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Digging Deep Into Curation: A Guide To Creating A Local Fossil Museum, Syd Joheim, René Shroat-Lewis May 2026

Digging Deep Into Curation: A Guide To Creating A Local Fossil Museum, Syd Joheim, René Shroat-Lewis

Research and Creative Works Expo

In Arkansas, the dissemination of educational content related to evolution and natural history is often constrained by prevailing cultural dynamics. This has resulted in a significant gap in the exposure of primary and secondary school students to foundational concepts in natural history and physical sciences. Even the state’s most popular science and history museums often intentionally exclude exhibits that risk upsetting a large portion of their visitors, further exacerbating the issue. As a consequence, many children do not receive proper education on natural history and physical sciences.  To address this educational deficiency, researchers identified and cataloged specimens from a collection …


A Herpetological Survey Of The Catawba Nation, York County, South Carolina, Courtney Skeldon, Marvin Bouknight, Steven E. Fields May 2026

A Herpetological Survey Of The Catawba Nation, York County, South Carolina, Courtney Skeldon, Marvin Bouknight, Steven E. Fields

Journal of the South Carolina Academy of Science

No abstract provided.


Behavioral And Molecular Effects Of 2,4-Dichlorophenoxyacetic Acid Exposure In Planarians, Emily Pintilie May 2026

Behavioral And Molecular Effects Of 2,4-Dichlorophenoxyacetic Acid Exposure In Planarians, Emily Pintilie

Graduate Student Dissertations, Theses, Capstones, and Portfolios

The widespread use of pesticides, such as 2,4-dichlorophenoxyacetic acid (2,4-D), raises serious concerns about their effects on non-target aquatic organisms. In vertebrates, 2,4-D has been associated with neurotoxic outcomes. Planarians offer a valuable system for toxicological studies because of their well-characterized nervous system, regenerative capacity, and measurable behavioral responses. This study investigated the neurotoxic effects of 2,4-D exposure in the planarian species Girardia dorotocephala, including the assessment of nociceptive behaviors, survival rate, phototactic responses, and DNA methylation. Worms were exposed to various concentrations of 2,4-D (0–100 mg/L) at multiple time points. Nociceptive responses, such as scrunching, C-shapes, and seizures, …


The Detection Of Forever Chemicals From Paper Cups Using Colorimetry, Sara Miller May 2026

The Detection Of Forever Chemicals From Paper Cups Using Colorimetry, Sara Miller

Graduate Student Dissertations, Theses, Capstones, and Portfolios

Per- and polyfluoroalkyl substances (PFAS) are synthetic chemicals added to food packaging products to increase water-resistance and durability. PFAS are a growing concern due to their migration into beverages and foods and have been linked to cancer, immunotoxicity, and thyroid disturbances. The purpose of this study was to quantify PFAS migration from paper cups into water and evaluate toxic effects using planarians. PFAS detection was performed using a colorimetric assay, which forms a colored complex with PFAS, on: perfluorononanoic acid (PFNA), heptadeca fluorooctane sulfonic acid potassium salt (PFOS), perfluorooctanoic acid (PFOA). PFNA and PFOS showed absorbance at 640 nm, while …


Virtualizing Practical Science Education: A Sociological Analysis Of Undergraduate Teaching And Learning, Sina J. Fakoyede, Femi E. Babalola, Oluwatoyin T. Obateru, Oluwayemisi D. Akomolafe, Deborah O. Alabi, Stella K. Ekundayo May 2026

Virtualizing Practical Science Education: A Sociological Analysis Of Undergraduate Teaching And Learning, Sina J. Fakoyede, Femi E. Babalola, Oluwatoyin T. Obateru, Oluwayemisi D. Akomolafe, Deborah O. Alabi, Stella K. Ekundayo

Higher Learning Research Communications

Objectives: The study aims to investigate the perspectives of university teachers and students on integrating practical science teaching with virtual learning in a multicultural and diverse society, specifically within the context of southwestern Nigeria.

Methods: A mixed-methods approach was employed, involving data collection from 100 university lecturers and 400 undergraduate students across 10 universities. Qualitative data were gathered through semi-structured interviews with lecturers and focus group discussions with students. Quantitative data were obtained via structured questionnaires, allowing for triangulation and comprehensive analysis.

Results: Findings revealed that certain lecturers showed preferential support influenced by students’ cultural backgrounds or academic behavior during …


Bio 340 Final Project: Effects Of Light Pollution On Bat Activity, Cael Schneider May 2026

Bio 340 Final Project: Effects Of Light Pollution On Bat Activity, Cael Schneider

Biology Student Scholarship

Cael Schneider - Environmental Biology ‘28

Artificial light at night (light pollution) is an increasing consequence of urbanization and can disrupt nocturnal wildlife. Bats are especially susceptible, as they rely on dark skies for navigation, predator avoidance, and foraging. The effects of artificial light may vary among species due to differences in foraging strategies, habitat use, and light tolerance.  Previous research has documented negative effects of artificial light on bat foraging, roosting, breeding, and hibernation. However, some species, such as red bats, may forage near artificial lights where insects are abundant. Species including the big brown bat, hoary bat, and …


Life Histories Of Two Sgcn Procambarus Species In East Texas, Mckenzie A. Adkins May 2026

Life Histories Of Two Sgcn Procambarus Species In East Texas, Mckenzie A. Adkins

Biology Theses

Within freshwater systems, crayfish play notable roles by shaping habitats and linking food web levels. Yet even with such influence, knowledge about numerous species is incomplete, which hinders protection strategies. Found only in the Neches River basin in eastern Texas, Procambarus nechesae and Procambarus nigrocinctus have different conservation statuses, mainly because data on their life history remains sparse. These two species are not closely related and are analyzed independently in this study. Research into traits tied to survival and environmental preferences was carried out so that informed decisions on conservation efforts can be made for these species. The goal of …


Proximity Data And Social Networks: A Validation Study Of Two Models Of Mobile Dataloggers In The Semi-Fossorial And Social Rodent, Octodon Degus, L. D. Hayes, A. A. Maldonado-Chaparro, Jessica Morales Valenzuela, L. Platt, L. A. Ebensperger May 2026

Proximity Data And Social Networks: A Validation Study Of Two Models Of Mobile Dataloggers In The Semi-Fossorial And Social Rodent, Octodon Degus, L. D. Hayes, A. A. Maldonado-Chaparro, Jessica Morales Valenzuela, L. Platt, L. A. Ebensperger

Biology Faculty Works

The use of mobile dataloggers to gather proximity data has generated novel insights into the social network structure of animals. The performance of mobile dataloggers may be impacted by different environmental and social conditions. Thus, validation studies that account for species-specific habitat use and behavior are essential. We conducted a series of validation studies of two brands of mobile dataloggers (Gipsy6, Technosmart, Italy; ProxLog, IoSA, Belgium) on the semi-fossorial rodent, Octodon degus. First, we determined the sensitivity of both types of dataloggers to track position and closeness in different habitats (Experiments 1 and 2). To determine if the structure …


The Ccu Disease Hunters: Putting Undergraduate Research To Work To Improve Community Health And Training The Next Generation Of Doctors, Paul E. Richardson May 2026

The Ccu Disease Hunters: Putting Undergraduate Research To Work To Improve Community Health And Training The Next Generation Of Doctors, Paul E. Richardson

Journal of the South Carolina Academy of Science

No abstract provided.


Beyond The Bug: How The Covid-19 Pandemic Exposed Longstanding Systemic Failures In The United States Workforce And Economy, Abbey M. Markley May 2026

Beyond The Bug: How The Covid-19 Pandemic Exposed Longstanding Systemic Failures In The United States Workforce And Economy, Abbey M. Markley

Honors Theses

The purpose of this thesis is to investigate and discuss how longstanding systemic failures in the United States exacerbated the effects the COVID-19 pandemic had on the American economy, essential healthcare workers, and overall public trust in institutions. A review of the existing literature surrounding this topic and the subsequent results of various studies were used to compose this thesis. The United States has a painful lack of worker protections and tends to view members of its workforce as expendable, even in times such as pandemic when they are deemed essential. Healthcare workers in particular faced increasing threats to their …


Antibiotic Resistance Patterns In Bacterial Isolates From Mississippi Gulf Coastal Environments, Lawler F. Horn May 2026

Antibiotic Resistance Patterns In Bacterial Isolates From Mississippi Gulf Coastal Environments, Lawler F. Horn

Honors Theses

Coastal environments host diverse microbial communities that play essential ecological roles but may also serve as reservoirs for antibiotic-resistant bacteria. This study investigated patterns of antibiotic resistance in microbial isolates collected from public beaches along the Mississippi Gulf Coast. A total of 143 bacterial isolates exhibited growth and were included in antibiotic susceptibility screening. High levels of antibiotic resistance were observed, with 137 of 143 isolates showing growth on vancomycin, 124 on amoxicillin, and 102 on erythromycin. Resistance to kanamycin (19/143 isolates) and tetracycline (33/143) was less common. Multidrug resistance was widespread, with 101 isolates (70.6%) resistant to three or …


Acquisition Behavior Response: Investigating The Ability Of Leptoglossus Phyllopus To Choose Between A “Good” Versus “Sub-Optimal” Strain Of Caballeronia, Irene L. Pham May 2026

Acquisition Behavior Response: Investigating The Ability Of Leptoglossus Phyllopus To Choose Between A “Good” Versus “Sub-Optimal” Strain Of Caballeronia, Irene L. Pham

2026 Spring Honors Capstones Projects

Leptoglossus phyllopus is a polyphagous pest that primarily feasts on citrus, tomatoes, and legumes. They must form a mutualistic symbiosis with Caballeronia every generation by ingesting the bacterium from their environment. However, not all species of Caballeronia confer the same fitness on the L. phyllopus host, raising the question of whether L. phyllopus possesses a partner choice mechanism that would allow it to distinguish between a good (GAOX1) and a suboptimal (Lep1P3) symbiosis partner. This was examined by conducting a choice experiment in which the two symbionts (GAOX1 vs. Lep1P3) were placed in a plastic box, and the bug’s roaming …


The Cranial Morphology Of Large Wild Versus Captive Felids, Bryttin Boyde, Hannah Selvey May 2026

The Cranial Morphology Of Large Wild Versus Captive Felids, Bryttin Boyde, Hannah Selvey

Caravel Undergraduate Research Journal

Studies have shown that carnivores in captivity that are fed soft diets have more periodontal problems and that the morphology of their skeletons, skulls specifically, responds to environmental and behavior factors. Considering that captive diet affects teeth and gums, we have conducted research on Dr. Adam Hartstone-Rose’s hypothesis that variation will also be discernible through the examination of cranial morphology. We compared captive and wild lions Panthera leo and tigers Panthera tigris to see if differences were present, then analyzed data collected by a microscribe using a 3D morphometric analysis program. Next, Principal Component Analysis was completed to account for …


Exploring Microbiome Communities Of Two Belize Coral Species Five Years After Reciprocal Transplant, Lily E. Kratz May 2026

Exploring Microbiome Communities Of Two Belize Coral Species Five Years After Reciprocal Transplant, Lily E. Kratz

Biology Honors Papers

Coral holobionts, which include the coral animal host, dinoflagellate algae, bacteria, etc., support coral reef biodiversity by providing habitat to many marine species. In the face of global change, identifying and understanding the microbial diversity of corals is extremely important because their survival indicates overall ocean health. Coral-associated bacterial and algal communities contribute to the health and survival of a coral species through creating resistance to stressors and supplying essential nutrients. As coral reef environments continue to change, microbial communities must respond to ensure the survival of reef-building corals. To investigate if coral microbiomes possess the ability to acclimatize to …


Bats In A Modified Suburban Landscape, Brittany Morgera May 2026

Bats In A Modified Suburban Landscape, Brittany Morgera

Biology Honors Papers

This research assesses bat species richness and activity across a gradient of suburban habitat modification surrounding Ursinus College and Green Lane Park in Montgomery County, located in southeastern Pennsylvania. Habitat modification was quantified using GIS-based land-cover metrics (e.g. impervious surfaces and canopy cover), and bat activity and species richness were measured though acoustic surveys conducted at 15 sites during the summers of 2024 and 2025. The project tests the hypothesis that increasing habitat modification has a negative correlation with bat species richness and activity.


Red Lines, Green Blooms: Assessing The Effects Of Historical Environmental Disparity On Algal Communities In The Twin Cities, Mn, Georgia C. Akins May 2026

Red Lines, Green Blooms: Assessing The Effects Of Historical Environmental Disparity On Algal Communities In The Twin Cities, Mn, Georgia C. Akins

Environmental Studies Honors Projects

Redlining, a discriminatory housing policy from the 1930s, shaped lastng urban inequalites that may extend to environmental conditons. This study examines links between redlining and urban water quality in the Twin Cites using algal communites as indicators. Across 28 water bodies sampled in 2022 and 2024, physical, chemical, and biological metrics were compared by redlining grade. Results show limited relatonships between redlining and most variables, with only surface water temperature differing significantly. Nutrients and algal diversity were unrelated to redlining, instead reflectng present-day conditons. These findings suggest contemporary factors play a stronger role than redlining in shaping urban freshwater ecosystems.


Employing Zebrafish To Understand Genetic Drivers Of Epilepsy-Related Comorbid Behaviors, Chinwendu Ononuju, Kharma Hall, Olivia Beatty, Scott C. Baraban, Colleen Carpenter-Swanson May 2026

Employing Zebrafish To Understand Genetic Drivers Of Epilepsy-Related Comorbid Behaviors, Chinwendu Ononuju, Kharma Hall, Olivia Beatty, Scott C. Baraban, Colleen Carpenter-Swanson

Biology Faculty Publications

Children with epilepsy frequently experience a range of significant comorbidities beyond seizures, such as motor dysfunction, cognitive impairment, and neurodevelopmental delays. In some cases, these comorbidities contribute more significantly to overall disease burden than the seizures themselves. To improve quality-of-life (QOL) for these children, treatment options should be selected that control seizures and ameliorate comorbidities. Unfortunately, such therapeutics remain largely elusive. Addressing this issue requires suitable preclinical models. Here we used CRISPR-generated zebrafish with single-gene mutations linked to pediatric epilepsy and applied clinically-relevant behavioral assays to study the effects of these mutations. Using high-throughput locomotion-based assays, we uncovered errors in …


Status Of The Potentially Invasive Australian Redclaw Crayfish (Cherax Quadricarinatus), Margaret (Addie) Munn May 2026

Status Of The Potentially Invasive Australian Redclaw Crayfish (Cherax Quadricarinatus), Margaret (Addie) Munn

Biology Theses

The introduction of invasive species, including crayfish, is common through aquaculture and aquarium trade pathways and is an important factor causing the decline and/or extinction of native species. Invasive species cause billions of dollars in damages annually and threaten almost half of the native species in the United States. Australian Redclaw Crayfish (Redclaws) are a species of large individuals that originated in Australia and New Guinea and have been introduced specifically for aquaculture and the aquarium trade. Redclaws are known to thrive in tropical and subtropical locations. They can tolerate high temperatures and low oxygen and are generalist omnivores that …


Ai×Life 2026: Advances In Computational Biology Across Scales For Life & Health Sciences, Suma Mohan S., Devika N. T., Pranavathiyani G. May 2026

Ai×Life 2026: Advances In Computational Biology Across Scales For Life & Health Sciences, Suma Mohan S., Devika N. T., Pranavathiyani G.

Conference Proceedings

Book of Abstracts of the International Conference, AIxLife 2026: Advances in Computational Biology Across Scales for Life & Health Sciences, held on 17–18 April 2026, SASTRA Deemed to be University, Thanjavur, Tamil Nadu, INDIA 

About the Conference

Conventional computational biology laid the foundation for biological data analysis through statistical models, algorithms, and curated databases, enabling insights into genes, proteins, and pathways. Today, the integration of artificial intelligence, spanning machine learning, deep learning, and large language models (LLMs) is driving a transformative shift across biological scales, from atomic and molecular interactions to pathways, cellular systems, organisms, and populations.

AI-powered approaches …


Assessing The Conservation Needs Of Native Texas Crayfish Species Through Distribution Analysis And Species Distribution Modeling, Elizabeth J. Croft May 2026

Assessing The Conservation Needs Of Native Texas Crayfish Species Through Distribution Analysis And Species Distribution Modeling, Elizabeth J. Croft

Biology Theses

Crayfish have essential roles in ecosystems because they are ecosystem engineers and keystone species. However, crayfish species are threatened by degradation and loss of habitats, introduction of invasive species, and high levels of endemism. Texas provides no legal protections for crayfish, despite its large number of species and endemics in the state. The overall goal of this study is to clarify the distributions of native Texas crayfish species using literature records, citizens science, and field sampling data, and highlight species in need of more conservation research. Occurrence records from all collected data were combined to visualize the distributions of species …


A Series Of Investigations On The L. Cornutus Host System And Bivesiculid Sp. Cercariae, Briana K. Zaffiro May 2026

A Series Of Investigations On The L. Cornutus Host System And Bivesiculid Sp. Cercariae, Briana K. Zaffiro

All HCAS Student Capstones, Theses, and Dissertations

Rising global temperatures are likely modulating parasite-host interactions, yet the direction and magnitude of these changes remain uncertain. Rarely are these interactions viewed from both a host and parasite perspective. Using both CTmax, or Critical Thermal Maximum, and Clark-type polarographic oxygen microelectrodes, the physiological metabolic effects are investigated. CTmax is examined in Common Shiners, Luxilis cornutus, and is shown to be modulated by parasitism: uninfected fish had the highest CTmax average of 37.0°C, whereas heavily infected fish had decreased CTmax averaging 35°C. Prior to the parasite metabolic investigation, a suitable medium was determined, as …


Shape Variation In The Mandible Of The Borophagine Canids And Implications For Predatory Behavior, Benjamin Neavear May 2026

Shape Variation In The Mandible Of The Borophagine Canids And Implications For Predatory Behavior, Benjamin Neavear

Theses, Dissertations and Culminating Projects

The borophagines are an extinct lineage of canids that inhabited North America for more than 20 million years. This group included genera such as Aelurodon, Borophagus, and Epicyon. Borophagines are commonly referred to as “bone-crushing dogs” because their skull morphology converged with that of extant hyenas, predators capable of consuming bone. The purpose of this study was to determine whether convergence was present in the mandible, an anatomical structure strongly correlated with feeding behavior, particularly in the genera Aelurodon and Borophagus. It was hypothesized that most borophagine taxa possessed mandibular shapes similar to those of hyenas. Numerous mandibular characters were …


Geospatial Science And The Changing Environment: Applied Methods For Sustainable Agriculture And Landscape Management, Harrison Wakefield Smith May 2026

Geospatial Science And The Changing Environment: Applied Methods For Sustainable Agriculture And Landscape Management, Harrison Wakefield Smith

Graduate Theses and Dissertations

Rapid environmental change is disrupting agricultural productivity, ecological function, and the long-term resilience of managed landscapes. At the same time, the rapid growth of geospatial data science has improved our understanding of environmental change and increasingly is being used to improve sustainability in agricultural and environmental management. However, critical gaps remain that limit the applicability of data-driven insights in landscape management. This dissertation investigates the potential of geospatial analytics for sustainable agriculture and landscape management, with a focus on applied methods that operate across spatial and temporal scales. Using field, landscape, regional, and national datasets, it explores the capabilities and …


Aggregation And Collation Of Texas Crayfish Museum Specimens To Determine Their Current Conservation Statuses, Adrian Alvarez May 2026

Aggregation And Collation Of Texas Crayfish Museum Specimens To Determine Their Current Conservation Statuses, Adrian Alvarez

Biology Theses

Crayfish account for over half of the macroinvertebrate biomass in many ecosystems and serve as crucial bioindicators because of their behavioral and physiological adaptability to environmental changes. Despite being home to approximately 52 native crayfish species, Texas currently lacks protective regulations. This gap is primarily because of insufficient information regarding their distribution and occurrences within the state, including their statuses as Species of Greatest Conservation Need (SGCN). This study focuses on compiling and comparing crayfish museum records with current iNaturalist field survey data across Texas to address these knowledge gaps and recommend updates to SGCN statuses. Five species not currently …


Using The Flexon Approach To Rnai For Analysis Of Gene Function In Gonadal Development In C. Elegans, Andrew D. Spalding May 2026

Using The Flexon Approach To Rnai For Analysis Of Gene Function In Gonadal Development In C. Elegans, Andrew D. Spalding

Undergraduate Theses

C. elegans, a nematode worm, was used as a model organism to analyze the developmental effects of knocking down the expression of genes that are both essential to worm viability and highly enriched in reproductive tissues. In earlier research, approximately 70 C. elegans genes were identified that are both gonad-enriched and essential. To prioritize which genes should be the focus of future studies, a new strain of C. elegans was produced with three specific genetic variations that: 1) silence essential, somatic gonad-enriched gene expression, 2) limit the effects of this gene silencing to the gonad, and 3) yield a …


Hormonal, Cytokinetic, And Skeletal Response To A Single Weighted Vest Exercise Treatment: A Pilot Study To Determine Weighted Vest “Rucking” Treatment For Osteoporosis In Post-Menopausal Women, Eleanor Pell May 2026

Hormonal, Cytokinetic, And Skeletal Response To A Single Weighted Vest Exercise Treatment: A Pilot Study To Determine Weighted Vest “Rucking” Treatment For Osteoporosis In Post-Menopausal Women, Eleanor Pell

Master's Theses

Osteoporosis is characterized by a deteriorating of bone tissue leading to brittleness and increased fracture risk that affects both men and women as they age. Postmenopausal women are of particular interest due to high risk of declining bone density, a characteristic of this population. Treatments for osteoporosis range from menopause hormone therapy, bisphosphonates, biologics, nutritional changes, and exercise. Load bearing exercise interventions have shown effectiveness for stimulating bone growth, but many factors such as lack of access, motivation, self-confidence or knowledge, prevent long term compliance. “Weighted vest rucking” refers to a variation of rucking, which is a form of exercise …


Investigating The Taxonomic Uncertainties Of Potentilla Rupincola, Natalie Elizabeth Hieber May 2026

Investigating The Taxonomic Uncertainties Of Potentilla Rupincola, Natalie Elizabeth Hieber

Master's Theses

Landscape stewardship is an integral role for land managers that becomes more important when rare or endangered species occur within managed areas. Conservation across the landscape is a primary concern for land managers, but resources allocated for conservation can be limited due to many competing goals. Taxonomic uncertainty surrounding rare and endangered taxa, particularly plant taxa, that have historically been classified solely using morphological features further complicates conservation planning. One such taxon is Potentilla rupincola, a rare plant endemic to the eastern Rocky Mountains in Colorado. For over a century, botanists have debated whether P. rupincola is a distinct species …


Developing Warning Signage For Harmful Algal Blooms At Lake Fayetteville, Abigail R. Evans May 2026

Developing Warning Signage For Harmful Algal Blooms At Lake Fayetteville, Abigail R. Evans

Biological Sciences Undergraduate Honors Theses

The presence of harmful algal blooms (HABs) poses many risk factors for human health, and once microcystin concentration has reached a certain level, that water source will be closed for the duration of the bloom. Agencies tend to use static signage with limited information to inform the public about the current status of the water, but a sign similar to the one used by the National Fire Danger Rating System can allow for constant updates detailing specific risk levels for the public. Using previously reported data on Lake Fayetteville, thresholds for risk factors that are easily measurable were determined. The …


The Genetic Basis Of High Transformation Efficiency In The Model Yeast Saccharomyces Cerevisiae, Mary Kathryn E. Cheek May 2026

The Genetic Basis Of High Transformation Efficiency In The Model Yeast Saccharomyces Cerevisiae, Mary Kathryn E. Cheek

Biological Sciences Undergraduate Honors Theses

Transformation, a type of horizontal gene transfer, describes the uptake, maintenance, and integration of environmental DNA by cells. Despite the known effects of transformation on evolutionary processes, little is known about the genetic basis of this phenomenon. Saccharomyces cerevisiae is a eukaryote able to transform at quantifiable rates when made chemically competent and treated with a heat shock. S. cerevisiae also has variation in transformation efficiencies between various strains, and high recombination rates allow for sufficient allele “shuffling” in intercross populations. This project utilizes a bulk segregant analysis approach to compare sequencing data between transformant pools and non-transformant pools of …


Mechanisms Of Life History Trade-Offs In The Evolution Of Eusociality, Frances Katherine Hunter May 2026

Mechanisms Of Life History Trade-Offs In The Evolution Of Eusociality, Frances Katherine Hunter

All Graduate Theses and Dissertations, Fall 2023 to Present

When faced with stressors such as pathogen infection, organisms must invest energy in defense or risk death. Because resources are limited, mounting an immune response often comes at the expense of other costly activities such as reproduction. The costs of immunity are shaped by factors including behavior, nutrition, age, and evolutionary history.

Social insects (e.g., ants, bees, and wasps) offer a powerful system for studying these trade-offs because queens in these colonies appear to escape them—living long lives while reproducing prolifically. Bees, which include species that range from those which nest solitarily to those with complex social systems and many …