using fish otolith samples previously concluded to have physiological and morphometric differences between MPA and fished reef (FR) fish, this thesis explores the potential of using microchemical analysis of zinc (Zn) to reveal impacts MPAs might …

Tile Fish (Lopholatilus Villarii) Neurocranium, Maeya C. Yeatman, Sebastian K. Chan, David W. Kerstetter

All Scans: Kerstetter Fisheries and Avian Ecology 3D Scan Series

Neurocranium from 12 cm total length (TL) Tile fish (Lopholatilus villarii), collected from Foodtown on 10/22/2021 by Maeya Yeatman and Sebastian Chan

Snook Neurocranium Scan Mello & Glanzmann, Christopher F. Glanzmann, Madison A. Mello

All Scans: Kerstetter Fisheries and Avian Ecology 3D Scan Series

This file is a 3D scan of a Snook Neurocranium. Snook head provided by DR. Kerstetter and scan performed by Chris Glanzmann and Madison Mello. Medium detail scan with a simplification ratio of 50.

Methodological Challenges In Studying Trust In Natural Resources Management, Antonia Sohns, Gordon M. Hickey, Jasper R. De Vries, Owen Temby

School of Earth, Environmental, & Marine Sciences Faculty Publications

Trust has been identified as a central characteristic of successful natural resource management (NRM), particularly in the context of implementing participatory approaches to stakeholder engagement. Trust is, however, a multi-dimensional and multi-level concept that is known to evolve recursively through time, challenging efforts to empirically measure its impact on collaboration in different NRM settings. In this communication we identify some of the challenges associated with conceptualizing and operationalizing trust in NRM field research, and pay particular attention to the inter-relationships between the concepts of trust, perceived risk and control due to their multidimensional and interacting roles in inter-organizational collaboration. The …

Binational Reflections On Pathways To Groundwater Security In The Mexico–United States Borderlands, Rosario Sanchez, José Agustin Breña-Naranjo, Alfonso Rivera, Randall T. Hanson, Antonio Hernández-Espriú, Rick J. Hogeboom, Anita Milman, Jude A. Benavides, Adrian Pedrozo-Acuña, Julio Cesar Soriano-Monzalvo

School of Earth, Environmental, & Marine Sciences Faculty Publications

Shared groundwater resources between Mexico and the United States are facing unprecedented stressors. We reflect on how to improve water security for groundwater systems in the border region. Our reflection begins with the state of groundwater knowledge, and the challenges groundwater resources face from a physical, societal and institutional perspective. We conclude that the extent of ongoing cooperation frameworks, joint and remaining research efforts, from which alternative strategies can emerge, still need to be developed. The way forward offers a variety of cooperation models as the future offers rather complex, shared and multidisciplinary water challenges to the Mexico–US borderlands.

Dioxins Contamination In Sediment And Their Effect On Accumulation In The Portuguese Oyster (Crassostrea Angulata) And White Shrimp (Litopenaeus Vannamei), Yin-Yu Chen, Yu-Sheng Wu, Hsiang-Yin Chen, Zhen-Hao Liau, Fan-Hua Nan

Journal of Marine Science and Technology–Taiwan

This research aims to realize whether the dioxin accumulation in the oyster and white shrimp which were cultured in the dioxin pollutant sediment. This presented evidence indicated that dioxin was detected in all experimental groups with high concentration accumulated in experiment animals. Observation of the concentration, the PCDD/F, 2,3,7,8-T4CDD was with high concentration in the sediment and declined with the time elapsed. Detection of the dioxin accumulated in the animal, the accumulative maximum amounts was in the oyster as 3746.16 (pg-TEQ/g d.w.) and in the shrimp as 705.44 (pg-TEQ/g d.w.). The detected amount of dioxin in the supernatant was quite …

A Meshless Method With Radial Basis Function For Solving Unsaturated Flow In Heterogeneous Porous Media, Cheng-Yu Ku, Chih-Yu Liu, Wei-Po Huang, Jing-En Xiao

Journal of Marine Science and Technology–Taiwan

This paper presents a study for solving unsaturated flow in heterogeneous porous media using the meshless method with the radial basis function (RBF). For modeling the nonlinear hydrological process in unsaturated zone, an exponential model is introduced in the Richards equation such that we may obtain the linearized Richards equation. We adopt the multiquadric function as the RBF in the meshless method for solving the linearized Richards equation. For simulating the unsaturated flow problems in layered heterogeneous soils, the flux and the head must satisfy the continuity condition at the interface. Several examples are carried out for modeling the hydrological …

Catch Rates And Distribution Pattern Of The Silky Shark, Carcharhinus Falciformis, Caught By The Taiwanese Large-Scale Longline Fishery In The Indian Ocean, Chia Yun Li, Xing-Han Wu, Shang Yin Vanson Liu, Sheng-Ping Wang, Wen-Pei Tsai

Journal of Marine Science and Technology–Taiwan

The silky shark, Carcharhinus falciformis, is widely distributed in tropical and temperate waters, and it is a common bycatch species for tuna longline fisheries. This study examined the distribution of and presents relative abundance indices of the silky shark in the Indian Ocean by using logbook and observer data from the Taiwanese large-scale tuna longline fishery between 2005 and 2019. Due to the high zero catch rate, a zero-inflated negative binomial (ZINB) model was used to standardize catch per unit effort. Due to a lack of detailed targeting information, the fishery strategy was identified by using cluster analysis based on …

Accumulation And Bioavailability Of Heavy Metals (Cadmium, Chromium, Nickel, Lead And Zinc) In Water And Sediment Samples In The Musa Creek, Northwest Coasts Of The Persian Gulf, Saghafiankhoo Sedigheh, Battaleb-Looie Sedigheh, Amanipoor Hakimeh, Dravishi Khatooni Javad, Ghanemi Kamal

Journal of Marine Science and Technology–Taiwan

The current study aimed to investigate the total concentrations of heavy metals in water and sediment samples as well as their bioavailability in sediments of Musa Creek. Twenty samples including 10 sediments and 10 water samples were collected from 10 stations. The concentration of heavy metals was measured using inductively coupled plasma–mass spectrometry. The bioavailability of heavy metals was carried out by first step of Tessier sequential extraction method so called single extraction in the manuscript. The results showed that the mean concentrations of cadmium, chromium, nickel, and zinc in water samples was 0.17, 7.44, 43.80, and 8.85 µg/L, respectively. …

Parametric Analyses Of The Dynamic Response And Fatigue Life Of A Compliant Vertical Access Riser With Internal Flow, Fuheng Li, Haiyan Guo, Xiaomin Li, Zhen Liu, Honglu Gu, Peng Cui

Journal of Marine Science and Technology–Taiwan

Subsea interventions on wellheads can be directly performed from a platform using a compliant vertical access riser (CVAR), thereby incurring tremendous economic benefits. The dynamic response and the fatigue damage of a CVAR subjected to regular waves are important for assessing its application feasibility. Therefore, a reasonable numerical model is presented for assessing the damage to a CVAR considering the internal flow using the slender rod theory. The finite element method combined with a Newmark-β scheme is employed to discretize the equations and update the time integration. The Palmgren-Miner rule is applied for fatigue life prediction, focusing on the wave-induced …

Using The Inverse Method To Investigate Flow Models For Mixed Convection Of Annular Finned Tube Heat Exchanger, Han-Taw Chen, Cheng-Hui You, Wei-Lun Hsu, Jiang-Ren Chang

Journal of Marine Science and Technology–Taiwan

The inverse heat conduction method (IHCM) and three-dimensional (3D) computational fluid dynamic (CFD) in combination with are used to select the appropriate flow model and near-wall treatment for the mixed convection of the annular finned tube heat exchanger. Thus, the numerical results obtained by all k-ε models and near-wall treatments are yielded for 4 m/s ≤ Va ≤ 5 m/s. First, IHCM combined with is applied to estimate and Q values. The obtained estimates of and Q are used as reference values for CFD. The results show that the Tk and results obtained by the standard (STD) k-ε model with …

Prediction Of Remaining Useful Life Of Wind Turbine Shaft Bearings Using Machine Learning, Jinsiang Shaw, Bingjie Wu

Journal of Marine Science and Technology–Taiwan

Wind turbines are a major trend in the current green energy market. Wind energy is abundant, and if utilized properly, can result in significant reductions in carbon emissions. Therefore, the development of wind power systems is urgently required. However, wind turbines are mainly built in unmanned areas. Regular inspections require substantial manpower and material resources, and doubts regarding the accuracy of the inspected data may occur. Therefore, it is necessary to establish an automatic diagnostic method for determining the remaining useful life (RUL) of a wind turbine to facilitate predictive maintenance. In this study, a multi-class support vector machine (SVM) …

Supply Chain Management For Superyacht Construction In Taiwan, Chun Pong Wong

Journal of Marine Science and Technology–Taiwan

In the last 20 years, Taiwan’s superyacht construction industry has performed well in the global market; however, numerous yacht builders still practice traditional supply chain management (SCM), which is often inefficient and regularly leads to time and cost overruns. This study aimed to answer two research questions: 1) what SCM challenges does the superyacht construction industry face? and 2) what SCM strategies and practices should be adopted to improve the efficacy and profitability of superyacht construction companies?

Numerous studies have explored supply chains (SCs) and SCM, but few have focused on the superyacht construction industry. Superyacht construction projects are highly …

The Impact Of Variable Curing Conditions On The Properties And Microstructures Of Mixtures Of Ground Granulated Blast Furnace Slag And Circulating Q1 Fluidized Bed Combustion Ash, Shao-Heng Hsieh, Wei-Chung Yeih, Ran Huang

Journal of Marine Science and Technology–Taiwan

The use of Portland cement, an important construction material, is encountering growing challenges because of its adverse environmental impacts. In this study, a new hydration system without Portland cement, which blends ground granulated blast furnace slag (GGBFS) and circulating fluidized bed combustion (CFBC) ash, was developed and tested. The variables included the effect of different types of CFBC ash, the mass ratios between CFBC ash and GGBFS, and different curing temperatures. The mortar properties were determined through the compressive strength test, absorption test, shrinkage test, X-ray diffraction, scanning electron microscopy, and energy-dispersive X-ray spectroscopy. The main results were as follows: …

A Marine Gis-Based Alert System To Prevent Vessels Collision With Offshore Platforms, Sheng-Long Kao, Ki-Yin Chang, Tai-Wen Hsu

Journal of Marine Science and Technology–Taiwan

How to effectively use an intelligent aid-to-navigation (AtoN) to prevent ships from accidentally colliding with offshore marine platforms (OMPs) is an important issue for maritime environment. This study proposes a novel collision alert system with three components to process Automatic Identification System (AIS) data in the cloud and broadcast AtoNs to ships in order to avoid collision with OMPs. The first component receives as input fuzzy linguistic variables (vessel size, relative speed, and approach distance) from the AIS data and calculates the radius of the guarding ring. The second component serves a Marine GIS (MGIS), where the system simulates the …

A Coral Spawning Calendar For Sesoko Station, Okinawa, Japan, Andrew H. Baird, Alasdair J. Edwards, James R. Guest, Saki Harii, Masayuki Hatta, Liam Lachs, Hanaka Mera, Frederic Sinniger, David Abrego, Or Ben-Zvi, Omri Bronstein, Patrick C. Cabaitan, Vivian R. Cumbo, Gal Eyal, Lee Eyal-Shaham, Bar Feldman, Joana Figueiredo, Jean-François Flot, Mila Grinblat, Andrew Heyward, Michio Hidaka, Mamiko Hirose, Akira Iguchi, Naoko Isomura, Robert A. Kinzie, Seiya Kitanobo, Alyson Kuba, Oren Levy, Yossi Loya, Takuma Mezaki, Amin R. Mohamed, Masaya Morita, Satoshi Nojima, Yoko Nozawa, Rian Prasetia, Eneour Puill-Stephan, Catalina Ramirez-Portilla, Hanna Rapuano, Yaeli Rosenberg, Yusuke Sakai, Kazuhiko Sakai, Tom Shlesinger, Tullia I. Terraneo, Irina Yakovleva, Hiromi H. Yamamoto, Kiyoshi Yamazato

Marine & Environmental Sciences Faculty Articles

Sesoko Station, Okinawa, has been the site of many significant advances in coral reproductive research and it continues to be a preferred destination for both Japanese and international researchers. Consequently, there are decades of spawning observations, which we present and explore here with the aim of making it easier to predict when species spawn at Sesoko Station. The data include over 700 spawning observations from 87 species of reef-building hermatypic corals. Almost all spawning occurred between dusk and dawn, with most spawning activity concentrated in the 2 to 4 hours after sunset. Some phylogenetic patterns were evident: most Acropora species …

Pelagic Tunicate Grazing On Marine Microbes Revealed By Integrative Approaches, Kelly R. Sutherland, Anne W. Thompson

Biology Faculty Publications and Presentations

Marine microorganisms comprise a large fraction of ocean carbon and are central players in global biogeochemical cycling. Significant gaps remain, however, in our understanding of processes that determine the fate, distribution, and community structure of microbial communities. Protists and viruses are accepted as being part of the microbial loop and a source of microbial mortality. However, pelagic tunicates (salps, doliolods, pyrosomes, and appendicularians), which are abundant in oceanic and coastal environments and consume microorganisms with higher individual grazing rates than other common grazers, remain underappreciated in their role controlling microbial communities, distributions, and flux through ecosystems. In spite of sampling …

Peruvian Fur Seals As Archivists Of El Niño Southern Oscillation Effects, Mickie Rae Edwards, Susana Cardenas-Alayza, Michael Adkesson, Mya Daniels-Abdulahad, Amy Hirons

Marine & Environmental Sciences Faculty Articles

Peru’s coastal waters are characterized by significant environmental fluctuation due to periodic El Niño- La Niña- Southern Oscillation (ENSO) events. This variability results in ecosystem-wide food web changes which are reflected in the tissues of the Peruvian fur seal (Arctocephalus australis). Stable isotope ratios (δ¹³C and δ¹⁵N) in Peruvian fur seal vibrissae (whiskers) are used to infer temporal primary production and dietary variations in individuals. Sea surface temperature anomaly (SSTA) recordings from the Niño 1+2 Index region captured corresponding ENSO conditions. Fluctuations in δ¹⁵N values were correlated to SSTA records, indicating that …

Resource Allocation And Phenotypic Plasticity Of Simultaneous Hermaphroditic Turtle Barnacles (Chelonibia Testudinaria), Kevin C. Cash

All HCAS Student Capstones, Theses, and Dissertations

This research addresses the knowledge gap of phenotypic plasticity in a commonly found and important species of epizoic barnacle, Chelonibia testudinaria. Limited research has been published regarding how phenotypic expression is mediated the spatial distribution of barnacles on a mobile host. To investigate this potential relationship, barnacles were collected from the backs of turtles along the beaches of Fort Lauderdale Florida. These barnacles were assessed for various phenotypic traits as well as their corresponding spatial distribution on the turtle carapace. Barnacles were safely removed from the carapace using a chisel before their preservation in ethanol. Barnacles were then numbered …

Prey Toxicity, Prey Refuge Use, And Stochasticity Increase Algal Blooms In A Phytoplankton-Zooplankton Mathematical Model, Naomi Cedeño, Erin Estes, Christopher Stieha

Annual Symposium on Biomathematics and Ecology Education and Research

No abstract provided.

Temporal Baseline Of Essential And Non-Essential Elements Recorded In Baleen Of Western Arctic Bowhead Whale (Balaena Mysticetus), Samantha Shore, Dimitri Giarikos, Lawrence K. Duffy, Mickie Rae Edwards, Amy Hirons

Marine & Environmental Sciences Faculty Articles

This study established the first baseline of changing elemental concentrations in bowhead whale baleen over time (1958–1999). From previously published stable isotope data, year, season (summer or winter), and location (Beaufort or Bering/Chukchi seas) were attributed to each sample. Thirteen elements (Al, Cd, Co, Cr, Cu, Fe, Hg, Mn, Ni, Pb, Se, V, Zn) in baleen from nine subsistence-harvested bowhead whales (n = 138) were detected. Al, Cu, and Fe were the highest concentrations while Cd and V were among the lowest. Our data supports absorption as the main route of exposure to environmental elements rather than biomagnification due to …

Who Are We? Highlighting Nuances In Asian American Experiences In Ecology And Evolutionary Biology, K. H. Nguyen, A. K. Akiona, C. C. Chang, V. B. Chaudhary, S. J. Cheng, S. M. Johnson, S. S. Kahanamoku, A. Lee, E. E. Deleon Sanchez, L. M. Segui, Richelle L. Tanner

Biology, Chemistry, and Environmental Sciences Faculty Articles and Research

"As ecologists and evolutionary biologists, it is important to recognize that issues surrounding AAPI do not exist in a vacuum and that these issues affect some of our students and colleagues in and outside of the classroom and lab. Recently, anti-Asian racism has been brought to the forefront of US society. Though media attention has primarily focused on how COVID-19 fears motivated both violent and nonviolent racist incidents against East Asians in 2020 (e.g., Chinese and Taiwanese Americans) (Tessler et al. 2020), attacks on other AAPI groups, such as South and Southeast Asians (e.g., Filipino, Pakistani, and Vietnamese Americans) (Truong …

First Ecological Characterization Of Whip Black Coral Assemblages (Hexacorallia: Antipatharia) In The Easter Island Ecoregion, Southeastern Pacific, Jan M. Tapia-Guerra, Cynthia M. Asorey, Erin E. Easton, Daniel Wagner, Matthias Gorny, Javier Sellanes

School of Earth, Environmental, & Marine Sciences Faculty Publications

One of the main priorities of marine scientific research is to provide practical information and guidance for biodiversity conservation and management. In this context, the identification of key and fragile habitat-forming species is particularly important. Despite their ecological importance, whip corals in the order Antipatharia remain poorly studied around the world, and especially in the South Pacific. During recent expeditions to explore mesophotic and deep benthic habitats around Rapa Nui (Easter Island), dense assemblages of whip black corals were observed in situ. We here report the first detailed description of the distribution and abundance of these black coral assemblages …

Ontogeny And Resource Use Of Invasive Lionfish, Pterois Volitans, In Shallow Water Habitats Surrounding Summerland Key, Emily Shallow

LSU Master's Theses

The invasive red lionfish, Pterois volitans, has been shown to considerably reduce reef biomass via predation and competition in non-native ranges. Primary maintenance of lionfish is through human removal (e.g., culling), which does not sufficiently control the population due to deep-water spawning, recolonization of recently culled reefs, and ontogenetic migrations to mesophotic waters. Here, I investigated the diets and primary energy pathways during ontogeny in lionfish from the Florida Keys using stomach contents and stable isotope (δ¹³C and δ¹⁵N) analysis to better understand their role in the trophic food web. Additionally, isotopic overlap between lionfish …

The Effects Of Partial Replacement Of Fish-Meal With Mesquite Pods On Growth Parameters Of Tilapia Fry, Oreochromis Mossambicus, Mohammed Abdullah Al-Dohai, Balqees Saleh Bin Breek, Mazan Saleh Bamatraf

Hadhramout University Journal of Natural & Applied Sciences

This study was carried out to determine the effect of partial replacement of fish-meal with 15%, 30% and 45% of Mesquite pods (MP) in testing diets on growth parameters of Tilapia fries, Oreochromis mossambicus which reared in glass aquarium for 12 weeks. Twenty O. mossambicus fries (Mean initial weight 3.33 g) were randomly distributed into each 12 glass aquariums (measuring 50 x 50 x 50 cm) filled with tap water to 40 cm depth. Four treatments were formulated in this experiment. The first treatment was controlled and three testing diets contain 15%, 30% and 45% of MP. The results showed …

Proximate And Fatty Acid Composition Of Three Tuna Species From Hadhramout Coast Of The Arabian Sea, Yemen, Osan Maroof Bahurmiz

Hadhramout University Journal of Natural & Applied Sciences

The Proximate and fatty acid composition were evaluated in three species of tuna; Yellowfin tuna (Thunnus albacares), longtail tuna (T. tonggol), little tuna (Euthynnus affinis) caught from Hadhramout coast of the Arabian Sea. The results of proximate composition showed high protein content in the flesh of all species, ranged from 22.52% to 24.36%. The average moisture, lipid and ash contents in the flesh of the three fish species were in the range of 70.13–74.0%, 2.34–4.66% and 1.25–1.37%, respectively. The fatty acid profile of all fish was dominated by saturated fatty acids, (31.76–36.77%), followed by polyunsaturated fatty acids (31.21–31.59%) and then …

Cetacean Acousticwelfare In Wild And Managed-Care Settings: Gaps And Opportunities, Paige E. Stevens, Heather Hill, Jason N. Bruck

Faculty Publications

Cetaceans are potentially at risk of poor welfare due to the animals’ natural reliance on sound and the persistent nature of anthropogenic noise, especially in the wild. Industrial, commercial, and recreational human activity has expanded across the seas, resulting in a propagation of sound with varying frequency characteristics. In many countries, current regulations are based on the potential to induce hearing loss; however, a more nuanced approach is needed when shaping regulations, due to other non-hearing loss effects including activation of the stress response, acoustic masking, frequency shifts, alterations in behavior, and decreased foraging. Cetaceans in managedcare settings share the …

Wrack Lines Volume 21, Number 2, Fall/Winter 2021-22, Nancy C. Balcom, Judy Benson, Syma A. Ebbin, Kira Goldenberg, Judy Preston, Howard "Mickey" Weiss

Wrack Lines

"Discovery, Rediscovery and Rebirth: new eyes, new understanding of familiar places" is the theme for the Fall-Winter 2021-22 issue. The main article package consists of five stories about the lands and waters that will comprise the Connecticut National Estuarine Research Reserve (CT NERR). These are: overview and introduction; Great Island; lower Thames River; Bluff Point State Park; and Haley Farm State Park. Other articles include one on research into the cause of invasive Cladophora seaweed dominating Little Narragansett Bay; and another on the transformation of the Yale Peabody Museum of Natural History.

Nutrient Cycling In Tropical And Temperate Coastal Waters: Is Latitude Making A Difference?, Christian Lønborg, Moritz Müller, Edward C. V. Butler, Shan Jiang, Seng Keat Ooi, Dieu Huong Trinh, Pui Yee Wong, Suryati M. Ali, Chun Cui, Wee Boon Siong, Erik S. Yando, Daniel A. Friess, Judith A. Rosentreter, Bradley D. Eyre, Patrick Martin

Biological Sciences Faculty Publications

Tropical coastal waters are highly dynamic and amongst the most biogeochemically active zones in the ocean. This review compares nitrogen (N) and phosphorus (P) cycles in temperate and tropical coastal waters. We review the literature to identify major similarities and differences between these two regions, specifically with regards to the impact of environmental factors (temperature, sunlight), riverine inputs, groundwater, lateral fluxes, atmospheric deposition, nitrogen fixation, organic nutrient cycling, primary production, respiration, sedimentary burial, denitrification and anammox. Overall, there are some similarities but also key differences in nutrient cycling, with differences relating mainly to temperature, sunlight, and precipitation amounts and patterns. …

Overfishing Drives Over One-Third Of All Sharks And Rays Toward A Global Extinction Crisis, Nicholas K. Dulvy, Nathan Pacoureau, Cassandra L. Rigby, Riley A. Pollom, Rima W. Jabado, David A. Ebert, Brittany Finucci, Caroline M. Pollock, Jessica Cheok, Danielle H. Derrick, Katelyn B. Herman, C. Samantha Sherman, Wade J. Vanderwright, Julia M. Lawson, Rachel H.L. Walls, John K. Carlson, Patricia Charvet, Kinattumkara K. Bineesh, Daniel Fernando, Gina M. Ralph, Jay H. Matsushiba, Craig Hilton-Taylor, Sonja V. Fordham, Colin A. Simpfendorfer

Biological Sciences Faculty Publications

The scale and drivers of marine biodiversity loss are being revealed by the International Union for Conservation of Nature (IUCN) Red List assessment process. We present the first global reassessment of 1,199 species in Class Chondrichthyes-sharks, rays, and chimeras. The first global assessment (in 2014) concluded that one-quarter (24%) of species were threatened. Now, 391 (32.6%) species are threatened with extinction. When this percentage of threat is applied to Data Deficient species, more than one-third (37.5%) of chondrichthyans are estimated to be threatened, with much of this change resulting from new information. Three species are Critically Endangered (Possibly Extinct), representing …