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Articles 31 - 60 of 124
Full-Text Articles in Behavior and Ethology
James Davidson Fawcett (1933–2020): Imbibing With The Kiwi, Louis A. Somma
James Davidson Fawcett (1933–2020): Imbibing With The Kiwi, Louis A. Somma
Papers in Herpetology
An obituary and summary of the life of James D. Fawcett (1933-2020), herpetologist and instructor and professor of Biology at University of Nebraska at Omaha, 1972-2015. Includes bibliography of his works, list of master's theses chaired, and recollections of former students.
Dog And Owner Characteristics Predict Training Success, Jeffrey R. Stevens, London M. Wolff, Megan Bosworth, Jill Morstad
Dog And Owner Characteristics Predict Training Success, Jeffrey R. Stevens, London M. Wolff, Megan Bosworth, Jill Morstad
Jeffrey Stevens Publications
Teaching owners how to train their dogs is an important part of maintaining the health and safety of dogs and people. Yet we do not know what behavioral characteristics of dogs and their owners are relevant to dog training or if owner cognitive abilities play a role in training success. The aim of this study is to determine which characteristics of both dogs and owners predict success in completing the American Kennel Club Canine Good Citizen training program. Before the first session of a dog training course, owners completed surveys evaluating the behavior and cognition of their dog and themselves. …
Are Capuchin Monkeys (Sapajus Spp.) Sensitive To Lost Opportunities? The Role Of Opportunity Costs In Intertemporal Choice, Elsa Addessi, Valeria Tierno, Valentina Focaroli, Federica Rossi, Serena Gastaldi, Francesca De Petrillo, Fabio Paglieri, Jeffrey R. Stevens
Are Capuchin Monkeys (Sapajus Spp.) Sensitive To Lost Opportunities? The Role Of Opportunity Costs In Intertemporal Choice, Elsa Addessi, Valeria Tierno, Valentina Focaroli, Federica Rossi, Serena Gastaldi, Francesca De Petrillo, Fabio Paglieri, Jeffrey R. Stevens
Jeffrey Stevens Publications
Principles of economics predict that the costs associated with obtaining rewards can influence choice. When individuals face choices between a smaller, immediate option and a larger, later option, they often experience opportunity costs associated with waiting for delayed rewards because they must forego the opportunity to make other choices. We evaluated how reducing opportunity costs affects delay tolerance in capuchin monkeys. After choosing the larger option, in the High cost condition, subjects had to wait for the delay to expire, whereas in the Low cost different and Low cost same conditions, they could perform a new choice during the delay. …
Characterizing The Host Usage And Development Of Ormia Lineifrons (Diptera: Tachinidae), Kyler J. Rogers
Characterizing The Host Usage And Development Of Ormia Lineifrons (Diptera: Tachinidae), Kyler J. Rogers
Murray State Theses and Dissertations
Insect communication systems are strongly driven by the evolution of signals or signal preferences. These systems rely on a signaler to truthfully emit signals for receivers to interpret. Often, these signals are conspicuously broadcasted. Conspicuous signals involved in animal communication are intended to attract mates, however, these signals are often exploited by eavesdroppers. Thus, many communication systems experience natural selection and sexual selection acting in opposite directions. New adaptations can arise in response to selective pressures, such as eavesdroppers, leading to co-evolving systems between eavesdroppers and hosts, for example. Understanding these systems can provide valuable insight into how unintended receivers …
Influence Of Testosterone On Male-Male Competition In The Red-Sided Garter Snake, Thamnophis Sirtalis Parietalis, Isabella M. G. Bukovich
Influence Of Testosterone On Male-Male Competition In The Red-Sided Garter Snake, Thamnophis Sirtalis Parietalis, Isabella M. G. Bukovich
Senior Honors Projects, 2020-current
Female mimicry in the red-sided garter snake, Thamnophis sirtalis parietalis, is described in several well-studied evolutionary advantages, but an aspect that remains elusive is how reproductive traits are linked to the 3-fold higher circulating androgens in males with this phenotype. In this study, we implanted male garter snakes (n=15 per group) with either a blank implant (SHAM; control) or a T implant, the latter simulating the female mimic phenotype. Following simulated low-temperature dormancy, males were scored for courtship intensity over three days of behavioral trials with females housed in the same facility. Males were sacrificed, and sperm were collected …
Hind Wing Eyespots Of Brassolini Butterflies (Lepidoptera, Nymphalidae): Evolutionary Diversification And Functions In Anti-Predator Defense And Mating Behavior, Logan Crees
LSU New Orleans Theses and Dissertations
Ventral hind wing eyespots are prominent pattern elements in Brassolini butterflies, likely functioning in predator-prey interactions and reproductive activities. Caligo and Opsiphanes differ in male mate-seeking behaviors and it has been suggested that Caligo females use the male cua1 eyespot as a mate-locating cue, but Opsiphanes females do not seem to do so. We predict Caligo males should have larger eyespots than congeneric females, but the sexes would not differ in eyespot size in Opsiphanes. Our analyses supported both these predictions. Displacing the eyespot to the center of the wing makes eyespots more conspicuous, we asked if eyespot position …
Encephalization In Commensal Raccoons: A Unique Test Of The Cognitive Buffer Hypothesis, Peter M. Anderson
Encephalization In Commensal Raccoons: A Unique Test Of The Cognitive Buffer Hypothesis, Peter M. Anderson
Electronic Theses and Dissertations
This study investigated selective pressures associated with encephalization in mammals and discussed broader implications. Relative brain size as measured by EQ (Encephalization Quotient) was compared between ecological categories. Omnivores had higher average EQ than ecological specialists. Since specialists are disproportionately affected by extinction events, selection for ecological generalism is proposed as encephalization mechanism. This mechanism may reinforce the more widely known Cognitive Buffer Hypothesis (CBH)—the idea that possessing relatively large brains has buffered lineages against environmental change. CBH is tested here by comparing EQs in Procyon lotor (raccoon) in urban and rural environments. CBH predicts that raccoons in the most …
From Organisms To Ecosystems: Impacts Of Limb Loss And Regeneration On Crayfish Behavior, Luc Arnaud Dunoyer
From Organisms To Ecosystems: Impacts Of Limb Loss And Regeneration On Crayfish Behavior, Luc Arnaud Dunoyer
Theses and Dissertations--Biology
The ability of some organisms to regenerate tissues and organs has fascinated naturalists since antiquity, dating back to the earliest accounts of scientific inquiry with Aristotle in Ancient Greece. Then, Darwin’s theory invigorated some scientists’ dream of stimulating (or reactivating) regenerative capacities in human beings by showing that we are related to highly regenerative organisms. More recently, a renewed interest in discovering the molecular and genetic basis for organ and tissue regeneration has led biologists to focus more specifically on a restrictive set of model organisms.
Although the process of limb regeneration is different between invertebrate and vertebrate organisms, it …
Extreme Offspring Ornamentation In American Coots Is Favored By Selection Within Families, Not Benefits To Conspecific Brood Parasites, Bruce E. Lyon, Daizaburo Shizuka
Extreme Offspring Ornamentation In American Coots Is Favored By Selection Within Families, Not Benefits To Conspecific Brood Parasites, Bruce E. Lyon, Daizaburo Shizuka
School of Biological Sciences: Faculty Publications
Offspring ornamentation typically occurs in taxa with parental care, suggesting that selection arising from social interactions between parents and offspring may underlie signal evolution. American coot babies are among the most ornamented offspring found in nature, sporting vividly orange-red natal plumage, a bright red beak, and other red parts around the face and pate. Previous plumage manipulation experiments showed that ornamented plumage is favored by strong parental choice for chicks with more extreme ornamentation but left unresolved the question as to why parents show the preference. Here we explore natural patterns of variation in coot chick plumage color, both within …
The Influence Of Breeding Density On Female Aggression, Parental Care, And Ornamentation In A Secondary-Cavity Nesting Warbler, Elsa B. Chen
The Influence Of Breeding Density On Female Aggression, Parental Care, And Ornamentation In A Secondary-Cavity Nesting Warbler, Elsa B. Chen
Theses and Dissertations
Tradeoffs between individual survival and reproductive success associated with aggressive behaviors are a driving force of evolution, but these tradeoffs are often overlooked for aggressive conspecific interactions between females. For avian males, it is well documented that more aggressive individuals tend to provide less parental care. In the few studies that address this in females, the tradeoffs seem to be more context-dependent, varying due to factors such as predation pressure and habitat quality. The relationship between female ornamentation and aggression is similarly understudied, but evidence suggests that both aggression and ornamentation are important traits involved in social selection – the …
Evaluating The Effect Of Time Of Day On Singing Behavior In Anna’S Hummingbirds, Adrian D. Macedo, Maxine R. Mota
Evaluating The Effect Of Time Of Day On Singing Behavior In Anna’S Hummingbirds, Adrian D. Macedo, Maxine R. Mota
IdeaFest: Interdisciplinary Journal of Creative Works and Research from Cal Poly Humboldt
No abstract provided.
The Evolution Of Reproductive Complexity In Fishes, Frieda Benun
The Evolution Of Reproductive Complexity In Fishes, Frieda Benun
Dissertations, Theses, and Capstone Projects
How does reproductive complexity evolve? In this dissertation, I investigate the evolution of parental investment and mating behavior, using both macro- and microevolutionary lenses. I use fishes, the most diverse group of vertebrates, comprising over 30,000 species, as a model to study the evolution of these traits.
In Chapter 1, I introduce the evolution of parental behaviors in fishes. I present a systematic review of parental care for 294 families (close to 60%) of bony fish and show that male-only care is the predominant form of care in this group. I summarize current theories on parental care, emphasizing external fertilization, …
Traffic Noise And Sexual Selection: Studies Of Anthropogenic Impact On Bird Songs And Undergraduate Student Reasoning Of Evolutionary Mechanisms, Sarah Spier
School of Natural Resources: Dissertations, Theses, and Student Research
Humans have transformed much of the natural landscape and are continuing to do so at an accelerated rate, compromising natural areas that serve as important habitat for many species. Roads impact much of the environment as they fragment habitat and introduce traffic noise into the acoustic environment, deferentially affecting wildlife in roadside habitat. I explored how traffic noise affects the detection of birds based on whether their vocalizations were masked by traffic noise. Masked species detection was not affected by an increase in traffic noise amplitude, while there was a negative effect of traffic noise amplitude on unmasked species detection, …
Ecology, Monogamy, And The Evolution Of Animal Families, Jacqueline Rae Dillard
Ecology, Monogamy, And The Evolution Of Animal Families, Jacqueline Rae Dillard
Theses and Dissertations--Biology
Family-living has been recognized as a necessary prerequisite for the evolution of advanced cooperative societies, yet the evolutionary and ecological processes that drive the coupling of different forms of cooperation in family-based societies are still poorly understood. In my dissertation, I investigate the correlated evolution of parental care, monogamy, and cooperative breeding in a variety of family-based taxa. I explore the mating and social behavior of family-living beetles with incipient cooperation to better understand the factors driving these social traits. Specifically, I evaluate different causes of extra-pair mating in socially monogamous beetles, the potential benefits that young adult offspring may …
Multiple Connections Between Amazonia And Atlantic Forest Shaped The T Phylogenetic And Morphological Diversity Of Chiasmocleis Mehely, 1904 (Anura: Microhylidae: Gastrophryninae), Rafael O. De Sá, João Filipe Riva Tonini, Hannah Van Huss, Alex Long, Travis Cuddy, Mauricio C. Forlani, Pedro L.V. Peloso, Hussam Zaher, Célio F.B. Haddad
Multiple Connections Between Amazonia And Atlantic Forest Shaped The T Phylogenetic And Morphological Diversity Of Chiasmocleis Mehely, 1904 (Anura: Microhylidae: Gastrophryninae), Rafael O. De Sá, João Filipe Riva Tonini, Hannah Van Huss, Alex Long, Travis Cuddy, Mauricio C. Forlani, Pedro L.V. Peloso, Hussam Zaher, Célio F.B. Haddad
Biology Faculty Publications
Chiasmocleis is the most species-rich genus of Neotropical microhylids. Herein, we provide the first comprehensive multilocus phylogeny for the genus, including all but 3 of the 34 recognized species and multiple individuals per species. We discuss cryptic speciation, species discovery, patterns of morphological evolution, and provide a historical biogeographic analysis to account for the current distribution of the genus. Diversification of Chiasmocleis from other New World microhylids began during the Eocene, app. 40 mya, in forested areas, and current diversity seems to be a product of recurrent connections between the Atlantic Forest and Amazonia. Small-sized species evolved independently three times …
Happy Or Hangry Families: Does Kinship Mediate Cooperation And Cannibalism In Drosophila Melanogaster Larvae?, Lucas Khodaei
Happy Or Hangry Families: Does Kinship Mediate Cooperation And Cannibalism In Drosophila Melanogaster Larvae?, Lucas Khodaei
Theses and Dissertations (Comprehensive)
The fruit fly Drosophila melanogaster has a long history as a research model for studies on behaviour and group dynamics. When individuals are grouped in an environment and resources decrease, then they may behave co-operatively with one another or antagonistically compete. Hamilton’s Law states that if the benefit of helping a related individual out-weighs the cost to its direct fitness, then the individual’s indirect fitness will increase when they help their kin compared to helping an unrelated stranger. Yet it is unknown whether kinship mediates larval behaviour to encourage co-operation and/or lessen antagonistic behaviours between consanguineous individuals. In this thesis …
Shaping The Tools Of Battle : Sexual Selection, Contest Dynamics, And Weapon Divergence In The Asian Rhinoceros Beetle Trypoxylus Dichotomus, Jillian F. Del Sol
Shaping The Tools Of Battle : Sexual Selection, Contest Dynamics, And Weapon Divergence In The Asian Rhinoceros Beetle Trypoxylus Dichotomus, Jillian F. Del Sol
Graduate Student Theses, Dissertations, & Professional Papers
Exaggerated weapons of sexual selection often diverge more rapidly and dramatically than other body parts, suggesting that relevant agents of selection may be discernible in contemporary populations. I examined the ecology, reproductive behavior, and strength of sexual selection on horn length in five recently diverged beetle populations that differ in relative horn size. I show that mating system ecology differs between these locations and corresponds with the local strength of contemporary selection on horn length. Comparisons of ecological conditions and selection strength across populations offer a critical first step towards meaningfully linking mating system dynamics, selection patterns, and diversity in …
Courtship Behavior, Communication, And Copulation In Tigrosa Annexa, Samuel White
Courtship Behavior, Communication, And Copulation In Tigrosa Annexa, Samuel White
Scholars Week
The evolution of multimodal communication, where signalers use multiple signal components in multiple sensory modalities, has become the subject of investigation by many researchers. Signaling puts males at risk of predation, so why do males of some species evolve extra signals that may increase this risk? In some wolf spider species, males incorporate many visual and vibrational signals into a display that they use to attract a female for mating. Female spiders are often aggressive toward courting males and so the male display also functions to decrease the odds of cannibalism. Female wandering spiders deposit silk containing pheromones that communicate …
Review Of The Family Rivulidae (Cyprinodontiformes, Aplocheiloidei) And A Molecular And Morphological Phylogeny Of The Annual Fish Genus Austrolebias Costa 1998, Marcelo Loureiro, Rafael O. De Sá, Sebastián W. Serra, Felipe Alonso, Luis Esteban Krause Lanés, Matheus Vieira Volcan, Pablo Calviño, Dalton Nielsen, Alejandro Duarte, Graciela Garcia
Review Of The Family Rivulidae (Cyprinodontiformes, Aplocheiloidei) And A Molecular And Morphological Phylogeny Of The Annual Fish Genus Austrolebias Costa 1998, Marcelo Loureiro, Rafael O. De Sá, Sebastián W. Serra, Felipe Alonso, Luis Esteban Krause Lanés, Matheus Vieira Volcan, Pablo Calviño, Dalton Nielsen, Alejandro Duarte, Graciela Garcia
Biology Faculty Publications
The family Rivulidae is the fourth most diverse clade of Neotropical fishes. Together with some genera of the related African family Nothobranchiidae, many rivulids exhibit a characteristic annual life cycle, with diapausing eggs and delayed embryonic development, which allows them to survive in the challenging seasonal ponds that they inhabit. Rivulidae also includes two species known as the only the self-fertilizing vertebrates and some species with internal fertilization. The first goal of this article is to review the systematics of the family considering phylogenetic relationships and synapomorphies of subfamilial clades, thus unifying information that is dispersed throughout the literature. From …
Transcriptomics Of Learning, Pablo Iturralde
Transcriptomics Of Learning, Pablo Iturralde
Theses
Learning is a basic and important component of behavior yet we have very little empirical information about the interaction between mechanisms of learning and evolution. In our work, we are testing hypotheses about the neurogenetic mechanisms through which animal learning abilities evolve. We are able to test this directly by using experimentally evolved populations of flies, which differ in learning ability. These populations were previously evolved within the lab by creating worlds with different patterns of change following theoretically predicted effects on which enhanced learning will evolve. How has evolution acted to modulate genes and gene expression in the brain …
Radical Social Ecology As Deep Pragmatism: A Call To The Abolition Of Systemic Dissonance And The Minimization Of Entropic Chaos, Arielle Brender
Radical Social Ecology As Deep Pragmatism: A Call To The Abolition Of Systemic Dissonance And The Minimization Of Entropic Chaos, Arielle Brender
Student Theses 2015-Present
This paper aims to shed light on the dissonance caused by the superimposition of Dominant Human Systems on Natural Systems. I highlight the synthetic nature of Dominant Human Systems as egoic and linguistic phenomenon manufactured by a mere portion of the human population, which renders them inherently oppressive unto peoples and landscapes whose wisdom were barred from the design process. In pursuing a radical pragmatic approach to mending the simultaneous oppression and destruction of the human being and the earth, I highlight the necessity of minimizing entropic chaos caused by excess energy expenditure, an essential feature of systems that aim …
Diel Rhythmicity Found In Behavior But Not Biogenic Amine Levels In The Funnel-Web Spider Agelenopsis Pennsylvanica (Araneae, Agelenidae), Alexander E. Demarco
Diel Rhythmicity Found In Behavior But Not Biogenic Amine Levels In The Funnel-Web Spider Agelenopsis Pennsylvanica (Araneae, Agelenidae), Alexander E. Demarco
Electronic Theses and Dissertations
Quantifying individual differences in behavior and the extent that behavior is influenced by circadian control is of paramount importance in behavioral ecology. In addition, the proximate mechanisms underlying behavior are also critical in order to obtain a more complete picture of how behavior evolves. Biogenic amines (BAs) are simple nitrogenous compounds derived from amino acids and have been consistently and extensively linked to behavior. For this study, we analyzed temporal patterns of BAs in relation to the antipredator (boldness) and aggressive behavior in female Agelenopsis pennsylvanica, a funnel-web spider. Using HPLC-ED, we compared behavioral responses to temporal patterns of …
Social Contact Patterns Can Buffer Costs Of Forgetting In The Evolution Of Cooperation, Jeffrey R. Stevens, Jan K. Woike, Lael J. Schooler, Stefan Lindner, Thorsten Pachur
Social Contact Patterns Can Buffer Costs Of Forgetting In The Evolution Of Cooperation, Jeffrey R. Stevens, Jan K. Woike, Lael J. Schooler, Stefan Lindner, Thorsten Pachur
Jeffrey Stevens Publications
Analyses of the evolution of cooperation often rely on two simplifying assumptions: (i) individuals interact equally frequently with all social network members and (ii) they accurately remember each partner's past cooperation or defection. Here, we examine how more realistic, skewed patterns of contact—in which individuals interact primarily with only a subset of their network's members—influence cooperation. In addition, we test whether skewed contact patterns can counteract the decrease in cooperation caused by memory errors (i.e. forgetting). Finally, we compare two types of memory error that vary in whether forgotten interactions are replaced with random actions or with actions from previous …
Mesotocin Influences Pinyon Jay Prosociality, Juan Duque, Whitney Leichner, Holly Ahmann, Jeffrey R. Stevens
Mesotocin Influences Pinyon Jay Prosociality, Juan Duque, Whitney Leichner, Holly Ahmann, Jeffrey R. Stevens
Jeffrey Stevens Publications
Many species exhibit prosocial behavior, in which one individual’s actions benefit another individual, often without an immediate benefit to itself. The neuropeptide oxytocin is an important hormonal mechanism influencing prosociality in mammals, but it is unclear whether the avian homologue mesotocin plays a similar functional role in birds. Here, we experimentally tested prosociality in pinyon jays (Gymnorhinus cyanocephalus), a highly social corvid species that spontaneously shares food with others. First, we measured prosocial preferences in a prosocial choice task with two different payoff distributions: Prosocial trials delivered food to both the subject and either an empty cage or a partner …
Ecological Uncertainty Influences Vigilance As A Marker Of Fear, Laurence E. A. Feyten, Grant E. Brown
Ecological Uncertainty Influences Vigilance As A Marker Of Fear, Laurence E. A. Feyten, Grant E. Brown
Animal Sentience
We expand on the factors that may shape the predictability of risk and the potential impacts on the links between vigilance and fear, primarily in aquatic prey communities. Uncertainty in predation risks has been shown to induce increased levels of neophobia among prey. As a result of this phenotypically plastic response, prey are faced with risk assessment cues that may vary widely in their reliability. We argue that decomposing predictability may provide useful insights into the relationship between vigilance and fear.
Competing Activities As Measures Of Fear And Vigilance, Ferenc Mónus
Competing Activities As Measures Of Fear And Vigilance, Ferenc Mónus
Animal Sentience
In animal behavioural research on vigilance, visual signs of alertness are usually used to estimate perceived risk (an internal “fear” state) of free-ranging animals. Different measures of vigilance and competing activities (e.g., predator vigilance, conspecific vigilance, feeding, food handling) provide clues for better understanding vigilance behaviour. How efficiently does an animal in a vigilant/non-vigilant posture devote attention to threats or invest in other activities, such as searching for or handling food? Several species regularly withdraw to a sheltered spot when feeding in an abundant food patch, spending short periods in complete safety. Frequencies of feeding interruptions or false-alarm flights provide …
The North American Quails, Partridges, And Pheasants, Paul A. Johnsgard
The North American Quails, Partridges, And Pheasants, Paul A. Johnsgard
Zea E-Books Collection
This book documents the biology of six species of New World quails that are native to North America north of Mexico (mountain, scaled, Gambel’s, California, and Montezuma quails, and the northern bobwhite), three introduced Old World partridges (chukar, Himalayan snowcock, and gray partridge), and the introduced common (ring-necked) pheasant. Collectively, quails, partridges, and pheasants range throughout all of the continental United States and the Canadian provinces. Two of the species, the northern bobwhite and ring-necked pheasant, are the most economically important of all North American upland game birds. All of the species are hunted extensively for sport and are highly …
Is Cooperative Memory Special? The Role Of Costly Errors, Context, And Social Network Size When Remembering Cooperative Actions, Jeffrey R. Stevens, Tim Winke
Is Cooperative Memory Special? The Role Of Costly Errors, Context, And Social Network Size When Remembering Cooperative Actions, Jeffrey R. Stevens, Tim Winke
Jeffrey Stevens Publications
Theoretical studies of cooperative behavior have focused on decision strategies, such as tit-for-tat, that depend on remembering a partner’s last choices. Yet, an empirical study by Stevens et al. (2011) demonstrated that human memory may not meet the requirements that needed to use these strategies. When asked to recall the previous behavior of simulated partners in a cooperative memory task, participants performed poorly, making errors in 10–24% of the trials. However, we do not know the extent to which this task taps specialized cognition for cooperation. It may be possible to engage participants in more cooperative, strategic thinking, which may …
The Altruistic Self, Nathan Dougherty
The Altruistic Self, Nathan Dougherty
Dialogue & Nexus
Altruism as a purely naturalistic phenomenon self-defeats the term altogether; however, theology also makes unsubstantiated claims that some behaviors are purely selfless. I will first define various conceptual forms of altruism and then offer explanations of the term from neurological, evolutionary and psychological investigations. Despite the position that altruism can be reduced to a fantastical impossibility bearing neither the arms of science nor theology, it is also a fallacy to separate it from a religiously derived supernatural altruism that carries no implications for the realm of morality.
A New Taxonomy Of Altruism In Terms Of Prosocial Behaviors, Kristin Kaiser
A New Taxonomy Of Altruism In Terms Of Prosocial Behaviors, Kristin Kaiser
Dialogue & Nexus
The definition of altruism has been studied, explained, and even confused by many scholars in various fields. The term itself has been inappropriately used to describe prosocial behaviors that do not fall within the definition of altruism. An evaluation of Grant Ramsey’s taxonomy of altruism, which includes biological altruism, psychological altruism, and helping altruism, proves that it is not adequate in categorizing organism’s behaviors. A new taxonomy, with the branches of kin selection, reciprocity, and aesthetic altruism, is presented and explained to clarify the definition of altruism and alleviate confusion about how to describe prosocial behaviors. Both naming systems are …