Forest Sciences Commons^™

Effects Of Nitrogen On Temporal And Spatial Patterns Of Nitrate In Streams And Soil Solution Of A Central Hardwood Forest, Frank S. Gilliam, Mary Beth Adams

Biological Sciences Faculty Research

This study examined changes in stream and soil water NO3−and their relationship to temporal and spatial patterns of NO3−in soil solution of watersheds at the Fernow Experimental Forest, West Virginia. Following tenfold increases in stream NO3−concentrations over a 13-year period (1969–1981) on untreated WS4, concentrations have declined through 2006. Followingfourfold increases in stream NO3−on treatment WS3 from pretreatment levels to a 1998 maximum, concentrations have declinedthrough 2006, despite additions of N. Concentrations of soil water NO3−were consistently lower for WS4 compared to WS3.Data for soil water NO3−on WS3 versus WS4 followed patterns of net mineralization and nitrification for these watersheds.Nitrogen …

Nitrogen Addition Shapes Soil Phosphorus Availability In Two Reforested Tropical Forests In Southern China, Xiankai Lu, Jiangming Mo, Frank S. Gilliam, Hua Fang, Feifei Zhu, Yunting Fang, Wei Zhang, Juan Huang

Biological Sciences Faculty Research

Scant information is available on how soil phosphorus (P) availability responds to atmospheric nitrogen (N) deposition, especially in the tropical zones. This study examined the effect of N addition on soil P availability, and compared this effect between forest sites of contrasting land-use history. Effects of N addition on soil properties, litterfall production, P release from decomposing litter, and soil P availability were studied in a disturbed (reforested pine forest with previous understory vegetation and litter harvesting) and a rehabilitated (reforested mixed pine/broadleaf forest with no understory vegetation and litter harvesting) tropical forest in southern China. Experimental N-treatments (above ambient) …

0784: L. D. Fowler Papers, 1900-1920, Marshall University Special Collections

Guides to Manuscript Collections

The papers in this collection are both the professional and personal records of L.D. Fowler from 1900-1920 while working out of Durbin, WV. As an employee of the Pocahontas Tanning Company, L.D. Fowler’s business papers relate primarily to timber and the acquisition of bark associated with the tanning process. Among the type of materials included in the collection are correspondences (i.e. letters, telegraphs, and postcards), well-documented business transactions with a wide range of business partners, business cards, diaries, maps, popular culture items (calendars, etc.), and materials related to legal issues and political affiliations. This collection is divided into two overarching …

Effects Of Experimental Nitrogen Additions On Plant Diversity In An Old-Growth Tropical Forest, Xiankai Lu, Jiangming Mo, Frank S. Gilliam, Guoyi Zhou, Yunting Fang

Biological Sciences Faculty Research

Response of plant biodiversity to increased availability of nitrogen (N) has been investigated in temperate and boreal forests, which are typically N-limited, but little is known in tropical forests. We examined the effects of artificial N additions on plant diversity (species richness, density and cover) of the understory layer in an N saturated old-growth tropical forest in southern China to test the following hypothesis: N additions decrease plant diversity in N saturated tropical forests primarily from N-mediated changes in soil properties. Experimental additions of N were administered at the following levels from July 2003 to July 2008: no addition (Control); …

Effects Of Experimental Freezing On Soil Nitrogen Dynamics In Soils From A Net Nitrification Gradient In A Nitrogen-Saturated Hardwood Forest Ecosystem, Frank S. Gilliam, Adam Cook, Salina Lyter

Biological Sciences Faculty Research

This study examined effects of soil freezing on N dynamics in soil along an N processing gradient within a mixed hardwood dominated watershed at Fernow Experimental Forest, West Virginia. Sites were designated as LN (low rates of N processing), ML (moderately low), MH (moderately high), and HN (high). Soils underwent three 7-day freezing treatments (0, –20, or –80 °C) in the laboratory. Responses varied between temperature treatments and along the gradient. Initial effects differed among freezing treatments for net N mineralization, but not nitrification, in soils across the gradient, generally maintained at LN < ML ≤ MH < HN for all treatments. Net N mineralization potential was higher following freezing at –20 and –80 °C than control; all were higher than at 0 °C. Net nitrification potential exhibited similar patterns. LN was an exception, with net nitrification low regardless of treatment. Freezing response of N mineralization differed greatly from that of nitrification, suggesting that soil freezing may decouple two processes of the soil N cycle that are otherwise tightly linked at our site. Results also suggest that soil freezing at temperatures commonly experienced at this site can further increase net nitrification in soils already exhibiting high nitrification from N saturation.

Effect Of Snake Populations On Salamanders As A Result Of Forest Fragmentation, Casey Renee Bradshaw

Theses, Dissertations and Capstones

Forest fragmentation is one of the main causes for the loss of native biodiversity. One consequence is increased proportion of edge habitat that introduces new “edge” species, and makes habitat for interior forest-living species less-suitable. This study was conducted at three sites in Tucker County, West Virginia and included one downhill ski slope, one cross country ski slope, and one gravel road. The main objectives of this study were to determine relative abundance of snake communities, how far species move from edge habitat into the forest and to determine whether snakes are a predatory threat to salamanders, specifically the federally …

Global Assessment Of Nitrogen Deposition Effects On Terrestrial Plant Diversity : A Synthesis, R. Bobbink, K. Hicks, J. Galloway, T. Spranger, R. Alkemade, M. Ashmore, M. Bustamante, S. Cinderby, E. Davidson, F. Dentener, B. Emmett, J. W. Erisman, M. Fenn, Frank S. Gilliam, A. Nordin, L. Pardo, W. Devries

Biological Sciences Faculty Research

Atmospheric nitrogen (N) deposition is a recognized threat to plant diversity in temperate and northern parts of Europe and North America. This paper assesses evidence from field experiments for N deposition effects and thresholds for terrestrial plant diversity protection across a latitudinal range of main categories of ecosystems, from arctic and boreal systems to tropical forests. Current thinking on the mechanisms of N deposition effects on plant diversity, the global distribution of G200 ecoregions, and current and future (2030) estimates of atmospheric N-deposition rates are then used to identify the risks to plant diversity in all major ecosystem types now …

The Ecological Significance Of The Herbaceous Layer In Temperate Forest Ecosystems, Frank S. Gilliam

Biological Sciences Faculty Research

Despite a growing awareness that the herbaceous layer serves a special role in maintaining the structure and function of forests, this stratum remainsan underappreciated aspect of forest ecosystems. In this article I review and synthesize information concerning the herb layer’s structure,composition, and dynamics to emphasize its role as an integral component of forest ecosystems. Because species diversity is highest in the herb layeramong all forest strata, forest biodiversity is largely a function of the herb-layer community. Competitive interactions within the herb layer candetermine the initial success of plants occupying higher strata, including the regeneration of dominant overstory tree species. Furthermore, …

Satellite Image Processing For Biodiversity Conservation And Environmental Modeling In Kyrgyz Republic National Park, Galina N. Fet

Theses, Dissertations and Capstones

There is a need for extensive surveys of living organisms at a global scale; digital data exchange and storage is an essential part of such studies. Biodiversity inventory of fungi, which play an essential role in the health of the mountainous conifer forests of a developing country – Kyrgyz Republic, was linked to the vegetation classification produced from the high-resolution satellite imagery. Terra ASTER and SRTM90 imagery was used as a base map for the ecosystem modeling of the species and habitat distribution and for the three-dimensional representation, especially valuable for the mountainous landscapes of the Ala Archa National Park. …

Response Of The Herbaceous Layer Of Forest Ecosystems To Excess Nitrogen Deposition, Frank S. Gilliam

Biological Sciences Faculty Research

• 1 This review brings into focus what is known about the response of the herbaceous layer of forest ecosystems to increasing nitrogen deposition. The emphasis on forests in general is important for two reasons. First, forests often occupy areas receiving high rates of atmospheric deposition of N. Second, compared with herb-dominated communities, about which much is known regarding response to excess N, forests generally display greater biological and structural complexity. The more specific focus on the herbaceous layer – here defined as all vascular (herbaceous and woody) plants ≤ 1 m in height – is warranted because most of the …

Natural Disturbances And The Physiognomy Of Pine Savannas : A Phenomenological Model, Frank S. Gilliam, William J. Platt, Robert K. Peet

Biological Sciences Faculty Research

Abstract. Question: The decline of the Pinus palustris ecosystems has resulted from anthropogenic influences, such as conversion to pine plantation forestry, agriculture and land development, all of which are closely related to increases in human populations. Other effects, however, have arisen from alterations in disturbance regimes that maintain the structure and function of these ecosystems. How have alterations of the disturbance regime altered the physiognomy of ‘old-growth’ stands, and what are the implications for ecosystem conservation and restoration?

Methods: In contrast to models that emphasize close interactions among the vertically complex strata, we develop a conceptual phenomenological model for the …

Conservation And Restoration Of The Pinus Palustris Ecosystem, Frank S. Gilliam, William J. Platt

Biological Sciences Faculty Research

The well-documented decline of the Pinus palustris ecosystem has resulted from several anthropogenic influences, such as forest clearing (e.g. pine plantation forestry, agriculture) and urban development, both of which are closely related to increases in human populations. Other impacts have arisen from alterations in disturbance regimes responsible for maintaining the structure and function of these ecosystems. Restoration and management of degraded pine savanna ecosystems is critical. Identification of ecological processes that determine the structure and function of the intact system are important because successful restoration efforts should be based on sound scientific understanding. In this paper, we introduce this special …

Effects Of Atmospheric Nitrogen Deposition On The Herbaceous Layer Of A Central Appalachian Hardwood Forest, Frank S. Gilliam, Anne W. Hockenberry, Mary Beth Adams

Biological Sciences Faculty Research

Additions of nitrogen (N) have been shown to alter species diversity of plant communities, with most experimental studies having been carried out in communities dominated by herbaceous species. We examined seasonal and inter-annual patterns of change in the herbaceous layer of two watersheds of a central Appalachian hardwood forest that differed in experimental treatment. This study was carried out at the Fernow Experimental Forest, West Virginia, using two adjacent watersheds: WS4 (mature, second-growth hardwood stand, untreated reference), and WS3. Seven circular 0.04-ha sample plots were established in eachwatershed to represent its full range of elevation and slope aspect. The herbaceous …

Acidification And Nutrient Cycling, Mary Beth Adams, William T. Peterjohn, Frank S. Gilliam

Biological Sciences Faculty Research

Additions of acid anions can alter the cycling of other nutrients and elements within an ecosystem. As strong acid ions move through a forest, they may increase the concentrations of nitrogen (N) and sulfur (S) in the soil solution and stream water. Such treatments also may increase or decrease the availability of other anions, cations and metal ions in the soil. A number of studies in Europe and North America have documented increases in base cation concentrations such as calcium (Ca) and magnesium (Mg) with increased N and S deposition (Foster and Nicolson 1988, Feger 1992, Norton et al. 1994, …

Soil Chemical Response To Experimental Acidification Treatments, Mary Beth Adams, David R. Dewalle, William T. Peterjohn, Frank S. Gilliam, William E. Sharpe, Karl W.J. Williard

Biological Sciences Faculty Research

One of the conclusions reached during the Congressionally mandated National Acid Precipitation Program (NAPAP) was that, compared to ozone and other stress factors, the direct effects of acidic deposition on forest health and productivity were likely to be relatively minor. However, the report also concluded “the possibility of long-term (several decades) adverse effects on some soils appears realistic” (Barnard et al. 1990). Possible mechanisms for these long-term effects include: (1) accelerated leaching of base cations from soils and foliage, (2) increased mobilization of aluminum (Al) and other metals such as manganese (Mn), (3) inhibition of soil biological processes, including organic …

Vegetation And Acidification, David R. Dewalle, James N. Kochenderfer, Mary Beth Adams, Gary W. Miller, Frank S. Gilliam, Frederica Wood, Stephanie S. Odenwald-Clemens, William E. Sharpe

Biological Sciences Faculty Research

In this chapter, the impact of watershed acidification treatments on WS3 at the Fernow Experimental Forest (FEF) and at WS9 on vegetation is presented and summarized in a comprehensive way for the first time. WS7 is used as a vegetative reference basin for WS3, while untreated plots within WS9 are used as a vegetative reference for WS9. Bioindicators of acidification impacts that will be considered include several measures of tree and stand growth rates, foliar chemistry, bolewood chemistry, and herbaceous species composition and diversity. These studies enhance our understanding of the inter-relationships of changes in soil conditions caused by the …

Interspecific Divergence In Foliar Nutrient Dynamics And Stem Growth In A Temperate Forest In Response To Chronic Nitrogen Inputs, Jeffrey D. May, Sarah Beth Burdette, Frank S. Gilliam, Mary Beth Adams

Biological Sciences Faculty Research

We studied the effects of excessive nitrogen (N) fertilization on foliar nutrient dynamics and stem growth in three important tree species in a mixed-deciduous forest. Stem diameter growth, foliar N concentrations, nitrogen–phosphorus (N/P) ratios, and nutrient resorption were determined for Acer rubrum L. (ACRU), Liriodendron tulipifera L. (LITU), and Prunus serotina Ehrh. (PRSE) on two 30-year-old watersheds at the Fernow Experimental Forest, West Virginia, USA: WS3, fertilized annually with 35 kg ammonium sulfate·ha^-1 since 1989, and WS7, an untreated control watershed. In an earlier (1992) study, foliar N concentrations of all three species averaged 11% higher in WS3 than …

Interspecific Divergence In Foliar Nutrient Dynamics And Stem Growth In A Temperate Forest In Response To Chronic Nitrogen Inputs, Jeffrey D. May, Sarah Beth Burdette, Frank S. Gilliam, Mary Beth Adams

Biological Sciences Faculty Research

We studied the effects of excessive nitrogen (N) fertilization on foliar nutrient dynamics and stem growth in three important tree species in a mixed-deciduous forest. Stem diameter growth, foliar N concentrations, nitrogen–phosphorus (N/P) ratios, and nutrient resorption were determined for Acer rubrum L. (ACRU), Liriodendron tulipifera L. (LITU), and Prunus serotina Ehrh. (PRSE) on two 30-year-old watersheds at the Fernow Experimental Forest, West Virginia, USA: WS3, fertilized annually with 35 kg ammonium sulfate ha–1 since 1989, and WS7, an untreated control watershed. In an earlier (1992) study, foliar N concentrations of all three species averaged 11% higher in WS3 than …

Effects Of Silvicultural Practices On Soil Carbon And Nitrogen In A Nitrogen Saturated Central Appalachian (Usa) Hardwood Forest Ecosystem, Frank S. Gilliam, David A. Dick, Michelle L. Kerr, Mary Beth Adams

Biological Sciences Faculty Research

Silvicultural treatments represent disturbances to forest ecosystems often resulting in transient increases in net nitrification and leaching of nitrate and base cations from the soil. Response of soil carbon (C) is more complex, decreasing from enhanced soil respiration and increasing from enhanced postharvest inputs of detritus. Because nitrogen (N) saturation can have similar effects on cation mobility, timber harvesting in N-saturated forests may contribute to a decline in both soil C and base cation fertility, decreasing tree growth. Although studies have addressed effects of either forest harvesting or N saturation separately, few data exist on their combined effects. Our study …

Effects Of Excessive Nitrogen Deposition On Foliar Nutrient Dynamics, Nutrient Concentrations, And Nutrient Ratios In A Central Appalachian Hardwood Forest, Sarah Beth Burdette

Theses, Dissertations and Capstones

Many forested systems of the eastern US are becoming significantly nitrogen (N) saturated due to chronic N deposition from the atmosphere. Nitrogen saturation has the potential to alter important internal plant processes, such as nutrient cycling. While extensive research has been conducted on the responses of soil to N saturation, considerably less research has focused on the response of plant nutrient dynamics, nutrient concentrations, and nutrient ratios to excess N. Research was conducted on two watersheds at the Fernow Experimental Forest in West Virginia: WS3 (fertilized with ammonium sulfate annually since 1989) and WS7 (untreated control). Presenescent and senesced leaves …

A Gis Approach For The Abandoned Mine Inventory With Haul Roads Of The Monongahela National Forest To Provide A Comparative Analysis Of Usfs/Usace Collection Procedures And Image Basemap Selection For Cartographic Representation, John R. Ferguson Ii

Theses, Dissertations and Capstones

The Monongahela National Forest spans 10 counties in eastern central West Virginia. It has been an area of high mining and timbering activities throughout much of the early to mid twentieth century. As a result, the United States Forest Service (USFS) has focused reclamation and remediation efforts on the abandoned mine land areas. Much of the area has been subjected to mining after effects such as acid mine drainage, structural remains, gob/spoil piles, garbage piles, mine portals, and highwalls. In 1998 the USFS contracted with the U.S. Army Corps of Engineers (USACE) to locate these mining remains and provide a …

Effects Of Harvesting On Herbaceous Layer Diversity Of A Central Appalachian Hardwood Forest, Frank S. Gilliam

Biological Sciences Faculty Research

Clearcutting is a common harvesting practice in many eastern hardwood forests. Among the vegetation strata of these forests, the herbaceous layer is potentially the most sensitive in its response to harvest-mediated disturbances and has the highest species diversity. Thus, it is important to understand the response of herbaceous layer diversity to forest harvesting. Previous work on clearcut and mature stands at the Fernow Experimental Forest (FEF), West Virginia, has shown that, although, harvesting did not alter appreciably herbaceous layer cover, it influenced the relationship of cover to biotic and abiotic factors, such as tree density and soil nutrients, respectively. The …

Temporal And Spatial Variation Of Nitrogen Transformations In Nitrogen-Saturated Soils Of A Central Appalachian Hardwood Forest, Frank S. Gilliam, Bradley M. Yurish, Mary Beth Adams

Biological Sciences Faculty Research

We studied temporal and spatial patterns of soil nitrogen (N) dynamics from 1993 to 1995 in three watersheds of Fernow Experimental Forest, W.V.: WS7 (24-year-old, untreated); WS4 (mature, untreated); and WS3 (24-year-old, treated with (NH4)2SO since 1989 at the rate of 35 kg Nha–1year–1). Net nitrification was 141, 114, and115 kg Nha–1year–1, for WS3, WS4, and WS7, respectively, essentially 100% of net N mineralization for all watersheds. Temporal (seasonal) patterns of nitrification were significantly related to soil moisture and ambient temperaturein untreated watersheds only. Spatial patterns of soil water NO3–of WS4 suggest that microenvironmental variabilitylimits rates of N processing in …

Factors Influencing Spatial Variability In Nitrogen Processing In Nitrogen-Saturated Soils, Frank S. Gilliam, Charles C. Somerville, Nikki L. Lyttle, Mary Beth Adams

Biological Sciences Faculty Research

Nitrogen (N) saturation is an environmental concern for forests in the eastern U.S. Although several watersheds of the Fernow Experimental Forest (FEF), West Virginia exhibit symptoms of Nsaturation, many watersheds display a high degree of spatial variability in soil N processing. This study examined the effects of temperature on net N mineralization and nitrification in N-saturatedsoils from FEF, and how these effects varied between high N-processing vs. low N-processingsoils collected from two watersheds, WS3 (fertilized with [NH4]2SO4) and WS4 (untreated control). Samples of forest floor material (O2 horizon) and mineral soil (to a 5-cm depth) were taken from three subplots …

Vertical, Longitudinal And Seasonal Variation In The Benthic Fauna At The Fernow Experimential Forest, Parsons, West Virginia, Robert William Hood

Theses, Dissertations and Capstones

Multi-level basket samplers were placed within the shallow hyporheic zone (0 - 30 cm) of first through fourth-order streams in the Elklick Run drainage at the Fernow Experimental Forest, Parsons, West Virginia. Samplers were colonized by macroinvertebrates over three month intervals and collected during three different seasons. Seasonal, longitudinal, and vertical variation in the macroinvertebrate assemblage were examined to better understand the structure and function of the interstitial macroinvertebrate assemblage and important underlying factors. Surber samples were collected for comparison between surface and hyporheic macroinvertebrate assemblages. Temperature, dissolved oxygen, water chemistry, and interstitial water movement were measured at sampler locations. …

Four-Year And Seasonal Patterns Of Herbaceous Layer Development In Hardwood Stands Of The Fernow Experimental Forest, Parsons, Wv, Anne Wallace Hockenberry

Theses, Dissertations and Capstones

This study described patterns of herbaceous layer characteristics in watersheds of the Fernow Experimental Forest (FEF) in Parsons, WV. The purpose of this study was to examine four-year and seasonal patterns of change in herb layer species composition, cover, and richness within three stands of mixed hardwood forest. Sampling was conducted within seven circular plots (0.04 ha) in each of three watersheds of the FEF: WS3 (24-yr-old stand, previously clear-cut, receiving experimental nitrogen additions), WS7 (24-yr-old stand, previously clear-cut and treated with herbicide), and WS4 (>85-yr-old stand, untreated, control). All vascular plants~ lm in height were identified to species …

Patterns Of Soil Nitrogen Dynamics In A Central Appalachian Hardwood Forest Ecosystem, Bradley Michael Yurish

Theses, Dissertations and Capstones

Increasing awareness of the potential effects of acidic deposition prompted a watershed manipulation project at the Fernow Experimental Forest (FEF), West Virginia, in 1987. In this project, an acidifying agent, (Nf4)2S04 , was applied to an entire watershed to better understand ecosystem responses to simulated acid deposition in excess of ambient levels. In ecosystems receiving elevated N levels, increased soil N may contribute to soil acidification, lead to nutrient imbalances in plants, increase rates of nitrification, increase leaching of No3• and pollute surface waters. This study was designed to assess changes of three watersheds that received varied N inputs. The …

The Effects Of Wildfire On The Herbaceous Layer Of A Southwestern West Virginia Mixed Hardwood Forest, Michael K. Nowlin

Theses, Dissertations and Capstones

The effects of fire on the mixed hardwood forests in West Virginia have long been of interest both biologically and economically. This study examined the effects of a Fall, 1991 wildfire in Wayne County, West Virginia on soil fertility and herbaceous layer response.

Fire intensity was determined by comparison of 1) remaining organic material, 2) understory survival and 3) height of charcoal on the stems of trees following the fire. Study plots were established in areas representing fires of high, moderate and low intensities with a control established in an adjacent unburned area of similar aspect and elevation.

Composite soil …

Factors Influencing Herbaceous Layer Vegetation Of A Central Appalachian Hardwood Forest Ecosystem, Nicole Leigh Turrill

Theses, Dissertations and Capstones

The herb layer (vascular plants ≤1 m in height) of montane forests responds to a variety of environmental gradient conditions. This study compared herb layer characteristics, soil fertility, and plant-soil nutrient interactions of two young (~20 yr) and two mature (~>70 yr) watersheds of the Fernow Experimental Forest, Parsons, West Virginia. Fifteen 0.04-ha circular plots were established in each of four watersheds [WS4 control, >80 yr; WS13 select cut, >65 yr; WS3 clearcut, ~20 yr; WS7 clearcut/herbicide, ~20 yr] to sample the extremes of watershed elevation and aspect. Cover (%) was measured for all vascular species (≤1 m in …

Vascular Flora Of Three Watersheds In The Fernow Experimental Forest And Factors Influencing Species Composition Of The Herbaceous Layer, Staci Dudash Smith Aulick

Theses, Dissertations and Capstones

Herbaceous layer communities respond to a number of environmental variables in forests: thus, study of the herb layer is essential for a more complete understanding of the forest ecosystem. The objectives of this study were to (1) describe the flora of the watersheds. (2) describe the species richness and diversity of watersheds 3, 4 , and 7 of the Fernow Experimental Forest, Parsons, West Virginia, and (3) examine stand and soil variables and their influence on the herbaceous communities or the forest. Watersheds were selected on the basis of stand age/history (WS3 "20yr, clearcut; WS7 "20 yr. clearcut and herbicide: …