Ecology and Evolutionary Biology Commons^™

The Ecology Of The Timber Rattlesnake (Crotalus Horridus) In Southeastern Virginia, Scott M. Goetz

Biological Sciences Theses & Dissertations

Few studies have examined the relationship between forest canopy structure and the ecology of the timber rattlesnake (Crotalus horridus). I used radiotelemetry to compare the movements, activity range sizes and behaviors of timber rattlesnakes before and after a large-scale natural disturbance that opened a previously closed canopy. The disturbance was Hurricane Isabel which made landfall in northeast North Carolina and southeastern Virginia in 2003. Isabel created gaps in the canopy through tree blowdown, resulting in a 16.6% opening in the forest canopy at my study site, in southeastern Virginia. I compared six years of female tracking data from …

A Review Of The Subgenus Pandinus Thorell, 1876 With Descriptions Of Two New Species From Uganda And Ethiopia (Scorpiones: Scorpionidae), František Kovařík

Euscorpius

Pandinus (Pandinus) ugandaensis sp. n. from Uganda and P. (P.) mazuchi sp. n. from Ethiopia are described and compared with other species of the subgenus. P. ugandaensis sp. n. is characterized by 3-4/3: 3-4/3: 4/3: 4/3 spiniform formula of tarsomere II and only two spines on the inclined anteroventral surface of tarsomere II; eight ventral trichobothria on the chela; 10–11 pectinal teeth in females and 13–14 in males; and 1.6–1.7 length to depth ratio of the fourth metasomal segment. P. (P.) mazuchi sp. n. is characterized by 7/4: 6-7/4: 6-7/5-6: 8/5 spiniform formula of tarsomere II and only …

Redescription Of Centruroides Granosus (Thorell, 1876) And Identity Of Centrurus Granosus Simplex Thorell, 1876 (Scorpiones: Buthidae), Luis F. De Armas, Rolando Teruel, František Kovařík

Euscorpius

Centruroides granosus (Thorell, 1876) is redescribed based on several specimens of both sexes from Panama. We demonstrate that the original description of this species was based on an immature (subadult) male, which is herein designated as the lectotype. We conclude that Centrurus granosus simplex Thorell, 1876 is a senior synonym of Centruroides testaceus arubensis (Bakker, 1963), and elevate it to species level: Centruroides simplex (Thorell, 1876), comb. nov.

Buthus Awashensis Sp. N. From Ethiopia (Scorpiones: Buthidae), František Kovařík

Euscorpius

Buthus awashensis sp. n. from the Awash region of Ethiopia, is described and compared with B. berberensis Pocock, 1900 from Somaliland. The new species is characterized chiefly by the total length of 50–65 mm. B. berberensis reaches only 45–55 mm and differs in coloration, with adults having the chela of pedipalp and chelicerae entirely yellow without dark reticulations, and by mophometric characters, mainly the shape of the chela of pedipalp. Pectinal marginal tips extend the to proximal end of the fourth sternite in males of B. awashensis sp. n. and to proximal end of the fifth sternite in males of …

Hemiscorpius Novaki Sp. N. From Somaliland (Scorpiones: Hemiscorpiidae), František Kovařík, Tomáš Mazuch

Euscorpius

Hemiscorpius novaki sp. n. from Somaliland is described and compared with other African species of the genus. The new species is characterized chiefly by the total length of 40–46 mm; yellow to yellowish-brown color with pedipalp fingers reddish brown to black, darker than the chela; slightly longer metasoma in the male than in the female. These characters distinguish H. novaki sp. n. from H. somalicus Lourenço, 2011. The third African species of the genus, H. tellinii Borelli, 1904, is known only from the female holotype, which differs from females of H. novaki sp. n. in having much less pronounced granulation …

Scorpions In The Modern Cuban Culture: An Introductory Iconography, Luis F. De Armas

Euscorpius

Some graphic examples of the role of scorpions in the present-day culture of Cuba are presented. They include tattoos, jewels, carpets, caps, and drawings by children, among other objects of personal or social usage. Increased use of this arachnid as a cultural element among Cuban people during last 50 years seems to be directly related with globalization, mostly through the TV and movies.

The First Record Of The Family Euscorpiidae (Arachnida: Scorpiones) From Central China, With A Key Of Chinese Species Of The Genus Scorpiops, Zhi-Yong Di, Ya-Wen He, Zhi-Jian Cao, Ying-Liang Wu, Wen-Xin Li

Euscorpius

The genus Scorpiops (Euscorpiidae) is recorded for the first time in Central China. Two immature specimens of a form belonging to Scorpiops hardwickii (Gervais, 1843) “complex” were collected from Huzhaoshan Mountains in Hubei Province. A discussion of Chinese species of genus Scorpiops is provided, as well as a key of Scorpiops from China.

Further Considerations On The Species Of The Genus Orthochirus Karsch, 1891 From Africa, With Description Of Three New Species (Scorpiones: Buthidae), Wilson R. Lourenço, Elise-Anne Leguin

Euscorpius

New considerations are proposed regarding the African species of the genus Orthochirus Karsch, 1891. Two species, Orthochirus aristidis (Simon, 1882) and Orthochirus innesi Simon, 1910 have been the subject of several publications in the past decades; however, doubts remain about their exact identity and range of geographical distribution. In this note, their taxonomic status is reinvestigated. The type material is revised and the lectotype and paralectotypes are designated for O. aristidis. Revised diagnoses and illustrations are proposed for both species, and these are confirmed as valid. Three new species are described from Algeria, Morocco, and Mauritania. The total number …

Redescription Of Tityus Pictus Pocock, 1893 And Tityus Smithii Pocock, 1893, With Notes On The Tityus Species From The Lesser Antilles (Scorpiones: Buthidae), Rolando Teruel

Euscorpius

In the present paper, the taxonomic status of Tityus pictus Pocock, 1893 and all of its junior synonyms is revised. Tityus smithii Pocock, 1893 is restored as a valid species, and the subspecies Tityus smithii microdon Pocock, 1893 (currently T. pictus microdon) is regarded as its junior synonym. Both T. pictus and T. smithii are redescribed, supplementary information on their morphological variability are given, and their geographical distribution is up- dated, including a new locality record for the latter. Also, the taxonomic position and relationships of all Lesser Antillean species of the genus are clarified: it is demonstrated that …

On Fabre’S Traces: An Important Contributor To The Knowledge Of Buthus Occitanus (Amoreux, 1789), Marco Colombo

Euscorpius

Jean-Henri Casimir Fabre (1823–1915) has probably been one of the most important entomologists of the world in the last two centuries, leaving to posterity a huge amount of manuscripts and books. The Languedoc yellow scorpion, Buthus occitanus (Amoreaux, 1789), has been one of his objects of study: many of its behaviours, including the famous “promenade a deux”, have been described by the careful eye of the French entomologist, giving an interesting contribution to modern scorpiology

Scorpions In Ancient Egypt, Hisham K. El-Hennawy

Euscorpius

The ancient Egyptians knew the scorpion and its toxicity, and venerated it since pre-dynastic era. They used the scorpion as a king's name, a name of a nome (county), and a symbol to their goddess, Serqet, that protects the body and the viscera of the dead, and that accompanies them in their journey to the afterlife. They had medical prescriptions and magical spells to heal the stings. Since the 5th dynasty, the title of a “Follower of Serket” was given to clever physicians. Scorpions are most famously depicted on Horus Cippus, a talisman featuring Horus the Child holding in his …

Pectine Development In Scorpion Embryos And First And Second Instars, Roger D. Farley

Euscorpius

The pectines are complex sensory organs that extend from the ventral surface of the anterior opisthosoma (mesosoma) in all extant scorpions and nearly all the fossil scorpions that have been examined. The pectines are synapomorphic for the Order Scorpiones. In this investigation, the scanning electron microscope (SEM) was used to study the development of the pectines in representatives from five scorpion families. In the more basal families (e.g., Vaejovidae) with apoikogenic development, the pectines start to develop early with enlargement of the limb buds on the third opisthosomal segment. The primordial pectines become elongate lobes attached to the ventro-lateral surface …

The First Record Of Upper Permian And Lower Triassic Scorpions From Russia (Chelicerata: Scorpiones), Victor Fet, Dmitry E. Shcherbakov, Michael E. Soleglad

Euscorpius

Several small fragments of fossil scorpions are reported from two localities in Vologda Province, Russia, representing the Upper Permian (Severodvinian, correlated to Wuchiapingian) (Isady) and Lower Triassic just above the Permian-Triassic boundary (Induan) (Nedubrovo). Most observed structures are not diagnostic at genus or family level. The Isady leg fragment possesses ungues (claws), which are both denticulated and setaceous, and resembles a Carboniferous Eobuthus sp. (Eobuthidae). It is the latest record of this type of ungues, which are known in some Paleozoic scorpions (extinct suborder Mesoscorpiones); all extant scorpions have smooth claws without denticulation or setation.

In Situ Infrared Videography Of Sand Scorpion Nighttime Surface Activity, Douglas D. Gaffin

Euscorpius

Sand scorpions make burrows from which they emerge at night to seek mates and prey. Previous ecological studies have documented scorpion activities on the sand surface. Few studies, however, have videotaped scorpion movements in the context of understanding their sensory neurobiology. Our objective is to understand which signals guide scorpions as they emerge from and return to their burrows. Candidate stimuli include chemical trails, footsteps, celestial patterns, geomagnetic cues, humidity and/or temperature gradients, distinctive scents from the burrow, seismic echolocation, memory, or landmark orientation. We videotaped scorpions to learn whether they return to their burrows by the same path they …

Polymorphism And Hybridization In Species Of Hottentotta Birula, 1908 (Scorpiones: Buthidae), Wilson R. Lourenço, Eric Ythier, Mark Stockmann, John L. Cloudsley-Thompson

Euscorpius

A new and well documented case of hybridization among scorpions is presented. It was obtained under laboratory conditions between Hottentotta jayakari (Pocock) and Hottentotta salei (Vachon) specimens of which had been collected in the northern and southern regions of Oman. Hybrids were successfully produced not only from F₀ males and females, but also from F₁ males and females, thereby attesting to the fact that the first generation obtained (F₁), was completely fertile. Both F₁ and F₂ broods were composed of dark and pale morphs, indicating that the juveniles could inherit either one or …

Factors That Affect Harem Stability In A Feral Horse (Equus Caballus) Population On Shackleford Banks Island, Nc, Jessa Madosky

LSU New Orleans Theses and Dissertations

Mammal species often live in social groups, but the factors that promote group cohesion can be difficult to analyze due to the prevalence of strong group affiliations. Feral horses maintain stable harems of one or two males and several females, and harem stability is strongly related to individual fitness. Anecdotal evidence and an early study in the non-breeding season suggest that management of the Shackleford Banks island horses with immunocontraception reduces harem stability in the population, providing an opportunity to study the factors that influence harem stability. I investigated the effects of the immunocontraceptive PZP on harem stability during the …

Exploring Sources Of Selection On The Multimodal Courtship Displays Of Two Sister Species Of Wolf Spiders: Schizocosa Crassipalpata And Schizocosa Bilineata, Mitch D. Bern

School of Biological Sciences: Dissertations, Theses, and Student Research

We examined multiple sources of selection on the multimodal courtship display of two sister species of wolf spiders, Schizocosa crassipalpata and Schizocosa bilineata. We first experimentally altered body condition in these two species by employing divergent diet treatments. We then tested for differences in adult male seismic courtship displays, visual courtship displays, and foreleg morphology; as well as adult female foreleg morphology. We then simultaneously examined both content and efficacy-based sources of selection on the visual and seismic multimodal courtship display of both species by conducting mate choice trials with high and low diet individuals across manipulated signaling environments …

The Complexities Of Wolf Spider Communication: Exploring Courtship Signal Function In Rabidosa Rabida, Dustin J. Wilgers

School of Biological Sciences: Dissertations, Theses, and Student Research

Evidence of signal complexity is seemingly pervasive across animal communication systems. Exploring signal function may provide insight into how these displays evolved and are maintained. This dissertation examines the courtship signal function in a grassland wolf spider. Rabidosa rabida lives in an extremely complex environment, and males use complex displays incorporating both visual and seismic modalities. Using several approaches I provide insight into the content and efficacy of the various signal components, as well as how variation in these displays influence female mating decisions in isolation and combined.

First, I manipulated male and female body condition using diet quantity manipulations …

Patterns Of Transience, Sex Bias, And Body Mass In Open-Habitat Rodent Populations, Stephen Edward Rice

Biological Sciences Theses & Dissertations

Rodents are assumed to live their lives in circumscribed (natal) areas with males being more prone to disperse than females and juveniles more prone to disperse than adults. To test these assumptions we examined the initial captures of geographic populations of hispid cotton rat, meadow vole, prairie vole, and marsh rice rat obtained through capture-mark-recapture methods. Capture records were obtained from Kansas and Illinois from long-term studies, and through live-trapping in Chesapeake, Virginia. I evaluated proportions of residents and transients, adults and juveniles, and males and females for significant differences among seasons, years, and geographic locations. The overall body masses …

Rats As Forest Pests In Southeastern Virginia: Girdling By The Hispid Cotton Rat Sigmodon Hispidus As A Significant Source Of Mortality Of Loblolly Pines (Pinus Taeda) In A Successional Pine Forest, Robyn M. Nadolny

Biological Sciences Theses & Dissertations

The hispid cotton rat (Sigmodon hispidus) is a common field rodent throughout the southeastern US, where volunteer loblolly pine trees (Pinus taeda) invade open space and begin the process of ecological succession from field to pine forest. Recent analysis of the diet of S. hispidus indicates that loblolly pine bark is stripped and eaten during the winter months. In this study, we explored the extent of rodent girdling on a 1.23 ha grid in a successional pine forest in southeastern Virginia. During the winter of 2005 we observed damage to 65% of trees in our study area, with 98% …

Redescription And Taxonomic Position Of Tityus Atriventer Pocock, 1897 (Scorpiones: Buthidae), Rolando Teruel, František Kovařík

Euscorpius

The largely neglected Lesser Antillean scorpion Tityus atriventer Pocock, 1897 is herein redescribed, after study of the two syntypes and one additional adult male. The latter was recently collected from Union Island (about 55 km northeast of Grenada), and it implies the first published finding of this species after its original description, and the first record of the species both outside the type-locality and from the Grenadines islands. The taxonomic position of T. atriventer is clarified (including designation of a lectotype and a paralectotype), its diagnosis is updated, a fully illustrated redescription is presented according to the current taxonomy of …

A New Species Of Vaejovis From Prescott, Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey, Michael E. Soleglad

Euscorpius

A new scorpion species, Vaejovis crumpi, sp. nov., is described from Prescott, Yavapai Co., Arizona. This species is related to V. paysonensis Soleglad, 1973, and the “sky island” species of southern Arizona. V. crumpi is compared to the seven Vaejovis species currently reported for Arizona, in particular to V. paysonensis.

An Assessment Of Stable Hydrogen-Isotope Analysis Methods To Assign Geographic Origin To Migratory Red-Tailed Hawks (Buteo Jamaicensis), Carla Marie Ahlschwede

Department of Environmental Studies: Undergraduate Student Theses

Stable-hydrogen isotopes are becoming an increasingly popular method of studying migratory birds, though sample preparation methods may affect results.  In this study I examined feathers from red-tailed hawks (Buteo jamaicensis) to determine the relationship between measure of δD due to inter-feather variation or drying methods, assessed the accuracy of results by using two birds of known-origin and estimated possible natal origins of migratory red-tailed hawks.  Two feathers per individual were taken from 81 wild hawks caught at Hitchcock Nature Center near Crescent IA and from 2 rescued red-tailed hawks, Raptor Recovery Nebraska near Eagle, NE. 119 of the …

Effects Of Gypsy Moth Outbreaks On North American Woodpeckers, Walter D. Koenig, Eric L. Walters, Andrew M. Liebhold

Biological Sciences Faculty Publications

We examined the effects of the introduced gypsy moth (Lymantria dispar) on seven species of North American woodpeckers by matching spatially explicit data on gypsy moth outbreaks with data on breeding and wintering populations. In general, we detected modest effects during outbreaks: during the breeding season one species, the Red-headed Woodpecker (Melanerpes erythrocephalus), increased over pre-outbreak levels, while during the winter one species, the Yellow-bellied Sapsucker (Sphyrapicus varius), increased and one, the Downy Woodpecker (Picoides pubescens), decreased from pre-outbreak levels. Responses following outbreaks were similarly variable, and in general we were unsuccessful …

A Mark–Recapture Technique For Monitoring Feral Swine Populations, Matthew M. Reidy, Tyler A. Campbell, David G. Hewitt

United States Department of Agriculture Wildlife Services: Staff Publications

Techniques to monitor populations of feral swine (Sus scrofa) relative to damage control activities are needed on rangelands. Our objectives were to describe and assess a mark–recapture technique using tetracycline hydrochloride (TH) for monitoring feral swine populations. We established bait stations at study sites in southern and central Texas. During 1 d, we replaced normal soured corn bait with bait containing TH and counted the number of feral swine that consumed bait with observers. We conducted feral swine removal using box-style traps and helicopters, at which time we collected teeth for TH analysis. In southern Texas, we estimated …

A Nebraska Bird-Finding Guide, Paul A. Johnsgard

Zea E-Books Collection

Nebraska lies in the transition zone between North American eastern and western avifaunas and is home to more than 200 breeding and 150 migrant species. This definitive guide to Nebraska birdwatching by the state’s preeminent ornithologist includes a county-by-county rundown of the best sites, a calendar of migrations, an annotated checklist of regularly occurring Nebraska birds, and recommendations for optical equipment, publications and reference materials, and contact information for conservation and ornithological groups. It features 48 maps as well as photographs and drawings by the author.

Paul Johnsgard, Foundation Professor Emeritus of Biological Sciences at the University of Nebraska–Lincoln, is …

Determining The Composition Of The Dwelling Tubes Of Antarctic Pterobranchs, Lukasz J. Sewera

Honors Projects

Pterobranchs are a group of marine invertebrates within the Hemichordata, which share characteristics with both chordates and echinoderms.  Pterobranchs live in colonies of secreted tubes, coenicia, which are composed of a gelatinous material of unknown composition.  Visually, the tubes appear similar to the tunic of tunicates, a group of invertebrates within the Chordata. The nonproteinaceous tunic of tunicates is composed of cellulose, which is unusual.  The goal of this study was to determine the composition of the pterobranch coenicium. Some aspects of pterobranch phylogeny are still unclear even after multiple molecular and morphological studies.  Identification of any new shared characteristics …

The Genus Butheoloides Hirst, 1925 (Scorpiones, Buthidae) In Morocco, With A Description Of А New Species, Wilson R. Lourenço, Oulaid Touloun, Ali Ali Boumezzough

Euscorpius

A new species belonging to the genus Butheoloides Hirst, 1925 (subgenus Butheoloides Hirst, 1925) (Scorpiones, Buthidae) is described from two localities in Morocco: north of Sidi Ifni in the southern coast and Sidi Moussa west of Tiznit. With the description of Butheoloides (Butheoloides) littoralis sp. n., the total number of species of this genus known from Morocco is raised to five. This makes Morocco the region in Africa with the highest diversity for Butheoloides species.

A Comparison Of Adrenal In Toad-Eating And Nontoad-Eating Snakes, Shabnam Mohammadi

Biological Sciences Theses & Dissertations

Toads are chemically defended by bufadienolides, a class of cardiotonic steroids lethal to most predators, including many snakes. Bufadienolides bind to Na+K +-ATPase, inhibiting their ability to transport ions. In cardiocytes, this inhibition cause arrhythmia and severely increased contraction strength, which, if prolonged, lead to death. However, several snakes are resistant to bufadienolides and consume toads with no ill effects. Adrenal glands produce hormones that are important for the maintenance of Na+K +ATPase, and may therefore play an important role in countering the negative effects of bufadienolides. Indeed, the toad-eating specialist Heterodon platirhinos has been known to possess enlarged, and …

Multilocus And Parametric Analyses Of The Evolutionary History Of The Amazonian Peacock Cichlids, The Genus Cichla (Teleostei: Cichlidae), Stuart Clayton Willis

School of Biological Sciences: Dissertations, Theses, and Student Research

Accurate knowledge of species boundaries and species phylogeny are fundamental to testing hypotheses of recent evolutionary processes, but the estimation of these partitions is challenging due both to inherent confusion about what is being estimated as well as the data available to estimate them. Using multilocus data from mtDNA, microsatellites, and nuclear locus sequences of over 1100 individuals, we delimited eight separately evolving species of Cichla rather than the 15 described. Among species we found evidence of rare but widespread introgressive hybridization, while within these species we observed evidence of long-term gene exchange and constrained evolutionary trajectories. In most cases …