Other Animal Sciences Commons^™

Feeding Ecology Of Asiatic Wild Ass Equus Hemionus, Jochen Lengger, Frieda Tataruch, Christian Walzer

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The Mongolian wild ass or khulan (Equus hemionus) is a potential competitor for forage to reintroduced Przewalski’s horses. To evaluate the major foraging plants of khulan we chose the alkane method that was first described by Mayes in 1984. Different plant species contain varying amounts and proportions of n-alkanes. This fact allows the determination of diet composition by comparing the plant alkane to the fecal alkane pattern. The major advantage is that the method is non-invasive and has been shown to be a reliable method in captive trials. Fecal and plant samples were collected simultaneously and preserved by …

Numbers, Distribution And Social Structure Of Kiang (Equus Kiang Moorcroft 1841) Population In The Southwestern Part Of Tibet, China, Natalia V. Paklina, Chris Van Orden

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In September - October 1998 we made an extensive survey in southwest Tibet to study numbers, distribution and social structure of the kiang population in this remote part of its range. Kiangs were sighted between 29º 40’ and 32º 50’ N, and 81º- 86º E. They preferred broad flat valleys with lakes at an altitude from 4,000 up to 5,000 m. Small kiang groups and solitary animals were sighted in Transhimalaya. There were no kiangs in Tsangpo (Brahmaputra) River valley, in the sand desert near Ali and in the Clay Mountains near Tsada. Seven old kiang corpses were found along …

Some Information On The Ecology Of Khulan (Equus Hemionus Pallas, 1775) In The Western Part Of The South Gobi Province, Mongolia, D. Tsendjav, S. Purevsuren

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

This short study provides information on the density and distribution of khulans in September 2003 and speculates about factors that negatively influence the khulan population in the sums of Khurmen, Bayandalai, Noyon, Sevrei, and Gurvantes in the South Gobi province.

Turkmenian Kulan (Equus Onager Kulan) Captive Population, Anna Mekarska

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The Turkmenian kulan (Equus onager kulan) is critically endangered, fewer then 1,500 animals are left in the wild. In 2004, there were additional 570 Turkmenian kulans living in captivity: 156 specimens in Euro-Asian zoos, 83 in Askania Nova Reserve and 331 in Canyon Colorado Equid Sanctuary. Population size in EAZA region is decreasing both in terms of numbers and genetic parameters. Those facts underline the urgent need for coordinating development of Eurasian kulan’s captive population. From 2004 onwards, large efforts to create healthy breeding groups in zoos raised bigger interest in this species in European zoos.

Identification Of Herder-Wild Equid Conflicts In The Great Gobi B Strictly Protected Area In Sw Mongolia, Petra Kaczensky, N. Enkhsaihan, O. Ganbaatar, Christian Walzer

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In protected areas of Mongolian overgrazing, competition with wild ungulates and poaching are important management concerns. The Great Gobi B Strictly Protected Area (SPA) in SW Mongolia is a re-introduction site for the Przewalski’s horse (Equus ferus przewalskii), a stronghold of the Asiatic wild ass (Equus hemionus), and remains an important grazing area for seminomadic herders. We show the power of simple inventory and monitoring methods to assess herder-wildlife conflicts, by combining data of: (1) human and livestock demographic data, (2) migration patterns, of semi-nomadic herders, (3) monthly surveys of wild- and domestic ungulates, and (4) …

Kiangs (Equus Kiang, Moorcroft 1841) In Sikkim, India, Natalia V. Paklina, Chris Van Orden

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Between the 5^th and 10^th of March 2001 we completed a 450 km route by car in Northern Sikkim. During the trip along the southern slopes of the Himalayas and Sikkim Plateau we didn’t see a single kiang. According to the local people and military personnel in Sikkim, kiangs are known only in the outer limits of the Sikkim Plateau, where wild animals have a possibility to migrate free over the Chinese-Indian border. In winter only small groups of kiang incidentally come to the Sikkim Plateau from South Tibet. But in May more of them come there and …

Current Status Of The Khulan (Equus Hemionus) In The Trans-Altai Gobi, Enkhbileg Dulamtseren, Adiya Yadamjav, Dovchindorj Ganbold, Mijidorj Batmunkh

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Since 1975, 25 wildlife surveys have been conducted in the Trans-Altai Gobi. In this paper we summarize results from the last 11 surveys, conducted between October 2000 and April 2005, and discuss our findings together with results from previous surveys.

Assessment Of Non-Metric Skull Characters And Age Determination In The Asiatic Wild Ass Equus Hemionus: A Methodological Approach, Hermann Ansorge, Annegret Stubbe, Nyamsuren Batsaikhan, Ravčigijn Samjaa, Michael Stubbe

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The report presents the first information on the use of non-metric skull characters and age determination by tooth section for Asiatic wild asses. It was initiated by a collection of more than 400 skulls of the Asiatic wild ass from the Bordzongijn-Gobi 2002-2004. The methodological overview gives a first research prospect in Asiatic Wild Ass concerning the use of non-metric skull characters as a morphological tool for assessing genetic variability, determining epigenetic divergences and evaluating fluctuating asymmetry. The time course of tooth eruption and replacement in the Asiatic wild ass is described with regard to its use for age determination. …

Territorial Behaviour Of Kiang (Equus Kiang Moorcroft, 1841) In Ladakh (India), Natalia V. Paklina, Chris Van Orden

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The observations of kiang behavior were made in Navokar Valley northeast of Tso Kar Lake (Ladakh, India) between July 30 and November 22, 2001. In the breeding season (end of July until the end of August) adult kiang males kept not overlapping, protected territories (about 10 km²), and marked by single defecation and urination marks. There were adult females with and without offspring on the territories (up to 12 animals, including the male). The distance between male and females on the territories was usually hundred times bigger, then a distance between stallion and his harem in horses. Females …

Semi-Wild Population Of Kulans In The Bukhara Breeding Centre And Their Co-Habitation With Przewalski’S Horses, Karim Bahloul, Olga B. Pereladova, Natalia V. Soldatova, Ekaterina V. Sidorenko, Antoine J. Sempere

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Asiatic wild asses and Przewalski's horses initially inhabited steppe, semi-desert and desert areas, but Przewalski's horses became extinct in the wild, and kulans are under threat of disappearance. The Bukhara Breeding Centre (Uzbekistan) was created in 1976 for conservation and reintroduction of wild ungulate species. In 1977-1978, five kulans (two males and three females) from Barsa-Kelmes Island on the Aral lake were introduced to the reserve. The group increased to 25-30 animals in 1995-1998, when five Przewalski's horses from Moscow and St. Petersburg zoos were introduced to the same territory. We analyzed the home ranges, preferred habitats and social interactions …

Social Networks In Wild Asses: Comparing Patterns And Processes Among Populations, Daniel I. Rubenstein, Siva Sundaresan, Ilya Fischhoff, David Saltz

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Asiatic wild asses inhabit some of the most arid environments in the world. All live in fissionfusion societies, but demography varies and the deserts in which they live often differ in subtle ways. Characterizing details of social structure of wild ass populations has been a challenge and has made it difficult to determine causes and consequences of any differences that might exist. We use network theory to compare the social structures of two populations of Asiatic asses/ onagers inhabiting the Negev desert, Israel and khur of the Little Rann of Kuch, India and show that populations differ in important structural …

An Overview On The State Of Equus Hemionus In Whole China, Weikang Yang

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The wild ass in China is divided into two species: Gobi khulan (Equus hemionus) and kiang (Equus kiang). Maps and charts of the distribution and population sizes in China and Mongolia of each species are shown from the 1980s through the early 2000s.

Similar Levels Of X-Linked And Autosomal Nucleotide Variation In African And Non-African Populations Of Drosophila Melanogaster, Nadia D. Singh, J. Michael Macpherson, Jeffrey D. Jensen, Dmitri A. Petrov

Biology, Chemistry, and Environmental Sciences Faculty Articles and Research

Background: Levels of molecular diversity in Drosophila have repeatedly been shown to be higher in ancestral, African populations than in derived, non-African populations. This pattern holds for both coding and noncoding regions for a variety of molecular markers including single nucleotide polymorphisms and microsatellites. Comparisons of X-linked and autosomal diversity have yielded results largely dependent on population of origin.

Results: In an attempt to further elucidate patterns of sequence diversity in Drosophila melanogaster, we studied nucleotide variation at putatively nonfunctional X-linked and autosomal loci in sub-Saharan African and North American strains of D. melanogaster. We combine our experimental …

Zur Nahrungsökologie Des Uhus Bubo Bubo In Der Mongolei = Feeding Ecology Of The Eagle Owl Bubo Bubo In Mongolia, Thomas Hoffmann, Michael Stubbe, D. Heidecke, R. Piechocki, R. Samjaa, J. Erfurt, D. Sumjaa

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

This paper presents further results on the food ecology of the eagle owl in western Mongolia for the years 1984 to 1992. Additional to the already known sites we investigated two new resting places. At least 27 mammal and 17 bird species were recorded. The results confirm previous studies. Small to medium sized mammals (Cricetidae, Dipodidae, Arvicolidae) dominate the prey of the eagle owl whereas birds and invertebrates are less important.

Ecology Of Soil Microarthropods In Gobi Gurvan Saykhan Mountains, Southern Mongolia, Tsedev Bolortuya, Badamdorj Bayartogtokh

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In the present paper, we describe the community structure, species diversity, population density, and the character of distribution of soil microarthropods of the Gobi Gurvan Saykhan mountains. In total 45 species of soil microarthropods belonging to 37 genera and 24 families were identified. Twenty-seven species of soil microarthropods belonging to 25 genera and 20 families were found in the Züün Saykhan, 31 species belonging to 27 genera and 19 families in the Dund Saykhan. Oribatid mites are the dominant group (35 species = 77.8% of total species richness), while mesostigmatid mites account for 7 species (15.6 %). Other groups are …

Avifaunistische Beobachtungen Im Westchentej, Ulrich Zöphel, Jochen Schulenberg, Wolf-Dieter Busching, Ingolf Todte

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

This paper reports the results of bird observations at four study sites with many different habitats in the western Chentej-mountains (forest steppe, taiga) at the end of the spring-migration and the beginning of the breeding time. Altogether 134 bird species were observed, about 107 of which are probably breeding birds. Additional birds were caught using 6 mist-nets. In this way 98 birds of 21 species were caught, measured, and ringed. Furthermore feathers have been collected (molting feathers and near two breeding places of Accipiter nisus).

The bird fauna of the larch forests shows a high species richness (108 species), …

Types Of Areas And The Origin Of Black Flies Fauna In Mongolia (Diptera: Simuliidae), Josef Halgoš

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The survey of Mongolian Black Flies species (Diptera: Simuliidae) is given in the presented work. The classification based on Emelyanov (1974) and Gorodkov (1984) appeared to be as most advantageous one. Black Flies species found on the Mongolian territory up to now were classified according to the character of their distribution among 5 groups: Holoarctic species (7 species), Eurasian (15), Eastern Palaearctic (20), Turanian-Mediterranean (7), Mongolian endemic species (8). Individual groups are divided according to the type of area into smaller subgroups. The zoogeographical analysis has shown big similarity with Black Flies zoogeography in Siberia and Far East (in percentage …

Beaver Research In The Uvs Nuur Region, Michael Stubbe, N. Dawaa, R. Samjaa, Annegret Stubbe, A. P. Saveljev, D. Heidecke, D. Sumjaa, H. Ansorge, S. Shar, J.-F. Ducroz

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In 1985, 1988, and 2002 Castor fiber birulai was introduced to the Tes Gol of the Uvs Nuur basin in North-western Mongolia. The beavers migrated through the Republic of Tyva and settled in the middle part of Tes Gol near the Tyvinian-Mongolian border. About 10 colonies were recorded in this region in 2002. Strict protection of Castor fiber birulai has to be ensured in Mongolia and the Republic of Tyva in future.

Vergleichende Untersuchungen Zur Besiedlung Von Ausgewählten Hochgebirgen Der Nordwest- Und Südmongolei Durch Laufkäfer (Coleoptera: Carabidae) = Comparing Investigations Of The Distribution Of Carabid Beetles (Coleoptera: Carabidae) In Different High Mountain Regions In Northwest- And South Mongolia, Peer Schnitter

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In the years 1977-1979 and 1986 two high mountain regions of Mongolia are observed, especially in regard of the carabid fauna within different altitudes. In one of the investigation areas--the mountains of Charchiraa in Northwest-Mongolia the first transect with any plots was arranged from the Mount Suslan-Chamar to the salt lake Uvs Nuur. The second transect in the mountains of Ich-bogd in South-Mongolia starts at the Mount Baruun-bogd and reaches the salt lake Orog Nuur. The species combinations are very different, only some species could be found in both areas. The carabid coenoses and their distribution within the different altitudes …

Das Arteninventar Der Avifauna Der Mongolei Während Einer Nord-Süd-Durchquerung 1997, Tobias Stenzel, Michael Stubbe, R. Samjaa, Annegret Stubbe, C. Dulamsuren

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Here we present a detailed report on bird biodiversity of ecosystems along a North-South transect through Mongolia in 1997whic h serves as a guideline for further research projects and training for students in all Mongolian vegetation zones from northern steppes to southern deserts.

Two hundred fifty-five bird species were observed between 21st of July and 10th of August 1997. The main study sites were located in the North the Selenge-Orchon-basin with meadows and dune woodlands 15 km southern from Suchbaatar, the Middle Gobi-Aimag with desert steppes and sajrs (covered with Ulmus pumila) and in the Bordzongijn-gobi in the South …

First Results Of Wild Ass Research In The South Gobi Aymag/Mongolia In 2003 And 2004, Annegret Stubbe, Michael Stubbe, N. Batsajchan, R. Samjaa, S. Doržderem

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The population of the Mongolian wild ass belongs to the nominate form Equus hemionus hemionus described by Pallas in 1775. This species is adapted to semidesert habitats in the Gobi region and is listed in the Red Book of Mongolia as well as on appendix I of the Washington Convention (CITES). Mapping of the northern border of the distributional range showed a remarkable decrease in size in comparison to the maps of Murzaev (1954) or Bannikov (1954). The pressure of civilization, settlements, motorization, and increasing numbers of domestic livestock in connection with the occupation of water resources and illegal hunting …

Quantitative Investigations On Bird Communities In Different Habitats In The Orkhon-Selenge-Valley In Northern Mongolia, Tobias Stenzel, Michael Stubbe, Ravčigijn Samjaa, Sundev Gombobaatar

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In the breeding seasons 2001 to 2003, the avifauna of a study area of about 80 km² in the Selenge Aymag was studied.

The area is characterized by loose groves of elms (Ulmus pumila) and pine trees (Pinus sylvestris), and a chain of barchan dunes which separate the Orkhon valley from the up to 70 m higher situated forest steppe. Steppe species (Stipa spp.) dominate in the south; the northern steppe part is characterized by Caragana bushes (Caragana microphylla, locally C. stenophylla). Between Orkhon Gol and the eastern dunes, dry steppe …

Beiträge Zur Ökologie Und Verbreitung Der Steppenwühlmaus Microtus Brandti Radde, 1861 In Der Mongolei = Ecology And Distribution Of The Steppe Vole Microtus Brandti Radde, 1861 In Mongolia, N. Dawaa, Michael Stubbe, Annegret Stubbe

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Microtus brandti is one of the most important pest species of Mongolian rodents. Since the publication of Bannikov (1954) the area of expansion is increased, in the Archangaj-Aimag 200 km westward, in the Central Aimag 70 km to the North and in the north-eastern part of Dornogov’- Aimag about 110 km to the South. In years of gradation the plant cover was exterminated up to 100%. We can distinguish a zone of exposed harmfulness with a damage of 70-100%. In this region 16 great centers of infestation were listed. Between 1925 and 1985 six important calamities were noticed, in intervals …

Der Einsatz Der Luzerne-Blattschneiderbiene, Megachile Rotundata (Hymenoptera: Megachilidae), Zur Bestäubung Von Luzerne In Der Mongolei = Management Of The Alfalfa Leaf-Cutter Bee, Megachile Rotundata (Hymenoptera: Megachilidae), For Alfalfa Pollination In Mongolia, Karsten Seidelmann, Čojrog Batchujag, Manfred Dorn

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The propagation of the alfalfa leaf-cutter bee, Megachile rotundata was tested from 1988 to 1991 on alfalfa seed growing fields at the Uvs-Nuur Basin in the North-West of Mongolia. We used the loose-cell technology in combination with wooden grooved boards (dimensions of the nesting holes: 8 × 110 mm). The climatic conditions at the investigation area proved to be favorable for the rearing of the bees. The average temperature was above 21 °C during 8.5 h by a mean sunshine duration of 6.4 h and sporadic rainfall. Nevertheless, the night temperatures felled below 10 °C and are responsible for a …

Die Parasitierung Der Luzerne-Blattschneiderbiene, Megachile Rotundata (Hymenoptera: Megachilidae), Durch Physocephala Pusilla (Diptera: Conopidae) In Der Mongolei = Parasitization Of The Alfalfa Leaf-Cutter Bee, Megachile Rotundata (Hymenoptera: Megachilidae), By Physocephala Pusilla (Diptera: Conopidae) In Mongolia, Karsten Seidelmann

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Physocephala pusilla Meigen (Diptera: Conopidae) was found to be a dangerous endoparasitoid of the alfalfa leafcutter bee Megachile rotundata F. (Hymenoptera: Megachilidae) in Mongolia. The conopid fly parasitizes adults and causes their early death. In 1989 and 1990, 16% and 15% respectively of the M. rotundata females were found dead in front of the bee shelter. However, the number of bees (females and males) dying in the field is unknown. These bees cause a contamination of the alfalfa fields due to an increase of the conopid population. Therefore, Ph. pusilla becomes a serious problem for the management of M. rotundata …

Facts From A Year Of Drought: Forage Competition Between Livestock And The Mongolian Pika (Ochotona Pallasi) And Its Effects On Livestock Densities And Body Condition, Vroni Retzer

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Burrowing small mammals in grasslands have long been regarded as pests because they compete for forage with livestock and reduce the forage availability for livestock by destroying pastures through their intensive digging activity.

In order to investigate forage competition between the Mongolian Pika (Ochotona pallasi) and livestock an exclosure experiment consisting of four different treatments was set up. The treatments were: 1) accessible only for pikas, (only pika) 2) accessible only for livestock, (only livestock) 3) accessible for both herbivore groups (pika & livestock) and 4) no grazing (no grazing). During the investigation period all requirements for forage …

Über Die Federmerkmale Der Mongolischen Sakerfalken Falco Cherrug Milvipes Im Vergleich Zu Den Westlichen Sakerfalken Der Unterart F. Ch. Cherrug Und Bemerkungen Über Die Gefiedermerkmale Des “Altaifalken”, Wolf-Dieter Busching

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The different plumage characteristics of the Mongolian Saker Falco cherrug milvipes and the nominate race F. ch. cherrug are discussed. It could be shown that both races are polymorphic. According to Baumgart (1997), Sakers don’t need uniform plumages because they live on nearly blind voles. Due to that there seems to be no selection pressure from victims side. The same phenomenon is observable with Buteo buteo and the typical Mongolian Buteo hemilasius.

Whereas young Sakers of the nominate race cherrug equal more or less already adults, are the differences in the plumage of juv. and ad. milvipes great enough. …

Wild Bactrian Camel Conservation, Richard P. Reading, Evan S. Blumer, Henry Mix, Jadamsuren Adiya

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The wild Bactrian camel (Camelus bactrianus ferus) is critically endangered throughout its range in China and Mongolia. Yet, wild camels remain poorly understood, with knowledge derived primarily from a few short studies and anecdotal information. We initiated a wild camel conservation project to determine the reasons for camel decline and to develop a program to address those problems.

We are employing satellite telemetry to gather data on wild camel movement patterns, home ranges, habitat use, and sources of mortality. We are also collecting feces from camels and wolves to determine important forage plants and to begin to assess …

Winter Pasture Conditions And Forage Use By Argali (Ovis Ammon) In Gobi Gurvan Saykhan National Park, Bayart Mandakh, Ganchimeg J. Wingard, Richard P. Reading

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Conservation of Argali sheep (Ovis ammon) presents a significant challenge to biologists and conservation managers in Mongolia. Argali are endangered and available data suggest that their numbers are declining. However, the mechanisms causing this decline are poorly understood. Recent research suggests that declines in Argali sheep numbers may be due in part to competition with domestic livestock for forage.

Following privatization of herd ownership in 1991, livestock numbers increased dramatically across Mongolia. With this increase, many herders expanded grazing onto more marginal rangeland resulting in increased competition with wild ungulates and displacement of Argali from their former population …

Ecology Of Argali In Ikh Nartiin Chuluu, Dornogobi Aymag, Richard P. Reading, Sukhiin Amgalanbaatar, Ganchimeg J. Wingard, Danny Kenny, Anthony Denicola

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Argali sheep (Ovis ammon) are listed as threatened in both Mongolia and internationally. Yet, little is known about the biology and ecology of this species. Available data suggests that Argali in Mongolia are declining due to direct poaching and competition with domestic livestock. We initiated several research projects to better understand and conserve the species. In this report we discuss Argali ecology using radio telemetry.

We captured and radio-collared 36 Argali using drive-nets, lamb captures, and dating from 2000–2004. Fifteen collared animals have died: 2 due to capture techniques, 8 from predation, 1 from starvation and exposure, 1 …