Ecology and Evolutionary Biology Commons^™

Introduction, L. David Mech, Luigi Boitani

United States Geological Survey, Northern Prairie Wildlife Research Center: Publications

THE WOLF IS TRULY a special animal. As the most widely distributed of all land mammals, the wolf, formally the gray wolf (Canis lupus), is also one of the most adaptable. It inhabits all the vegetation types of the Northern Hemisphere and preys on all the large mammals living there. It also feeds on all the other animals in its environment, scavenges, and can even eat fruits and berries. Wolves frequent forests and prairies, tundra, barren ground, mountains, deserts, and swamps. Some wolves even visit large cities, and, of course, the wolf's domesticated version, the dog, thrives in …

Wolf Population Dynamics, Todd K. Fuller, L. David Mech, Jean Fitts Cochrane

United States Geological Survey, Northern Prairie Wildlife Research Center: Publications

A LARGE, DARK WOLF poked his nose out of the pines in Yellowstone National Park as he thrust a broad foot deep into the snow and plowed ahead. Soon a second animal appeared, then another, and a fourth. A few minutes later, a pack of thirteen lanky wolves had filed out of the pines and onto the open hillside.

Wolf packs are the main social units of a wolf population. As numbers of wolves in packs change, so too, then, does the wolf population (Rausch 1967). Trying to understand the factors and mechanisms that affect these changes is what the …

Wolves: Behavior, Ecology, And Conservation: Photo Section Ii, Douglas W. Smith, L. David Mech, Isaac Babcock, Melissa Mccaw

United States Geological Survey, Northern Prairie Wildlife Research Center: Publications

Adult wolves are very attentive to the pups. Both parents feed and care for them. Any older siblings similarly participate in pup care and feeding. Kin selection is probably the best explanation for the latter behavior. Top: Photograph by Isaac Babcock. Bottom: Photograph by L. David Mech.

Nontarget Effects—The Achilles’ Heel Of Biological Control? Retrospective Analyses To Reduce Risk Associated With Biocontrol Introductions, Svata M. Louda, R. W. Pemberton, M. T. Johnson, P. A. Follett

Papers in Ecology

Controversy exists over ecological risks in classical biological control. We reviewed 10 projects with quantitative data on nontarget effects. Ten patterns emerged: (a) Relatives of the pest are most likely to be attacked; (b) host-specificity testing defines physiological host range, but not ecological range; (c) prediction of ecological consequences requires population data; (d) level of impact varied, often in relation to environmental conditions; (e) information on magnitude of nontarget impact is sparse; ( f ) attack on rare native species can accelerate their decline; (g) nontarget effects can be indirect; (h) agents disperse from agroecosystems; (i) whole assemblages of species …

Wolves: Behavior, Ecology, And Conservation: Photo Section I, Douglas W. Smith, Rolf Peterson, L. David Mech, Barron Crawford

United States Geological Survey, Northern Prairie Wildlife Research Center: Publications

Wolves in Yellowstone National Park's restored population often must compete with grizzly bears at the carcasses of their kills. Especially after their first feeding at a kill, wolves tend to relinquish their kills to bears. Photograph by Doug Smith.

Elk are one of the most common prey of wolves in the northern Rocky Mountains of North America, primarily because in most areas they tend to outnumber other prey. This large bull standing his ground escaped this pack. Photograph by Doug Smith.

Incidence Of Mink, Mustela Vison, And River Otter, Lutra Canadensis, In A Highly Urbanized Area, L. David Mech

United States Geological Survey, Northern Prairie Wildlife Research Center: Publications

Mink (Mustela vison) frequently inhabited or traversed a residential. business. and industrial part of the Twin Cities. Minnesota, with little water or natural vegetation. At least one River Otter (Lutra canadensis) also resided on a small pond on a golf course in the area for several winter months.

Wolf Social Ecology, L. David Mech, Luigi Boitani

United States Geological Survey, Northern Prairie Wildlife Research Center: Publications

THE FIRST REAL BEGINNING to our understanding of wolf social ecology came from wolf 2204 on 23 May 1972. State depredation control trapper Lawrence Waino, of Duluth, Minnesota, had caught this female wolf 112 km ( 67 mi) south of where L. D. Mech had radio-collared her in the Superior National Forest 2 years earlier. A young lone wolf, nomadic over 100 km² (40 mi²) during the 9 months Mech had been able to keep track of her, she had then disappeared until Waino caught her. From her nipples it was apparent that she had just been …

Wolves And Humans, Steven H. Fritts, Robert O. Stephenson, Robert D. Hayes, Luigi Boitani

United States Geological Survey, Northern Prairie Wildlife Research Center: Publications

TRY TO IMAGINE a small group of wolves sitting at a table engaged in vigorous debate. These wolves are from various parts of the globe and are perhaps a bit more scholarly than most. In fact, they are especially knowledgeable about the biology of that notorious two-legged species, Homo sapiens. They have been brought together to document their relationship with humans over the last several millennia. Pause for a few moments and consider what they might say ...

Perhaps the wolves' discussion would chronicle the evils of the human species, including details of atrocities committed against lupine ancestors down …

Wolf Interactions With Non-Prey, Warren B. Ballard, Ludwig N. Carbyn, Douglas W. Smith

United States Geological Survey, Northern Prairie Wildlife Research Center: Publications

WOLVES SHARE THEIR ENVIRONMENT with many animals besides those that they prey on, and the nature of the interactions between wolves and these other creatures varies considerably. Some of these sympatric animals are fellow canids such as foxes, coyotes, and jackals. Others are large carnivores such as bears and cougars. In addition, ravens, eagles, wolverines, and a host of other birds and mammals interact with wolves, if only by feeding on the remains of their kills.

Conclusion, L. David Mech, Luigi Boitani

United States Geological Survey, Northern Prairie Wildlife Research Center: Publications

WOLVES CAN LIVE almost anywhere in the Northern Hemisphere, and almost everywhere they do, they are an issue. In the vast emptiness of the northern tundra or the Arabian desert, on the outskirts of a European town or in the safety of an American national park, in meager agricultural lands in India or mountains in rich Norway or Switzerland, wolves always attract people's attention. Wolves form a key part of many ecosystems, and they are considered charismatic creatures by most human cultures. Thus they polarize public opinion and make headlines year after year.

If we look back 6o years to …

References, L. David Mech, Luigi Boitani

United States Geological Survey, Northern Prairie Wildlife Research Center: Publications

Abrams, P. A. 2000. The evolution of predator-prey interactions. Annu. Rev. Ecol. Syst. 31:79-105.

Abuladze, K. I. 1964. Osnovy Tsestodologii. Vol. IV. Teniatylentochnye gel' minty zhivotnykh i cheloveka i vyzyvaevaniia. Nauka, Moscow. 530 pp.

Achuff, P. L., and R. Petocz. 1988. Preliminary resource inventory of the Arjin Mountains Nature Reserve, Xinjiang, People's Republic of China. World Wide Fund for Nature, Gland, Switzerland. 78 pp.

Ackerman, B. B., F. A. Leban, M. D. Samuel, and E. 0. Garton. 1990. User's manual for program Home Range. 2d ed. Technical Report no. 15. Forestry, Wildlife, and Range Experiment Station, University ofldaho, Moscow.

Acorn, …

Survival Of The Western Pond Turtle (Emys Marmorata) In An Urban California Environment, Phillip Q. Spinks, Gregory B. Pauly, John J. Crayon, H. Bradley Shaffer

United States Geological Survey: Staff Publications

The western pond turtle Emys (formerly Clemmys marmorata) is declining throughout its range, primarily due to loss of habitat via urbanization and conversion to agriculture. Urban waterways present several important challenges to freshwater turtle populations, but they also present an opportunity to maintain declining species in a ubiquitous habitat that has high public visibility. The arboretum waterway on the University of California, Davis campus is an example of an extensively altered urban habitat that supports a relatively large E. marmorata population. Over the last 6 years, we monitored the turtle population inhabiting the arboretum waterway to determine the …

Bats Of Nevis, Northern Lesser Antilles, Scott C. Pedersen, Hugh H. Genoways, Matthew N. Morton, James W. Johnson, Siân E. Courts

University of Nebraska State Museum: Mammalogy Papers

Only one species of bat, Molossus molossus, previously has been documented as occurring on the northern Lesser Antillean island of Nevis. Field research and reviews of existing museum collections have provided documentation based on voucher specimens for an additional seven species occurring on the island — Noctilio leporinus, Brachyphylla cavernarum, Monophyllus plethodon, Ardops nichollsi, Artibeus jamaicensis, Natalus stramineus, and Tadarida brasiliensis. The biological diversity of the chiropteran fauna on Nevis is similar to that found on other islands in the northern Lesser Antilles. Ecologically, this is a simple chiropteran fauna, including one …

Taxonomy And Distribution Of Butterflies (Papilionoidea) Of The Skardu Region, Pakistan, Muhammad Abbas, Muhammad Ather Rafi, Mian Inayatullah, Muhammad Rafique Khan, Harry Pavulaan

The Taxonomic Report of the International Lepidoptera Survey

A study was conducted in six different localities around Skardu, Pakistan to document the butterfly fauna of that region. The study revealed that 16 species in 5 families and 14 genera occur in the area. The families include: Papilionidae (represented only by the genus Parnassius); Pieridae (genera Pieris, Pontia, and Colias are represented); Lycaenidae (genera Lycaena, Everes, Aricia, Plebejus, Zizeeria, and Zizina are represented); Nymphalidae (only two genera, Aglais and Cynthia are represented); and Satyridae (represented by the genera Pararge and Maniola).

The Prairie Naturalist Volume 34, No. 3/4 September/December 2002

The Prairie Naturalist

SEASONAL FOOD HABITS OF COYOTES IN NORTHEASTERN KANSAS ▪ J. F. Kamler, P. S. Gipson, and C. C. Perchellet

SELECTION OF NESTING HABITAT BY SHARP-TAILED GROUSE IN THE NEBRASKA SANDHILLS ▪ B. L. Prose, B. S. Cade, and D. Hein

PATTERNS OF REPRODUCTION FOR WESTERN HARVEST MICE IN NORTH-CENTRAL KANSAS ▪ J. R. Goheen, G. A. Kaufman, and D. W. Kaufman

THE IMPACT OF RECREATIONAL TRAILS AND GRAZING ON SMALL MAMMALS IN THE COLORADO PIEDMONT ▪ C. A. Meaney, A. K. Ruggles, N. W. Clippinger, and B. C. Lubow

CHRISTMAS BIRD COUNTS FOR NORTH DAKOTA 2001 ▪ R. N. Randall …

A Review Of Poanes Hobomok (Hesperiidae: Hesperiinae), With The Description Of A New Subspecies From The Southern Appalachians, Ronald R. Gatrelle

The Taxonomic Report of the International Lepidoptera Survey

Poanes hobomok monofacies is described from the southern limits of the Appalachian Mountains in western North Carolina, northern Georgia and eastern Tennessee, United States. The type locality is Chestnut Mountain, 3,400’, Macon County, North Carolina, United States. This new subspecies is characterized by the similarity of males and females and overall much darker and subdued marking when compared to both P. h. hobomok (eastern to mid-, northern United States and eastern to mid-, southern Canada) and P. h. wetona (Colorado and New Mexico, United States). The northern and western range of this new taxon and the …

How Many Black-Tailed Prairie Dogs Were There? A Dialogue Demonstrates Both The Process Of Science And The Role Of Science In Environmental Issues, Svata M. Louda

Svata M. Louda Publications

In this Dialogue, Dallas Virchow and Scott Hygnstrom challenge the present, widely held view that the numbers of black-tailed prairie dogs have declined significantly in post-settlement times. Using quotes and numbers from historical accounts of the occurrence of the prairie dog by the early explorers and settlers, they build a case that occurrence and abundance of the prairie dog in the shortgrass prairies and midgrass regions to the west of the tallgrass region in the Great Plains cannot be used to estimate the presettlement abundance of the species in tallgrass areas. They argue that the activities of early settlers, including …

Pterourus Appalachiensis (Papilionidae: Papilioninae), A New Swallowtail Butterfly From The Appalachian Region Of The United States, Harry Pavulaan, David M. Wright

The Taxonomic Report of the International Lepidoptera Survey

A new univoltine species of Tiger Swallowtail, Pterourus appalachiensis, is described from the southern Appalachian Mountain region of the eastern United States. This distinct swallowtail has remained unrecognized by lepidopterists since the description of its sympatric congener Pterourus glaucus (Linnaeus) in 1758. Historical accounts of Tiger Swallowtails from this region pertain specifically to glaucus and cannot be attributed to this new species. Morphology, voltinism, phenology, distribution, and behavioral traits indicate species level status of appalachiensis. Although appalachiensis shares several phenotypic characters with the recently elevated species canadensis (Rothschild & Jordan), preliminary mitochondrial DNA analysis indicates that appalachiensis is …

The Prairie Naturalist Volume 34, No. 1/2 March/June 2002

The Prairie Naturalist

LOCALIZED MOVEMENTS AND SITE FIDELITY OF WHITETAILED DEER IN THE NORTHERN GREAT PLAINS ▪ B. J. Kernohan, J. A. Jenks, and D. E. Naugle

SPAWNING-SEASON HOMING OF COMMON CARP AND RIVER CARPSUCKER ▪ J. L. Bonneau and D. L. Scamecchia

HOME RANGES OF RING-NECKED PHEASANTS IN NORTHWESTERN KANSAS ▪ R. D. Applegate, B. E. Flock, P. S. Gipson, M. W. McCoy, and K. E. Kemp

RESPONSES OF BIGHORN SHEEP TO DISTURBANCE IN LOW-ELEVATION GRASSLANDS ▪ R. W. Sayre, R. W. Seabloom, and W. F. Jensen

DISTRIBUTION OF CHANNEL CATFISH LIFE STAGES IN A PRAIRIE RIVER BASIN ▪ . J. C. …

Unexpected Ecological Effects Of Distributing The Exotic Weevil, Larinus Planus (F.), For The Biological Control Of Canada Thistle, Svata M. Louda, Charles W. O'Brien

Svata M. Louda Publications

Rhinocyllus conicus, a weevil introduced for biological control of exotic weeds, has had major nontarget ecological effects on native thistles. Some practitioners have argued that this is an isolated case. We report, however, that another Eurasian weevil (Larinus planus), currently being distributed in North America for the control of Canada thistle (Cirsium arvense), is significantly reducing seed production by a native thistle (Tracy’s thistle, Cirsium undulatum var. tracyi) in Colorado. In 1999 we discovered L. planus feeding in flower heads of Tracy’s thistle near a 1992–1993 biocontrol release site. Of the heads collected, 74% …

Name-Bearing Types And Taxonomic Synopsis Of Three Lycaenid Butterfly Taxa From Western Canada (Lepidoptera: Lycaenidae), Norbert G. Kondla, Crispin S. Guppy

The Taxonomic Report of the International Lepidoptera Survey

We clarify which of two designated lectotypes of Chrysophanus florus Edwards, 1884 is valid. We also show that the putative holotype of Plebeius saepiolus insulanus Blackmore, 1920 is actually a lectotype. A valid neotype designation for Lycaena saepiolus amica Edwards, 1863 is provided. Publication dates are corrected and we also briefly review the various taxonomic interpretations and distributions that appear in the literature with respect to these taxa.

Ecology Of The Missouri River. Progress Report, Dingell-Johnson Project F-75-R-19, Supplement 1- Missouri River Creel Survey, Camp Creek To Kansas State Line, 31 March Through 12 October 2001, Gerald Mestl

Nebraska Game and Parks Commission: White Papers, Conference Presentations, and Manuscripts

This report describes Missouri River activities and results related to a channelized Missouri River creel survey conducted from 31 March through 12 October 2001. This is the second of a planned annual creel survey to be conducted on alternating sections of the channelized Missouri River to measure changes in recreational fishing activity, especially those changes due to large scale habitat restoration efforts. Future reports will contain additional analyses of these data. Anglers spent over 22,000 hours fishing the Missouri River from Camp Creek (rkm 883.5) to the Kansas state line (rkm 790.2) during the survey period. Effort peaked during the …

Visual Predators Select For Crypticity And Polymorphism In Virtual Prey, Alan B. Bond, Alan Kamil

Papers in Behavior in Biological Sciences

Cryptically colored animals commonly occur in several distinct pattern variants. Such phenotypic diversity may be promoted by frequency-dependent predation, in which more abundant variants are attacked disproportionately often, but the hypothesis has never been explicitly tested. Here we report the first controlled experiment on the effects of visual predators on prey crypticity and phenotypic variance, in which blue jays (Cyanocitta cristata) searched for digital moths on computer monitors. Moth phenotypes evolved via a genetic algorithm in which individuals detected by the jays were much less likely to reproduce. Jays often failed to detect atypical cryptic moths, confirming frequency- …

Distribution And Prevalence Of Echinococcus Multilocularis In Wild Predators In Nebraska, Kansas, And Wyoming, S. T. Stroandt, D. R. Virchow, M. W. Dryden, Scott E. Hygnstrom, K. R. Kazacos

School of Natural Resources: Faculty Publications

To further determine the distribution and prevalence of Echinococcus multilocularis in the central United States, 245 wild canids (125 red foxes, 120 coyotes) and 33 bobcats were collected from Nebraska, Kansas, and Wyoming and examined for this parasite. Animals examined included 11 red foxes from the western panhandle of Nebraska; 5 red foxes and 30 coyotes from southern Nebraska; 56 red foxes and 1 coyote from northeastern Nebraska; 20 red foxes, 63 coyotes, and 13 bobcats from northern Kansas; 2 red foxes, 26 coyotes, and 20 bobcats from southern Kansas; and 31 red foxes from eastcentral Wyoming. Of these, 27 …

Diel And Seasonal Habitat Use By Red Shiner (Cyprinella Lutrensis), Shyi-Liang Yu, Edward J. Peters

School of Natural Resources: Faculty Publications

Fish distribution and microhabitat use were quantified over a 24-h period 15 times from July 1988 to October 1990 at a site in the lower Platte River, Nebraska. The objectives of this study were to determine the effects of availability on habitat use by the red shiner (Cyprinella lutrensis) and to describe variations in red shiner use of depth, velocity, cover, and substrate by seasonal and diel periods. Day-night collections showed a difference in selection or avoidance of depths and substrates in summer. Red shiners selected depths of less than 30 cm during the night and fine substrate during the …

Effects Of Management Practices On Grassland Birds: Horned Lark, Meghan F. Dinkins, Amy L. Zimmerman, Jill A. Dechant, Barry D. Parkin, Douglas H. Johnson, Lawrence D. Igl, Christopher M. Goldade, Betty R. Euliss

United States Geological Survey: Effects of Management Practices on Grassland Birds

Information on the habitat requirements and effects of habitat management on grassland birds were summarized from information in more than 4,000 published and unpublished papers. A range map is provided to indicate the relative densities of the species in North America, based on Breeding Bird Survey (BBS) data. Although birds frequently are observed outside the breeding range indicated, the maps are intended to show areas where managers might concentrate their attention. It may be ineffectual to manage habitat at a site for a species that rarely occurs in an area. The species account begins with a brief capsule statement, which …

Cognition As An Independent Variable: Virtual Ecology, Alan C. Kamil, Alan B. Bond

Papers in Behavior in Biological Sciences

On close examination, human cultural artifacts bear the unmistakable impress of the structure of the human mind: our tools, habitations. and methods of communication have been molded to suit the strengths and limitations of the human cognitive system (Nosman 1988). It has not commonly been emphasized. however. that similar shaping processes have taken place over the course of biological evolution in response to the cognitive features of other, nonhuman species (Bonner 1980; von Frisch 1974).
Cognitive influences are particularly evident in the modifications of color patterns and behavior of prey species that take advantage of biases and constraints in the …

A Comparative Study Of Geometric Rule Learning By Nutcrackers (Nucifraga Columbiana), Pigeons (Columba Livia), And Jackdaws (Corvus Monedula ), Juli E. Jones, Elena Antoniadis, Sara J. Shettleworth, Alan Kamil

Papers in Behavior in Biological Sciences

Three avian species, a seed-caching corvid (Clark’s nutcrackers; Nucifraga columbiana), a non-seed-caching corvid (jackdaws; Corvus monedula), and a non-seed-caching columbid (pigeons; Columba livia), were tested for ability to learn to find a goal halfway between 2 landmarks when distance between the landmarks varied during training. All 3 species learned, but jackdaws took much longer than either pigeons or nutcrackers. The nutcrackers searched more accurately than either pigeons or jackdaws. Both nutcrackers and pigeons showed good transfer to novel landmark arrays in which interlandmark distances were novel, but inconclusive results were obtained from jackdaws. Species differences in this …

Relating The Unique Sensory System Of Amblypygids To The Ecology And Behavior Of Phrynus Parvulus From Costa Rica (Arachnida, Amblypygi), Eileen Hebets

Eileen Hebets Publications

Abstract: While the unique sensory system and neuroanatomy of amblypygids suggest that olfaction is important in their lives, to date no behavioral data exist to support this suggestion. To gain insight into amblypygid ecology and behavior, an individual mark–recapture study was conducted on the Costa Rican amblypygid Phrynus parvulus. Within two 50 by 25 m plots, the distribution and movement patterns of individual amblypygids were recorded for over 2 months. A total of 88 adult individuals (60 males and 28 females) were marked, with an average 51% resighted. Females were resighted more frequently than males (75% females, 40% males). …

Clone Size Of Andropogon Gerardii Vitman (Big Bluestem) At Konza Prairie, Kansas, Kathleen H. Keeler, Charles F. Williams, Linda S. Vescio

School of Biological Sciences: Faculty Publications

Clone size of plants of Andropogon gerardii from Konza Prairie Biological Station, Manhattan, Kansas was estimated from spatial patterns of genetic variation, using proteins detected by starch gel electrophoresis and DNA content (ploidy) measured by flow cytometry. Unique multi-locus protein banding patterns and differences in ploidy were used to exclude plants as members of the same clone. Individual clones averaged about 2 m in diameter and areas of prairie of 100 m² were calculated to contain an average of 31.8 genetic individuals.