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Full-Text Articles in Other Ecology and Evolutionary Biology

Prairie Dogs, Gary Witmer, Jon Grant, Kendra Cross Mar 2023

Prairie Dogs, Gary Witmer, Jon Grant, Kendra Cross

Wildlife Damage Management Technical Series

Prairie dogs (Cynomys spp.) occur throughout the prairie states of middle North America from Mexico northward into Canada. They occupy a variety of habitats from prairies to high mountain valleys and sage brush-dominated deserts. The most common species is the black-tailed prairie dog (Cynomys ludovicianus; Figure 1). Prairie dogs are considered a “keystone species.” They provide habitat for many other native, grassland species. Prairie dogs live in colonies or “towns” that can span hundreds to thousands of acres. Depending on the species, their presence is evident by their burrow system. Despite the many ecosystem benefits prairie dogs provide by modifying …


Table Of Contents Jan 2023

Table Of Contents

Human–Wildlife Interactions

This is the table of contents.


Call For Associate Editors, S. Nicole Frey Jan 2023

Call For Associate Editors, S. Nicole Frey

Human–Wildlife Interactions

This is a call for associate editors for Human-Wildlife Interactions.


Carnivore And Ungulate Occurrence In A Fire-Prone Region, Sara J. Moriarty-Graves Jan 2023

Carnivore And Ungulate Occurrence In A Fire-Prone Region, Sara J. Moriarty-Graves

Cal Poly Humboldt theses and projects

Increasing fire size and severity in the western United States causes changes to ecosystems, species’ habitat use, and interspecific interactions. Wide-ranging carnivore and ungulate mammalian species and their interactions may be influenced by an increase in fire activity in northern California. Depending on the fire characteristics, ungulates may benefit from burned habitat due to an increase in forage availability, while carnivore species may be differentially impacted, but ultimately driven by bottom-up processes from a shift in prey availability. I used a three-step approach to estimate the single-species occupancy of four large mammal species: mountain lion (Puma concolor), coyote …


Individual And Population Responses To Hydrologic Variability In A Headwater Stream Salamander, Madaline Cochrane Jan 2023

Individual And Population Responses To Hydrologic Variability In A Headwater Stream Salamander, Madaline Cochrane

Graduate Student Theses, Dissertations, & Professional Papers

Understanding how organisms respond to environmental variability is a central goal in ecology – a goal made even more pressing by the herculean challenge global climate change presents to all organisms. Climate change is increasing the frequency and intensity of floods and droughts, which will likely have disproportionate effects on freshwater organisms. Many stream-associated species have multi-stage life histories. However, we lack an empirical understanding of life history and movement responses of these organisms to hydrologic disturbances, and how these responses may influence demographic rates. In my dissertation, I used a combination of growth, developmental, movement, and demographic data to …


Quantitative Song Variety In Relation To Genotype In A Hybridizing Chickadee Population, Shelby Madison Palmer Jan 2023

Quantitative Song Variety In Relation To Genotype In A Hybridizing Chickadee Population, Shelby Madison Palmer

Graduate Theses/Dissertations

The Black-capped chickadee (Poecile atricapillus) and Carolina chickadee (P. carolinensis) are North American songbird species that hybridize in a narrow contact zone stretching latitudinally from New Jersey to Kansas, USA. The association between genetic ancestry and song type in this hybrid zone has been studied independently several times and found to be minimal or absent, likely due to the influence of cultural transmission on learned song in the oscine passerine clade to which the chickadees belong. Despite this, the song of both species remains remarkably distinct in allopatry, suggesting a genetic constraint on certain qualities of …


The Role Of Interspecific Competition In The Range Collapse Of A Songbird After Rapid Range Expansion, Zachary H. Vickers Jan 2023

The Role Of Interspecific Competition In The Range Collapse Of A Songbird After Rapid Range Expansion, Zachary H. Vickers

Graduate Theses/Dissertations

There is a wealth of evidence demonstrating interspecific competition in birds, but often this evidence is localized and may or may not entirely explain range dynamics over large geographic extents. Bewick’s Wrens (Thrymanes bewickii) and House Wrens (Troglodytes aedon) are small passerine birds of the family Troglodytidae. Previous experimental evidence has demonstrated that Bewick’s Wrens suffer from asymmetric interference competition from House Wrens in Kansas, and this evidence has been cited as the likely reason for the historically recent range collapse of the Bewick’s Wren. However, I argue that localized experimental evidence is insufficient to explain abundance trends over large …


Cover, Editorial Staff, Journal Information Jan 2023

Cover, Editorial Staff, Journal Information

Human–Wildlife Interactions

This includes the cover, editorial staff, and journal information.


Differences In Drifting Invertebrate Communities Across Arctic Ecozones And The Influence On Potential Growth Of Grayling (Thymallus Arcticus), Charles Chanyi Jan 2023

Differences In Drifting Invertebrate Communities Across Arctic Ecozones And The Influence On Potential Growth Of Grayling (Thymallus Arcticus), Charles Chanyi

Theses and Dissertations (Comprehensive)

Invertebrate drift is a key process that potentially affects multiple levels of food web organization within stream environments. However, our understanding of the mechanistic drivers of drift in high latitude streams and subsequent bottom-up control that drift may have on fish predators in these environments remains understudied. This project aimed to gain the baseline knowledge of how drift functions across two major high latitude ecozones, the boreal forest and tundra, and how those possible differences in drifting community characteristics may impact drift-feeding Arctic grayling (Thymallus arcticus). These objectives were accomplished by characterizing stream environments across both ecozones, sampling …


Geographic Distribution: Anolis Sagrei (Brown Anole). Usa: Arizona., Addie Leimroth, Louis A. Somma, Dyrana N. Russell, Logan P. Cutts, Mason Ryan, Randall D. Babb, Karen E. Hajek, Andrew T. Holycross Dec 2022

Geographic Distribution: Anolis Sagrei (Brown Anole). Usa: Arizona., Addie Leimroth, Louis A. Somma, Dyrana N. Russell, Logan P. Cutts, Mason Ryan, Randall D. Babb, Karen E. Hajek, Andrew T. Holycross

Papers in Herpetology

We discovered the first distributional records for the exotic, Caribbean lizard, Anolis sagrei (Brown Anole), in Arizona.


The Ecology And Evolution Of Species Rarity In Oaks (Quercus Spp.), Yingtong Wu Oct 2022

The Ecology And Evolution Of Species Rarity In Oaks (Quercus Spp.), Yingtong Wu

Dissertations

Rare species are susceptible to extinction due to ecological and genetic factors. Understanding the distribution, ecology, and evolution of rare species can provide useful information for effective conservation. To investigate species rarity, this dissertation focuses on a species-rich and ecologically diverse genus, Quercus (oaks). In Chapter 1, I aimed to understand how interactions between hosts and soil microbes contribute to habitat restriction in oak species. I performed a soil inoculum experiment on two pairs of sister oak species that show habitat divergence. I found that host-specific soil microbes contribute to habitat divergence and exclusion among sister species of oaks, but …


Full Issue, Frank R. Thompson Iii Sep 2022

Full Issue, Frank R. Thompson Iii

National Quail Symposium Proceedings

No abstract provided.


The Evolution Of Quail Research: Trends In Themes, Cognitive Extent, And Lexical Diversity, Fidel Hernández Sep 2022

The Evolution Of Quail Research: Trends In Themes, Cognitive Extent, And Lexical Diversity, Fidel Hernández

National Quail Symposium Proceedings

Language is an interesting characteristic that is unique to humans. Language represents a method of human communication and is believed to reflect a person’s view of reality (Kramsch 2004). The words used by a person or a community provide insight into the ideas, concepts, and worldview held by people (Pennebaker et al. 2003). In science, publications represent a primary form of communication of ideas among scientists. Publications provide a historical record of a discipline and reflect the relative interest of a scientific community in particular concepts during a given era (Kim et al. 2018). Consequently, how word use in publications …


Effects Of Climate Change On Northern Bobwhite Nesting Chronology And Clutch Size, Justin A. Rectenwald, D. Clay Sisson, James A. Martin Sep 2022

Effects Of Climate Change On Northern Bobwhite Nesting Chronology And Clutch Size, Justin A. Rectenwald, D. Clay Sisson, James A. Martin

National Quail Symposium Proceedings

Widespread changes to breeding bird phenology in response to climate change have been apparent in North America for several decades. While the impact of an earlier breeding season may be minimal by itself, changes in community-level interactions can be greatly influenced because of varying responses to climate change in different trophic levels. Climate change has been shown to alter the onset of breeding season and chick survival, and lead to population declines for game birds in high latitudes, at high elevations, and on the periphery of their range. The topic of climate change in relation to northern bobwhite (Colinus …


Examining A Habitat-Weather Threshold For Northern Bobwhite Populations In The Southwestern United States, John T. Edwards, Fidel Hernández, Andrea Montalvo, Masahiro Ohnishi Sep 2022

Examining A Habitat-Weather Threshold For Northern Bobwhite Populations In The Southwestern United States, John T. Edwards, Fidel Hernández, Andrea Montalvo, Masahiro Ohnishi

National Quail Symposium Proceedings

In semiarid portions of the northern bobwhite (Colinus virginianus; hereafter, bobwhite) geographic distribution, weather is a strong driver of interannual abundance. However, the strength of this relationship may depend on habitat amount. Given this habitat–weather dependence, there is likely to be a threshold value for habitat that determines how strongly a bobwhite population responds to weather. Our objective was to evaluate the relationship between habitat amount and the relative influence of weather on bobwhite abundance in Texas and Oklahoma, USA and determine a potential land-cover threshold value. We collected bobwhite abundance and land-cover data from the Breeding Bird …


Climate Impacts On North American Quail, Shelby M. Perry, Erin Moser, Jeffrey G. Whitt, Kelly S. Reyna Sep 2022

Climate Impacts On North American Quail, Shelby M. Perry, Erin Moser, Jeffrey G. Whitt, Kelly S. Reyna

National Quail Symposium Proceedings

North America’s quail population trends are often linked to regional climate. Extreme climate events such as severe drought, hard freezes, or excessive winter precipitation can reduce quail populations by as much as 84%. Above-average spring and summer temperatures coincident with drought can reduce the laying season for quail by ≤60 days. Exposure of quail eggs to high temperatures during preincubation can initiate and alter embryonic development. Here, we review the impacts of extreme climate events and a changing climate on the survival, reproduction, and population trends of 6 North American quail species: California quail (Callipepla californica), Gambel’s quail …


Northern Bobwhite Response To Control Of Red Imported Fire Ants In The Gulf Coast Prairie Of Texas, Kelly M. Redmond, Eric D. Grahmann, Fidel Hernández, Leonard A. Brennan, Michael E. Morrow, Timothy Anderson Sep 2022

Northern Bobwhite Response To Control Of Red Imported Fire Ants In The Gulf Coast Prairie Of Texas, Kelly M. Redmond, Eric D. Grahmann, Fidel Hernández, Leonard A. Brennan, Michael E. Morrow, Timothy Anderson

National Quail Symposium Proceedings

The population decline of northern bobwhite (Colinus virginianus; hereafter, bobwhite) on the Texas Gulf Coast Prairie, USA is largely attributed to habitat loss. However, red imported fire ants (Solenopsis invicta) occur throughout the region and are considered a possible contributing factor to the bobwhite decline. The objectives of our study were to determine the influence of red imported fire ants on bobwhites by comparing bobwhite nest success, survival, and density between sites treated with fire ant bait (treatment) and reference (control) sites. Our study was conducted on 3 private ranches in Goliad and Refugio counties, Texas. …


Northern Bobwhite And Fire: A Review And Synthesis, David A. Weber, Evan P. Tanner, Theron M. Terhune Ii, J. Morgan Varner, James A. Martin Sep 2022

Northern Bobwhite And Fire: A Review And Synthesis, David A. Weber, Evan P. Tanner, Theron M. Terhune Ii, J. Morgan Varner, James A. Martin

National Quail Symposium Proceedings

Our understanding of the relationship between northern bobwhite (Colinus virginianus; hereafter, bobwhite) and fire began with Herbert Stoddard’s work in the early 20th century. Research on the topic has continued, but our application of fire is deeply rooted in Stoddard’s work, even as it has become evident that fire regimes must be adapted to variable environmental conditions that are evolving with a changing landscape and climate. A comprehensive review and synthesis of the literature on this topic would help formalize research advancements since Stoddard and identify knowledge gaps for future research. Results from experiments suggest fire creates favorable …


Bobwhite Response To Cattle Grazing In South Texas, Bradley K. Johnston, J. Alfonso Ortega-S., Leonard A. Brennan, Fidel Hernández, Humberto L. Perotto-Baldivieso Sep 2022

Bobwhite Response To Cattle Grazing In South Texas, Bradley K. Johnston, J. Alfonso Ortega-S., Leonard A. Brennan, Fidel Hernández, Humberto L. Perotto-Baldivieso

National Quail Symposium Proceedings

Range management practices to improve habitat for wildlife by reducing brush and increasing herbaceous plants, coupled with reduced stocking rates, can lead to dense stands of dominant grasses, such as four-flower trichloris (Trichloris pluriflora). This monoculture of trichloris creates dense vegetation unsuitable for northern bobwhite (Colinus virginianus; hereafter, bobwhite), reduces plant species diversity, and alters ecosystem functions. The objectives of this study are to 1) evaluate the effects of a proper cattle grazing regime to improve bobwhite habitat and 2) develop a management guide documenting how cattle grazing can be used as a tool to reduce …


Northern Bobwhite Response To Vegetation Management And Recovery In South Texas, Rachel A. Smith, Leonard A. Brennan, Humberto L. Perotto-Baldivieso, Fidel Hernández Sep 2022

Northern Bobwhite Response To Vegetation Management And Recovery In South Texas, Rachel A. Smith, Leonard A. Brennan, Humberto L. Perotto-Baldivieso, Fidel Hernández

National Quail Symposium Proceedings

The northern bobwhite (Colinus virginianus; hereafter, bobwhite) requires habitat structure and composition with grass cover for nesting, predator avoidance, and thermal refuge and forb cover for feeding on phytophagous arthropods and seeds. During the past 2 decades, many land managers with interest in promoting quail hunting opportunities have reduced or completely eliminated livestock across South Texas, USA, rangelands. Resting the land from grazing allows vegetation—especially grasses and forbs—to recover and thus provide nesting and foraging habitat for bobwhite and other birds. How bobwhite respond to postgrazing vegetation recovery is of keen interest to rangeland quail managers, but this …


A Simulation Model Of Sustained-Yield Harvest For Northern Bobwhite In South Texas, Joseph P. Sands, Stephen J. Demaso, Fidel Hernández, Leonard A. Brennan, Matthew J. Schnupp, Trent W. Teinert, Dale Rollins, Robert M. Perez Sep 2022

A Simulation Model Of Sustained-Yield Harvest For Northern Bobwhite In South Texas, Joseph P. Sands, Stephen J. Demaso, Fidel Hernández, Leonard A. Brennan, Matthew J. Schnupp, Trent W. Teinert, Dale Rollins, Robert M. Perez

National Quail Symposium Proceedings

Recommended sustainable harvest rates for northern bobwhite (Colinus virginianus) vary greatly and range from 25% to 70% of the prehunt population. Because northern bobwhite populations have declined across their geographic range, determining sustainable harvest levels is critical for effective management. Our objectives were to use simulation modeling to identify sustainable rates of bobwhite harvest, probability of population persistence, and minimum viable population estimates. We also conducted a sensitivity analysis to evaluate the impacts of harvest on northern bobwhite populations in Texas, USA. We constructed a simulation model using Program STELLA 9.0 for a hypothetical northern bobwhite population on …


Spatial And Temporal Analyses Of Bobwhite Hunting Dynamics, D.Abraham Woodard, Leonard A. Brennan, Fidel Hernández, Humberto L. Perotto-Baldivieso, Neal Wilkins Sep 2022

Spatial And Temporal Analyses Of Bobwhite Hunting Dynamics, D.Abraham Woodard, Leonard A. Brennan, Fidel Hernández, Humberto L. Perotto-Baldivieso, Neal Wilkins

National Quail Symposium Proceedings

A variety of factors influence the harvest of northern bobwhites (Colinus virginianus) and where that harvest occurs on a landscape. Many of these factors can be quantified and manipulated to distribute harvest pressure across time and space to meet desired spring densities. We collected spatial hunting metrics using global positioning system units on trucks and hunting dogs, along with detailed hunting logs from 211 quail hunts during the 2018–2019, 2019–2020, and 2020–2021 statewide hunting seasons in Jim Hogg County, Texas, USA. We found that hunting parties effectively covered 23.8 ± 0.3 hectares per hour, with hunts lasting 3.5 …


Evaluating The Harvest Rate Recommendation For Northern Bobwhites In South Texas, D. Abraham Woodard, Leonard A. Brennan, Fidel Hernández, Humberto L. Perotto-Baldivieso, Neal Wilkins, Andrea Montalvo Sep 2022

Evaluating The Harvest Rate Recommendation For Northern Bobwhites In South Texas, D. Abraham Woodard, Leonard A. Brennan, Fidel Hernández, Humberto L. Perotto-Baldivieso, Neal Wilkins, Andrea Montalvo

National Quail Symposium Proceedings

The current harvest rate recommendation for northern bobwhites (Colinus virginianus; hereafter, bobwhite) in South Texas, USA is 20% of the autumn population, including crippling loss. This recommendation is based on population simulations of empirical data. We completed the first field evaluation of the 20% harvest recommendation by comparing prehunting and posthunting bobwhite density estimates on a hunted and nonhunted site in South Texas during the 2018–2019, 2019–2020, and 2020–2021 statewide bobwhite hunting seasons in Jim Hogg County, Texas. We conducted line-transect distance sampling surveys on 4 occasions per year (early November, mid-December, late January, early March) from a …


Detection Rates Of Northern Bobwhite Coveys Using A Small Unmanned Aerial System-Mounted Thermal Camera, Megan E. Martin, Lori A. Hearon, Kristine O. Evans, Raymond B. Iglay, Jesse I. Morrison, Mark D. Mcconnell Sep 2022

Detection Rates Of Northern Bobwhite Coveys Using A Small Unmanned Aerial System-Mounted Thermal Camera, Megan E. Martin, Lori A. Hearon, Kristine O. Evans, Raymond B. Iglay, Jesse I. Morrison, Mark D. Mcconnell

National Quail Symposium Proceedings

The northern bobwhite (Colinus virginianus; hereafter, bobwhite) requires intensive monitoring to evaluate management efforts and determine harvest rates. However, traditional monitoring techniques (e.g., covey-call surveys) are labor-intensive and imprecise. Small unmanned aerial systems (sUASs) mounted with thermal cameras have demonstrated promise for monitoring multiple avian species and could provide a less intensive and more effective approach to monitoring bobwhite coveys, assuming coveys produce a recognizable heat signature. To assess sUAS monitoring, we evaluated the influence of bobwhite covey size (3, 6, and 12) and cover type (grass, shrub, and forest) on covey detectability by a sUAS equipped with …


Applying Multistate Mark-Recapture Models With State Uncertainty To Estimate Survival And Reproduction Of Quail, Rebekah E. Ruzicka Sep 2022

Applying Multistate Mark-Recapture Models With State Uncertainty To Estimate Survival And Reproduction Of Quail, Rebekah E. Ruzicka

National Quail Symposium Proceedings

Obtaining unbiased estimates of vital rates and understanding how vital rates change in response to environmental stimuli are a continual pursuit of ecologists. Multistate mark-recapture (MSMR) models provide a flexible framework for evaluating dependent vital rates in a comprehensive analysis. For example, a bird must remain alive during breeding season to initiate a nest (i.e., transition from a nonbreeding to a breeding state); thus, the probability that a bird initiates a nest is dependent on the probability that it is still alive. Traditional MSMR models allow only for the estimation of survival, detection, and state transition parameters and depend on …


Exploring The Relationships Between South Texas Northern Bobwhite Populations And Cecal Worms Via System Dynamics, Nicole J. Traub, Benjamin L. Turner, Leonard A. Brennan, Alan M. Fedynich Sep 2022

Exploring The Relationships Between South Texas Northern Bobwhite Populations And Cecal Worms Via System Dynamics, Nicole J. Traub, Benjamin L. Turner, Leonard A. Brennan, Alan M. Fedynich

National Quail Symposium Proceedings

Community ecology historically focused on plants and free-living organisms; however, problems such as defining habitat boundaries and obtaining adequate sample sizes arise when evaluating such communities. The unique nature of host-helminth systems allows parasite community ecologists to avoid these problems when testing ecological hypotheses. Unlike free-living communities that have artificially constructed boundaries, parasite communities have well-defined unambiguous boundaries within host individuals. Due to the inherently complex and dynamic nature of ecological systems, traditional experimental methods often require expensive, long-term trials beyond investigators’ time and resource budgets. Conversely, a system dynamics approach facilitates learning about such systems via simulation of ecosystem …


Exploring A Plant-Diversity Hypothesis To Explain Helminth Prevalence In Northern Bobwhite In Texas, John E. Herschberger, Shaelyn T. Rainey, Fidel Hernández, Kristyn G. Stewart, Andrea Montalvo, Lindsey K. Howard Sep 2022

Exploring A Plant-Diversity Hypothesis To Explain Helminth Prevalence In Northern Bobwhite In Texas, John E. Herschberger, Shaelyn T. Rainey, Fidel Hernández, Kristyn G. Stewart, Andrea Montalvo, Lindsey K. Howard

National Quail Symposium Proceedings

Helminths, in particular eyeworms (Oxyspirura petrowi), may be a factor influencing northern bobwhite (Colinus virginianus; hereafter, bobwhite) populations in Texas, USA. Mean eyeworm prevalence in Texas appears to be greater in the Rolling Plains (55.1%) than the Rio Grande Plains (16.4%), a pattern generally attributed to possible differences in the occurrence of insects, the intermediate hosts of eyeworms. We explored an alternative hypothesis centered on plant diversity. Many plants possess phytochemicals with anthelmintic properties. Because wild animals suffering from parasitic infestations are capable of self-medicating via diet selection, organisms foraging in diverse communities may be expected …


Histopathological Analysis Of Quails In The Trans-Pecos Ecoregion Of Texas, Trey E. Johnson, Carlos E. Gonzalez, Dale Rollins, Josh G. Cross, Ryan S. Luna Sep 2022

Histopathological Analysis Of Quails In The Trans-Pecos Ecoregion Of Texas, Trey E. Johnson, Carlos E. Gonzalez, Dale Rollins, Josh G. Cross, Ryan S. Luna

National Quail Symposium Proceedings

Quail populations in Texas, USA, have declined over the past few decades due primarily to habitat loss. The role that parasites may play in such declines has been a recent topic of concern. To help address this question, we collected 12 scaled quail (Callipepla squamata), 8 Gambel’s quail (Callipepla gambelii), and 3 Montezuma quail (Cyrtonyx montezumae) from across the Trans-Pecos ecoregion of Texas via hunter harvest, funnel traps, and night netting. Quail samples were necropsied to determine the abundance of eyeworms (Oxyspirura petrowi). Histopathological analyses were conducted on quail eyeballs and periocular …


Analysis Of Predator Avoidance Behavior In California Valley Quail, Curt Vandenberg, Jeffrey G. Whitt, Kelly S. Reyna Sep 2022

Analysis Of Predator Avoidance Behavior In California Valley Quail, Curt Vandenberg, Jeffrey G. Whitt, Kelly S. Reyna

National Quail Symposium Proceedings

Quail populations have been in decline across the United States, primarily due to habitat loss and climate. For remedy, landowners and game managers have attempted to restore populations by releasing captive-reared quail. These releases were largely unsuccessful, presumably due to high predation losses. Recently, there has been an increased interest in quail translocations, which tend to have lower mortality rates than captive-reared bird releases. Translocations are expensive and unpredictable, and require many person-hours; releasing captive-reared quail would be more efficient if the practice were successful. We compared predator avoidance behavior between captive-reared and wild-translocated California quail (Callipepla californica) …


Habitat Of The Long-Tailed Wood-Partridge In Central Mexico, David García-Solózano, Crisma Lopez-Sanchez, Edgardo Lopez-González, Carlos González-Rebeles Islas Sep 2022

Habitat Of The Long-Tailed Wood-Partridge In Central Mexico, David García-Solózano, Crisma Lopez-Sanchez, Edgardo Lopez-González, Carlos González-Rebeles Islas

National Quail Symposium Proceedings

The long-tailed wood-partridge (Dendrortyx macroura; hereafter, wood-partridge) is a forest quail endemic to the temperate forests of the Trans-Mexican Volcanic Belt of central Mexico and is considered threatened according to the Secretariat of the Environment of Mexico. We studied 34 sites within the Natural Resources Protection Area River Basins of the Valle de Bravo, Malacatepec, Tilostoc and Temascaltepec in central Mexico to evaluate wood-partridge habitat during September–December 2019. We evaluated attributes of tree, shrub, and herbaceous vegetation, canopy cover, humidity, slope, and altitude. We also identified vegetation used for nesting, food, and shelter. The scant information published for …