Evolution Commons^™

A Phylogeographic Study Of The Kentucky Endemic Cave Beetle Neaphaenops Tellkampfii Erichson (1844), Mahamad Miya

Masters Theses & Specialist Projects

Cave beetles are an excellent group to study evolutionary history and speciation by isolation within cave habitats. However, they are underrepresented in such studies, particularly in North America, one of the hotspots of cave biodiversity where beetles account for a significant proportion. The present research addresses this issue by focusing on the Kentucky endemic cave beetle, Neaphaenops tellkampfii, a species hypothesized to comprise four subspecies. Based on allozyme data, previous studies have noted high genetic variation within and high similarity among local populations of this beetle, suggesting complex evolutionary dynamics. In this regard, this study aims to elucidate the evolutionary …

Alfred Russel Wallace Notes 31. Wallace’S Personal Copy Of “Life Of Alexander Von Humboldt.”, Charles H. Smith

Faculty/Staff Personal Papers

Alfred Russel Wallace owned a personal copy of the two-volume 1873 English edition of the Life of Alexander von Humboldt, now held by the library of the Linnean Society of London. Each volume is highly annotated, and the nature of these notes, and their several-directioned importance, is here discussed.

Alfred Russel Wallace Notes 30. Wallace A Theist? Part I., Charles H. Smith

Faculty/Staff Personal Papers

Alfred Russel Wallace (1823−1913) has been portrayed as a ‘theist’ on a large number of occasions from his own time on to the present. In this, the first of a two part work, this assessment is questioned. In part one, the matter of Wallace’s personal philosophy and spiritual orientation is explored, the conclusion being that Wallace was a lifelong agnostic who can hardly be aligned with theism.

Bibliography, Charles H. Smith

Faculty/Staff Personal Papers

Bibliography of publications by Charles H. Smith.

Alfred Russel Wallace Notes 29. Extraterrestrial Entertainment: Are We Being Monitored By Alien Beings?, Charles H. Smith

Faculty/Staff Personal Papers

Alfred Russel Wallace (1823−1913) was an early advocate for the rational assessment of likelihood of extraterrestrial life. Current efforts to evaluate the situation have ranged from the heavily self-indulgent to the more objective, and it is still often difficult to decide just how much of the information we are receiving on the subject is dependable. An attempt is made here to cut through the haze and reduce the matter to elementals.

Twelve Wallace Myths, Charles H. Smith

Faculty/Staff Personal Papers

Alfred Russel Wallace’s (1823-1913) bicentennial year is a good time to take stock. In this presentation I discuss twelve Wallace-related issues that I feel have been poorly taken up. These range from the biological to the biographical, including subjects such as social criticism, human evolution, autobiographical memory, natural selection, national affinities, spiritualism, and wokeism.

Alfred Russel Wallace Notes 28. Wallace's 'Change Of Mind', Revisited, Charles H. Smith

Faculty/Staff Personal Papers

Alfred Russel Wallace (1823−1913) is best known for his work on the theory of evolution by natural selection, and studies on biogeography. This fame has not, however, prevented appraisals of his work that sometimes depart from rational interpretations of his actual words. In this study, the frequently-offered notion that his adoption of spiritualism in late 1866 caused him to reverse himself on the universality of natural selection is taken to task, with arguments linked to his own words on the subject.

Alfred Russel Wallace Notes 27. When Wallace Broke With Darwin., Charles H. Smith

Faculty/Staff Personal Papers

The year 1866 was the first year Alfred Russel Wallace showed definite signs of breaking with Darwin over the limits of natural selection. Attention is drawn to a July 1866 exchange of letters between the two, and how this foreshadowed what followed.

Alfred Russel Wallace Notes 25. Wallace And The 'Physical Environment'., Charles H. Smith

Faculty/Staff Personal Papers

Alfred Russel Wallace’s natural selection essay of 1858 has been held to frame a greater role for the physical environment in forcing selection regimes than we find in Darwin’s writings, but here that verdict is challenged by a re-examination of both the essay itself, and period usage of the term ‘physical.’

The Systems Measurement Of Mammalian Biotas, Part Two, Charles H. Smith, Patrick Georges, Ngoc Nguyen

Faculty/Staff Personal Papers

For a recent publication, the authors identified a seven-region model of mammal family distribution patterns, in which each unit contributes equally to the system’s overall statistical characteristics of diversity, despite its individual units having measurably different levels of diversity and endemism. This systemization presents a highly efficient descriptive model that can possibly be interpreted as a form of natural classification. An additional analysis of the same mode is described here, in which the seven-region model of the distribution of mammal families’ spatial affinities is shown to closely approach a most-probable-state arrangement, as assessed through combinatorics, raising some important questions about …

Alfred Russel Wallace Notes 24. Wallace At 200: Potential Subjects For Student Theses, Charles H. Smith

Faculty/Staff Personal Papers

The bicentennial of Alfred Russel Wallace’s birth in 2023 will likely produce a wide array of reviews of his life and work; here, we pause for a short look at some Wallace-related questions that might be adapted for student theses and dissertations. Some of the subjects treated fall in with established lines of research, while others are suggested by other Wallace interests or activities that have not been much explored.

Description, Diversity, And Phylogeography Of The North American Spider Beetle (Coleoptera: Ptinidae) Genus Coleotestudo, Robert Chambliss

Masters Theses & Specialist Projects

A new genus, Coleotestudo, is proposed for the American species currently placed in the genus Niptus LeConte and are transferred herein. Two new species from Mexico are also described as C. potosi n. sp. and C. nahua n. sp. Distinguishing morphological characteristics for the North American Coleotestudo species are given. The combined set of morphological evidence, DNA sequence data, and a Western New World distribution strongly support the hypothesis that Coleotestudo does not share a recent common ancestor with N. hololeucus and both represent unrelated clades of spider beetles.

During the summer of 2021, Coleotestudo was collected mainly from …

Alfred Russel Wallace Notes 22: Stumbling Blocks To An Understanding Of Wallace's Worldview, Charles H. Smith

Faculty/Staff Personal Papers

The writings of Alfred Russel Wallace (1823−1913) present a challenge to modern readers, both for the large range of subjects he entertained, and his unique point of view. In this short treatment, ten ‘stumbling blocks’ to an appreciation of Wallace’s thought are outlined which, it is suggested, have caused problems of interpretation in the past.

Alfred Russel Wallace Notes 21: Wallace & The Doorway To The Universe, Charles H. Smith

Faculty/Staff Personal Papers

An important yet largely unrecognized theme in the thought of Alfred Russel Wallace (1823−1913) was his insistence that all dependably-reported phenomena, even those of aberrant nature, were worthy of a respectful kind of attention: that is, a kind which did not automatically banish difficult subjects to the realm of myth or superstition. In this work, Wallace’s philosophy in this direction is documented, and linked to the world of post-Age-of-Enlightenment revisionism.

Alfred Russel Wallace Notes 19: Social Evolution's Useful Idiots, Charles H. Smith

Faculty/Staff Personal Papers

In today’s world liberals look at conservatives as the villains, and vice versa. How did this come to pass? In this essay a model of the biological roots of liberalism and conservatism is advanced; this is followed by a discussion of why cognitive dissonance may represent the key process in our social evolution. Alfred Russel Wallace’s experience with cognitive dissonance is then detailed, including how he dealt with it.

Alfred Russel Wallace Notes 18: Wallace On The Balance Of Nature, Charles H. Smith

Faculty/Staff Personal Papers

Alfred Russel Wallace (1823−1913) had a unique perspective on the ‘balance of nature’: he avoided classical thoughts on the subject, but nevertheless seems to have adopted elements of the ‘balance’ concept while acknowledging that irreversible change occurs at both the biological and environmental levels. Wallace’s position can be understood from his grounding in Humboldtian ‘equilibrium of forces’ thinking, and his turn toward ‘final causes’-based interpretations.

Diversity And Host Specificity Of Nycteribiid Bat Flies (Diptera: Nycteribiidae) In Kenya, Taylor Verrett

Masters Theses & Specialist Projects

Bat flies (Diptera: Nycteribiidae and Streblidae) are hematophagous ectoparasites of bats distributed globally. Members of Nycteribiidae are morphologically constrained relative to streblids, and are united by their lack of wings, dorso-ventrally compressed bodies, and a primary distribution across the Eastern hemisphere. Bats are principal reservoirs of infectious diseases, including viral zoonoses of important consideration to human health, but the overall high host specificity of bat flies has largely been thought to curb their potential as inter-specific vectors of bat-borne pathogens. However, nycteribiid diversity and host associations remain critically understudied in some geographic regions, and rare examples of nycteribiid bat flies …

Alfred Russel Wallace Notes 17: More On The South Asian Connection, Charles H. Smith

Faculty/Staff Personal Papers

During his school years the arts and cultural historian and metaphysician Ananda Coomaraswamy (1877-1947) was clearly influenced by Alfred Russel Wallace, and is known later to have corresponded with, and supplied information to, him. Up to now the connections involved had been obscure, but a new investigation suggests that Wallace had previously interacted with Ananda’s mother and/or father.

Alfred Russel Wallace Notes 16: The Flexible Wallace, Charles H. Smith

Faculty/Staff Personal Papers

Alfred Russel Wallace (1823−1913), though a naturalist of considerable renown, has sometimes been accused of inflexibility and inconsistency in his thinking. Many of his critics, however, ignore his basic approach to knowledge accumulation, which was explicitly incremental, based on a perceived need for constant reconsideration. Here, I: (1) identify five groups of Wallace critics who seem to be operating with prior agendas, and (2) provide lists of subjects on which he either changed his mind, or remained constant, according to individual situation.

Alfred Russel Wallace Notes 15. Wallace's Many "Hats": What Should We Call Him?, Charles H. Smith

Faculty/Staff Personal Papers

Alfred Russel Wallace (1823−1913) has been referred to by name through the aid of a variety of labels – some thirty or more, in fact – that link him to his emphases of attention. How many of these labels are/were justifiable? The assessment here is that he is probably best remembered as a ‘naturalist,’ ‘geographer,’ ‘evolutionist,’ and ‘social critic.’

Alfred Russel Wallace Notes 8: Wallace's Earliest Exposures To The Writings Of Alexander Von Humboldt, Charles H. Smith

Faculty/Staff Personal Papers

For many years it has been known that the German geographer and naturalist Alexander von Humboldt (1769-1859) exercised a considerable influence on Alfred Russel Wallace’s (1823-1913) decision to become a travelling naturalist. Almost no attention, however, has been given to the real possibility that von Humboldt exerted an even greater influence on Wallace’s general worldview. In this note, Wallace’s probable earliest contacts with von Humboldt’s writings are described.

Alfred Russel Wallace Notes 14: Background For Wallace's 1845 Kington Essay, Charles H. Smith

Faculty/Staff Personal Papers

‘An Essay, On the Best Method of Conducting the Kington Mechanic's Institution’ is one of the earliest written works by Alfred Russel Wallace (1823−1913) known to have reached print. The circumstances of its writing are still not clear, though I here review pertaining facts and theories that seem to indicate Wallace actually was the author of the essay, which had been questioned.

Alfred Russel Wallace Notes 12: How Good Was Wallace's Memory?, Charles H. Smith

Faculty/Staff Personal Papers

Alfred Russel Wallace (1823−1913) continues to be criticized for inconsistencies in his recollections of the earlier events in his life. This criticism, while not entirely unjust, has nevertheless been overplayed. Critics have not attended to the general understanding that self-biography is under the influence of two aspects of memory: that pertaining to remembrance of the qualities of past situations, and a secondary ability to assign absolutes of name or date to such memories. All evidence suggests that Wallace’s memory in the first sense was excellent throughout his life, but that he was prone to lapses of the second type.

Alfred Russel Wallace Notes 13: Wallace On Prayer, Charles H. Smith

Faculty/Staff Personal Papers

Alfred Russel Wallace (1823−1913) is known to most for his natural history explorations and theoretical biology, but he also developed thoughts on a number of subjects relatable to a wider appreciation of evolutionary cosmology. His adoption of spiritualism, for one, was attuned to this mission, and in turn his otherwise difficult-to-interpret two-sided position on prayer.

Many Miles Away: A Cautionary Tale, Charles H. Smith

Faculty/Staff Personal Papers

Reporter Kerry Phillips is in for the surprise of her life: her television station has been contacted on a matter so important that 'scarcely anything could be of greater interest,' and she has been asked to follow up on the story. Little does she know just how interesting her job is about to get!

A Phylogenetic Analysis Of Bostrichoidea (Coleoptera) And Revisions Of The Southern African Spider Beetle Genera Meziomorphum And Eutaphroptinus (Ptinidae: Coleoptera), Olivia M. Gearner

Masters Theses & Specialist Projects

Proposals for the internal relationships and classification of the bostrichoids are currently poorly supported, and almost all are based on morphology alone. This study improves upon on previous phylogenetic analyses of the group by including more taxa using the standard genes for many phylogenetic analyses. Cytochrome C oxidase subunit 1 (CO1), 28S small subunit rRNA, and 16S small subunit rRNA mitochondrial genes were sequenced or obtained from Genbank, then analyzed using parsimony and Bayesian analyses. Topologies differed depending on genes used. A three gene tree and a two gene (28S and CO1) tree both supported relationships in which a basal …

Species Distribution And Conservation Genetics Of The Upland And Midland Chorus Frogs (Pseudacris) In Kentucky, Tucker Cambridge

Masters Theses & Specialist Projects

The upland (Pseudacris feriarum) and midland (P. triseriata) chorus frogs are closely related cryptic species that are best distinguished genetically. The distribution of these species within the Commonwealth of Kentucky has previously been defined by only a handful of genetic samples, making delineation of range limits for each species difficult. Accurate understanding of species distributions, and the genetic structure within them, are vitally important for conservation management of amphibian species. In this study, I have collected genetic samples from across the putative ranges of P. triseriata and P. feriarum in Kentucky and used next-generation sequencing technology to generate more fine-scale …

The Mechanism Of Biotremor Production In The Veiled Chameleon (Chamaeleo Calyptratus), Samuel Tegge

Masters Theses & Specialist Projects

Vibratory communication has evolved in numerous animal groups, including insects, spiders, fishes, mammals, and was recently discovered in veiled chameleons (Chamaeleo calyptratus). I examined the mechanism by which C. calyptratus produce these biotremors. Muscle activity data were gathered during simulated anti-predator responses via electromyography (EMG) with simultaneous recordings of biotremor production using an accelerometer. I correlated EMG data with the accelerometer data to implicate the muscles responsible for the production of the biotremors. Mixed-effect linear regression models described the mechanism, and a model selection framework determined which model fit the data best. I then used an analysis of variance to …

Body Shape Diversification Of Pecos Pupfish (Cyprinodon Pecosensis) On Varying Habitats As Evaluated By Geometric Morphometrics, Qianna Xu

Masters Theses & Specialist Projects

During the 19th and 20th centuries, alterations to the Pecos River in New Mexico and Texas, USA due to anthropogenic activities, including damning and river channelization, vast water extraction for irrigation, as well as pollution of associated habitats, have greatly impacted the fish fauna within the drainage. One of the endemic fish species, the Pecos pupfish (Cyprinodon pecosensis), might be the most affected. Historically abundant and widespread large populations have been disrupted and became a series of small isolated subpopulations that persist at a few highly fragmented habitats restricted to a small area in southern New Mexico. The connectivity among …

Wallace And Incipient Structures: A World Of "More Recondite" Influences, Charles H. Smith

DLPS Faculty Publications

Alfred Russel Wallace is well-known for his co-discovery of the principle of natural selection. Natural selection is usually considered a process, but it is not clear that Wallace regarded it in exactly these terms. In fact he more likely thought of the relationships involved as representing what we would now term a “state space,” a negative feedback loop wherein populations are maintained at healthy levels through elimination of the unfit. Both before and after the advent of natural selection Wallace clung to the idea that “more recondite forces” were shaping the nature and direction of evolution; this is especially evident …