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Articles 1 - 29 of 29
Full-Text Articles in Forest Sciences
The Biltmore Forest School And The Establishment Of Forestry Education In America, Dan Barry Croom
The Biltmore Forest School And The Establishment Of Forestry Education In America, Dan Barry Croom
Journal of Research in Technical Careers
The Biltmore Forest School, despite its unusual existence within the affluent Biltmore Estate, played a crucial role in the early 20th-century American forestry movement. Founded by Carl A. Schenck and supported by George Vanderbilt II, the school aimed to educate foresters and promote sustainable forest management. However, many aspects of the Biltmore experiment failed due to the new and untested nature of forestry science in America. This experiment exposed a fundamental divide in forestry education, with Gifford Pinchot advocating for conservation-centered teaching while Schenck believed in the economic viability of lumber production. Ultimately, the Biltmore Forest School offered valuable vocational …
Palm Oil And Exported Deforestation, Anja Marcusiu, Alexis Osorio
Palm Oil And Exported Deforestation, Anja Marcusiu, Alexis Osorio
Undergraduate Research Symposium Posters
Palm oil is used in our everyday lives through: Beautify/hygiene products; Bioenergy; Food. Indonesia and Malaysia account for 85% of the world's palm oil production, making them the largest exporters. Because palm oil is a highly productive crop and is in high demand, how do companies integrate sustainability to limit deforestation and preserve biodiversity?
The Aboveground And Belowground Growth Characteristics Of Juvenile Conifers In The Southwestern United States, N. L. Pirtel, R. M. Hubbard, J. B. Bradford, T. E. Kolb, M. E. Litvak, Scott Abella, S. L. Porter, Matthew Petrie
The Aboveground And Belowground Growth Characteristics Of Juvenile Conifers In The Southwestern United States, N. L. Pirtel, R. M. Hubbard, J. B. Bradford, T. E. Kolb, M. E. Litvak, Scott Abella, S. L. Porter, Matthew Petrie
Life Sciences Faculty Research
Juvenile tree survival will play an important role in the persistence of coniferous forests and woodlands in the southwestern United States (SWUS). Vulnerability to climatic and environmental stress declines as trees grow, such that larger, more deeply rooted juveniles are less likely to experience mortality. It is unclear how juvenile conifers partition the aboveground and belowground components of early growth, if growth differs between species and ecosystem types, and what environmental factors influence juvenile carbon allocation above- or belowground. We developed a novel data set for four juvenile conifer groups (junipers, piñon pines, ponderosa pines, firs; 1121 juveniles sampled, 221 …
Hawai‘I Forest Review: Synthesizing The Ecology, Evolution, And Conservation Of A Model System, Kasey E. Barton, Andrea Westerband, Rebecca Ostertag, Elizabeth Stacy, Kawika Winter, Donald R. Drake, Lucas Berio Fortini, Creighton M. Litton, Susan Cordell, Paul Krushelnycky, Kapua Kawelo, Kealoha Feliciano, Gordon Bennett, Tiffany Knight
Hawai‘I Forest Review: Synthesizing The Ecology, Evolution, And Conservation Of A Model System, Kasey E. Barton, Andrea Westerband, Rebecca Ostertag, Elizabeth Stacy, Kawika Winter, Donald R. Drake, Lucas Berio Fortini, Creighton M. Litton, Susan Cordell, Paul Krushelnycky, Kapua Kawelo, Kealoha Feliciano, Gordon Bennett, Tiffany Knight
Life Sciences Faculty Research
As the most remote archipelago in the world, the Hawaiian Islands are home to a highly endemic and disharmonic biota that has fascinated biologists for centuries. Forests are the dominant terrestrial biome in Hawai‘i, spanning complex, heterogeneous climates across substrates that vary tremendously in age, soil structure, and nutrient availability. Species richness is low in Hawaiian forests compared to other tropical forests, as a consequence of dispersal limitation from continents and adaptive radiations in only some lineages, and forests are dominated by the widespread Metrosideros species complex. Low species richness provides a relatively tractable model system for studies of community …
Non-Native Spiders Change Assemblages Of Hawaiian Forest Fragment Kipuka Over Space And Time, Julien Pétillon, Kaїna Privet, George K. Roderick, Rosemary G. Gullespie, Don K. Price
Non-Native Spiders Change Assemblages Of Hawaiian Forest Fragment Kipuka Over Space And Time, Julien Pétillon, Kaїna Privet, George K. Roderick, Rosemary G. Gullespie, Don K. Price
Life Sciences Faculty Research
We assessed how assemblages of spiders were structured in small Hawaiian tropical forest fragments (Hawaiian, kipuka) within a matrix of previous lava flows, over both space (sampling kipuka of different sizes) and time (comparison with a similar study from 1998). Standardized hand-collection by night was carried out in May 2016. In total, 702 spiders were collected, representing 6 families and 25 (morpho-)species. We found that the number of individuals, but not species richness, was highly correlated with the area of sampled forest fragments, suggesting that kipuka act as separate habitat islands for these predatory arthropods. Species richness was significantly lower …
Varieties Of The Highly Dispersible And Hypervariable Tree, Metrosideros Polymorpha, Differ In Response To Mechanical Stress And Light Across A Sharp Ecotone, Jill M. Ekar, Donald K. Price, Melissa A. Johnson, Elizabeth A. Stacy
Varieties Of The Highly Dispersible And Hypervariable Tree, Metrosideros Polymorpha, Differ In Response To Mechanical Stress And Light Across A Sharp Ecotone, Jill M. Ekar, Donald K. Price, Melissa A. Johnson, Elizabeth A. Stacy
Life Sciences Faculty Research
Premise: The drivers of isolation between sympatric populations of long‐lived and highly dispersible conspecific plants are not well understood. In the Hawaiian Islands, the landscape‐dominant tree, Metrosideros polymorpha, displays extraordinary phenotypic differences among sympatric varieties despite high dispersibility of its pollen and seeds, thereby presenting a unique opportunity to investigate how disruptive selection alone can maintain incipient forms. Stenophyllous M. polymorpha var. newellii is a recently evolved tree endemic to the waterways of eastern Hawai'i Island that shows striking neutral genetic differentiation from its ancestor, wet‐forest M. polymorpha var. glaberrima, despite sympatry of these forms. We looked for evidence for, …
Biological Effects On Serpentinite Weathering, Mary H. Evert, Julie Baumeister, Elisabeth Hausrath
Biological Effects On Serpentinite Weathering, Mary H. Evert, Julie Baumeister, Elisabeth Hausrath
Undergraduate Research Opportunities Program (UROP)
Serpentinites, perhaps more than any other rock type, control the composition and evolution of the development of the surrounding ecosystems. The bulk chemistry of serpentinite rocks, high in Mg and trace elements, and low in nutrients such as Ca, K, P, and N, causes an extreme and stressful environment for ecosystems. However, the role that those serpentine ecosystems play in development of serpentine soils has not been examined.
Due to the unusual chemistry of serpentine soils, serpentine ecosystems have deeper and better-developed root systems than other ecosystems. The rhizosphere of serpentine systems, documented to produce abundant organic acids and siderophores, …
How Well Do U.S. Forest Service Terrestrial Ecosystem Surveys Correspond With Measured Vegetation Properties?, Scott R. Abella
How Well Do U.S. Forest Service Terrestrial Ecosystem Surveys Correspond With Measured Vegetation Properties?, Scott R. Abella
Life Sciences Faculty Research
No abstract provided.
Characterization Of Ips Pini Ipsdienol Dehydrogenase (Idol Dh), Heidi Pak, Claus Tittiger, William Welch
Characterization Of Ips Pini Ipsdienol Dehydrogenase (Idol Dh), Heidi Pak, Claus Tittiger, William Welch
Festival of Communities: UG Symposium (Posters)
Ipsdienol is an important pheromone component for pine engraver beetle, Ips pini. Ipsdienol is a ten carbon monoterpenoid secondary alcohol and ipsdienone is the corresponding ketone. We are characterizing the activity of recombinant IDOL DH produced in Sf9 (insect) cells. The enzyme has a high stereospecificity: (-) ipsdienol was found to be a substrate while (+)-ipsdienol was neither a substrate nor inhibitor. Closely related monoterpenoids, such as nerol, geraniol, and citral, were neither substrates nor inhibitors. Smaller compounds, such as 2-propanol, also failed to act as an inhibitor or substrate. This indicates the binding site of this enzyme is highly …
Research Poster: Eco-Hydrological Pathways Inferred From Stable Isotopes In A Pinus Ponderosa And Pinus Monophylla Woodland Of The Sheep Range, Southern Great Basin, Usa, Kelli Hoover, Simon Poulson, Franco Biondi, Jeffrey Underwood
Research Poster: Eco-Hydrological Pathways Inferred From Stable Isotopes In A Pinus Ponderosa And Pinus Monophylla Woodland Of The Sheep Range, Southern Great Basin, Usa, Kelli Hoover, Simon Poulson, Franco Biondi, Jeffrey Underwood
2010 Annual Nevada NSF EPSCoR Climate Change Conference
Research poster
Research Poster: Tree Population Dynamics At Ash Meadows National Wildlife Refuge: Influences Of Environmental Stress And Disturbance, Sarah L. Karam, Peter J. Weisberg, Stephanie O. Sunderman
Research Poster: Tree Population Dynamics At Ash Meadows National Wildlife Refuge: Influences Of Environmental Stress And Disturbance, Sarah L. Karam, Peter J. Weisberg, Stephanie O. Sunderman
2010 Annual Nevada NSF EPSCoR Climate Change Conference
Research poster
Mojave Applied Ecology Notes Fall 2009, Tom Monaco, Scott R. Abella, Amber Lee, James E. Deacon
Mojave Applied Ecology Notes Fall 2009, Tom Monaco, Scott R. Abella, Amber Lee, James E. Deacon
Mojave Applied Ecology Notes
USDA working to manage invasive annual grasses, effects of heat and smoke on red brome soil seed bank, how burial depth and substrate affect germination of Sahara mustard and red brome, environmental effects of the southern Nevada groundwater project
Tree Canopy Types Constrain Plant Distributions In Ponderosa Pine- Gambel Oak Forests, Northern Arizona, Scott R. Abella
Tree Canopy Types Constrain Plant Distributions In Ponderosa Pine- Gambel Oak Forests, Northern Arizona, Scott R. Abella
Public Policy and Leadership Faculty Publications
Trees in many forests affect the soils and plants below their canopies. In current high-density southwestern ponderosa pine (Pinus ponderosa) forests, managers have opportunities to enhance multiple ecosystem values by manipulating tree density, distribution, and canopy cover through tree thinning. I performed a study in northern Arizona ponderosa pine-Gambel oak (Quercus gambelii) forests to measure the influences of tree canopy types on understory plant communities and soil properties. On ten 2.5-acre (1-ha) sites, I sampled five 43-ft2 (4-m2) plots below each of the following five canopy types: openings; single ponderosa pine; and Gambel oak single stems, dispersed clumps, and thickets. …
Managing Gambel Oak In Southwestern Ponderosa Pine Forests: The Status Of Our Knowledge, Scott R. Abella
Managing Gambel Oak In Southwestern Ponderosa Pine Forests: The Status Of Our Knowledge, Scott R. Abella
Public Policy and Leadership Faculty Publications
Gambel oak (Quercus gambelii) is a key deciduous species in southwestern ponderosa pine (Pinus ponderosa) forests and is important for wildlife habitat, soil processes, and human values. This report (1) summarizes Gambel oak’s biological characteristics and importance in ponderosa pine forests, (2) synthesizes literature on changes in tree densities and fire frequencies since Euro-American settlement in pine-oak forests, (3) suggests management prescriptions for accomplishing various oak management objectives (for example, increasing diameter growth or acorn production), and (4) provides an appendix containing 203 Gambel oak literature citations organized by subject. Nine studies that reconstructed Gambel oak density changes since settlement …
Estimating Soil Seed Bank Characteristics In Ponderosa Pine Forests Using Vegetation And Forest-Floor Data, Scott R. Abella, Judith D. Springer
Estimating Soil Seed Bank Characteristics In Ponderosa Pine Forests Using Vegetation And Forest-Floor Data, Scott R. Abella, Judith D. Springer
Public Policy and Leadership Faculty Publications
Soil seed banks are important for vegetation management because they contain propagules of species that may be considered desirable or undesirable for site colonization after management and disturbance events. Knowledge of seed bank size and composition before planning management activities facilitates proactive management by providing early alerts of exotic species presence and of abilities of seed banks to promote colonization by desirable species. We developed models in ponderosa pine (Pinus ponderosa) forests in northern Arizona to estimate the size and richness of mineral soil seed banks using readily observable vegetation and forestfloor characteristics. Regression models using three or fewer predictors …
Gambel Oak Growth Forms: Management Opportunities For Increasing Ecosystem Diversity, Scott R. Abella
Gambel Oak Growth Forms: Management Opportunities For Increasing Ecosystem Diversity, Scott R. Abella
Public Policy and Leadership Faculty Publications
Gambel oak (Quercus gambelii) clones have several different growth forms in southwestern ponderosa pine (Pinus ponderosa) forests, and these growth forms each provide unique wildlife habitat and resource values. The purposes of this note are to review published growth-form classifications for Gambel oak, provide examples of ecological effects of different growth forms, and summarize management strategies for promoting desired growth forms. Four different growth-form classifications have been published, which generally recognize variants of three basic forms: shrubby thickets of small-diameter stems, pole-sized clumps, and large trees. These growth forms exemplify ecological and management tradeoffs. For example, shrubby forms provide browse …
Changes In Gambel Oak Densities In Southwestern Ponderosa Pine Forests Since Euro-American Settlement, Scott R. Abella, Peter Z. Fule
Changes In Gambel Oak Densities In Southwestern Ponderosa Pine Forests Since Euro-American Settlement, Scott R. Abella, Peter Z. Fule
Public Policy and Leadership Faculty Publications
Densities of small-diameter ponderosa pine (Pinus ponderosa) trees have increased in southwestern ponderosa pine forests during a period of fire exclusion since Euro-American settlement in the late 1800s. However, less well known are potential changes in Gambel oak (Quercus gambelii) densities during this period in these forests. We reviewed published literature to summarize changes in oak density in ponderosa pine forests over the past 140 years and evaluated the possibility that large-diameter oaks have decreased in density. All nine studies examining oak density changes found that densities of small-diameter oaks have escalated. Increases ranged from 4- to more than 63-fold. …
Woody Biomass Energy Solutions, Scott Bell
Woody Biomass Energy Solutions, Scott Bell
UNLV Renewable Energy Symposium
The 2008 UNLV Renewable Energy Symposium was presented by the Office of Strategic Energy Programs and co-sponsored by the Division of Research and Graduate Studies on August 20, 2008 on the UNLV campus.
The event focused on renewable energy production in Nevada, the US Southwest, and renewable research projects nationwide. It was a great opportunity for anyone working on renewable projects to collaborate with others in this field and exchange information. Over 230 individuals attended the event this year.
Plant Recruitment In A Northern Arizona Ponderosa Pine Forest: Testing Seed- And Leaf Litter- Limitation Hypotheses, Scott R. Abella
Plant Recruitment In A Northern Arizona Ponderosa Pine Forest: Testing Seed- And Leaf Litter- Limitation Hypotheses, Scott R. Abella
Public Policy and Leadership Faculty Publications
Seed availability and leaf litter limit plant establishment in some ecosystems. To evaluate the hypothesis that these factors limit understory plant recruitment in Pinus ponderosa forests, I conducted a seeding and litter removal experiment at six thinned sites in the Fort Valley Experimental Forest, northern Arizona. Experimental seeding of four native species (Penstemon virgatus, Erigeron formosissimus, Elymus elymoides, and Festuca arizonica) and raking of litter occurred in 2005. Seeding resulted in a substantial recruitment of 14 to 103 seedlings/m2 (1 to 10/ft2) one month after seeding for two species (P. virgatus and E. elymoides), but these densities subsequently declined by …
Fire Effects On Gambel Oak In Southwestern Ponderosa Pine-Oak Forests, Scott R. Abella, Peter Z. Fule
Fire Effects On Gambel Oak In Southwestern Ponderosa Pine-Oak Forests, Scott R. Abella, Peter Z. Fule
Public Policy and Leadership Faculty Publications
Gambel oak (Quercus gambelii) is ecologically and aesthetically valuable in southwestern ponderosa pine (Pinus ponderosa) forests. Fire effects on Gambel oak are important because fire may be used in pine-oak forests to manage oak directly or to accomplish other management objectives. We used published literature to: (1) ascertain historical fire regimes in pine-oak forests, (2) discern prescribed burning effects on Gambel oak survival and diameter growth, and (3) provide suggestions for using fire to manage oak. Frequent fire is part of Gambel oak’s historical environment, as historical fire return intervals often averaged less than 10 years in pine-oak forests. More …
Fire History And Forest Structural Change In The Spring Mountains, Scott R. Abella
Fire History And Forest Structural Change In The Spring Mountains, Scott R. Abella
Public Policy and Leadership Faculty Publications
Since early 2006 we have been working to develop a partnership with the Spring Mountains District of the Humboldt-Toiyabe National Forest to provide science support for understanding fire history and forest structural changes in support of ecologically based management strategies. We teamed up with the Ecological Restoration Institute (ERI) at Northern Arizona University and the University of Arizona Tree Ring Lab to deliver a workshop on March 6, 2008 at the interagency office in Las Vegas, Nevada. On September 16-18, we again teamed up with colleagues at ERI to conduct a preliminary field assessment of forest change at 10 sites …
A Unique Old-Growth Ponderosa Pine Forest In Northern Arizona, Scott R. Abella
A Unique Old-Growth Ponderosa Pine Forest In Northern Arizona, Scott R. Abella
Public Policy and Leadership Faculty Publications
Old-growth ponderosa pine (Pinus ponderosa) forests are uncommon in the Southwest, and only one oldgrowth forest (the Gus Pearson Natural Area [GPNA]) has been researched in the ponderosa pine belt surrounding the city of Flagstaff in northern Arizona. The purpose of this study was to measure soil characteristics, current and pre-Euro-American settlement (1885) tree structure, and understory plant composition in a 6-ha remnant old-growth forest on volcanic, red cinder soils. Soil bulk density was extremely low (0.21 Mg/m3) in this forest because of high volumetric contents of cinders >2 mm diameter. As a result, volumetric soil moisture, organic C, and …
Species Richness And Soil Properties In Pinus Ponderosa Forests: A Structural Equation Modeling Analysis, Daniel C. Laughlin, Scott R. Abella, W. Wallace Covington, James B. Grace
Species Richness And Soil Properties In Pinus Ponderosa Forests: A Structural Equation Modeling Analysis, Daniel C. Laughlin, Scott R. Abella, W. Wallace Covington, James B. Grace
Public Policy and Leadership Faculty Publications
Question: How are the effects of mineral soil properties on understory plant species richness propagated through a network of processes involving the forest overstory, soil organic matter, soil nitrogen, and understory plant abundance? Location: North-central Arizona, USA. Methods: We sampled 75 0.05-ha plots across a broad soil gradient in a Pinus ponderosa (ponderosa pine) forest ecosystem. We evaluated multivariate models of plant species richness using structural equation modeling. Results: Richness was highest at intermediate levels of understory plant cover, suggesting that both colonization success and competitive exclusion can limit richness in this system. We did not detect a reciprocal positive …
Past, Present, And Future Old Growth In Frequent-Fire Conifer Forests Of The Western United States, Scott R. Abella, W. Wallace Covington, Peter Z. Fule, Leigh B. Lentile, Andrew J. Sanchez Meador, Penelope Morgan
Past, Present, And Future Old Growth In Frequent-Fire Conifer Forests Of The Western United States, Scott R. Abella, W. Wallace Covington, Peter Z. Fule, Leigh B. Lentile, Andrew J. Sanchez Meador, Penelope Morgan
Public Policy and Leadership Faculty Publications
Old growth in the frequent-fire conifer forests of the western United States, such as those containing ponderosa pine (Pinus ponderosa), Jeffrey pine (P. jeffreyi), giant sequoia (Sequioa giganteum) and other species, has undergone major changes since Euro-American settlement. Understanding past changes and anticipating future changes under different potential management scenarios are fundamental to developing ecologically based fuel reduction or ecological restoration treatments. Some of the many changes that have occurred in these forests include shifts from historically frequent surface fire to no fire or to stand-replacing fire regimes, increases in tree density, increased abundance of fire-intolerant trees, decreases in understory …
Forest-Floor Treatments In Arizona Ponderosa Pine Restoration Ecosystems: No Short-Term Effects On Plant Communities, Scott R. Abella, W. Wallace Covington
Forest-Floor Treatments In Arizona Ponderosa Pine Restoration Ecosystems: No Short-Term Effects On Plant Communities, Scott R. Abella, W. Wallace Covington
Public Policy and Leadership Faculty Publications
Leaf litter accumulation during fire exclusion and increases in tree density in postsettlement southwestern Pinus ponderosa forests may limit the establishment of understory vegetation. We performed an experiment in P. ponderosa forests of northern Arizona to ascertain plant community responses to forest-floor scarification and Oi removal on thirty-six 100-m2 plots overlaid on an existing ecological restoration experiment that involved tree thinning and prescribed burning. Constrasting with findings from many other forest types, forest-floor treatments had no effect on community diversity or composition during the 2-year experiment. Sørensen similarities were as high as 97% between posttreatment years within treatments; and successional …
Canopy-Tree Influences Along A Soil Parent Material Gradient In Pinus-Ponderosa-Quercus Gambelii Forests, Northern Arizona, Scott R. Abella, Judith D. Springer
Canopy-Tree Influences Along A Soil Parent Material Gradient In Pinus-Ponderosa-Quercus Gambelii Forests, Northern Arizona, Scott R. Abella, Judith D. Springer
Public Policy and Leadership Faculty Publications
ABELLA, S. R. (Public Lands Institute and School of Life Sciences, University of Nevada, Las Vegas, Las Vegas, NV 89154-2040) AND J. D. SPRINGER (Ecological Restoration Institute, Northern Arizona University, Flagstaff, AZ 86011-5017). Canopy-tree influences along a soil parent material gradient in Pinus ponderosa- Quercus gambelii forests, northern Arizona. J. Torrey Bot. Soc. 135: 26–36. 2008.—The distribution of canopy trees can impose within-site patterns of soil properties and understory plant composition. At ten sites spanning a soil parent material gradient in northern Arizona Pinus ponderosa-Quercus gambelii forests, we compared soils and plant composition among five canopy types: openings, Pinus ponderosa …
Tree Thinning And Prescribed Burning Effects On Ground Flora In Arizona Ponderosa Pine Forests: A Review, Scott R. Abella
Tree Thinning And Prescribed Burning Effects On Ground Flora In Arizona Ponderosa Pine Forests: A Review, Scott R. Abella
Public Policy and Leadership Faculty Publications
Ground flora is an important response variable to monitor after tree thinning and prescribed burning treatments designed to restore Arizona ponderosa pine (Pinus ponderosa P.& C. Lawson) forests. This paper reviews published literature on the effects of thinning and burning on ground flora in Arizona ponderosa pine forests in five main categories of research: ground flora biomass, species diversity, plant community composition, population processes, and individual species ecology. Research published to date suggests that thinning and burning generally increase ground flora biomass, whereas other categories of research such as community composition and population processes have been little studied in Arizona …
Quantifying Ecosystem Geomorphology Of The Southern Appalachian Mountains, Scott R. Abella
Quantifying Ecosystem Geomorphology Of The Southern Appalachian Mountains, Scott R. Abella
Public Policy and Leadership Faculty Publications
Geomorphology is a dominant factor influencing vegetation distribution in the southern Appalachians, and quantifying landform characteristics is increasingly important for forest ecosystem classification. This study used slope gradient and two previously published geomorphic indices, terrain shape index and landform index that quantify landform shape and protection, to develop a field-based landform quantification system at four study areas in the southern Appalachians. Six major landform types (ridgetops, nose slopes, linear hillslopes, coves, stream ravines, and stream bottoms) exhibited quantitatively different characteristics, and these differences among landforms were not evident when using only categorical landform descriptions (e.g., convex, concave) that have been …
Spatial And Temporal Patterns Of Eastern White Pine Regeneration In The Northwestern Ohio Oak Stand, Scott R. Abella, Neil W. Macdonald
Spatial And Temporal Patterns Of Eastern White Pine Regeneration In The Northwestern Ohio Oak Stand, Scott R. Abella, Neil W. Macdonald
Public Policy and Leadership Faculty Publications
Eastern white pine (Pinus strobus L.) was often associated with oaks (Quercus spp.) on upland sites in presettlement forests of the upper Great Lakes region, but widespread logging and subsequent fires in the late 1800s converted these upland sites to fire-tolerant oak forests. Although white pine regeneration is occurring in these second-growth oak forests, white pine regeneration patterns in oak forests of the Great Lakes region are not well documented. We examined white pine regeneration in the southern Great Lakes region in an oak stand within the Oak Openings region of northwestern Ohio, where white pine plantations established in the …