Geography Commons^™

On The Importance Of Pollen Morphology In Classification Of Chenopodiaceae In Mongolia, Gaadan Punsalpaamuu, Frank Schluetz, Tserendorj Gegeensuvd, Davaadorj Saindovdon

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The pollen morphology of 18 species from 12 genera of Chenopodiaceae native to Mongolia was studied. Plants were collected during field courses and research projects on useful plants and are deposited in the Herbarium of the Mongolian State University of Education. Chemical preparation was carried out in the Palynological Laboratories at the School of Natural Sciences of the Mongolian State University of Education and the Department of Palynology and Climate Dynamics of the University of Göttingen in Germany. The results show, that all investigated species are distinguishable in terms of pollen morphology bases on their pollen grain sizes, exine thickness …

Home Range Characteristics Of Corsac And Red Foxes In Mongolia, Tserendorjiin Munkhzul, J. D. Murdoch, R. P. Reading

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Red (Vulpes vulpes) and corsac foxes (V. corsac) live sympatrically throughout most of Mongolia, but few details of their home range characteristics exist. We captured and radio-tagged 13 red fox (♂ = 5, ♀ = 8) and 15 corsac foxes (♂ = 8, ♀ = 7) between 2004 and 2008. We tracked their movements to estimate home range sizes and examined the effects of four factors on home range size, including sex, age, season, and year. We determined mean home range size for 12 red and 10 corsac foxes that had sufficient data using fixed kernel …

Pleistocene Faunal Fossils From Bayangol I Site, Bulgan Aimag, Mongolia, Takao Sato, Fedora Khenzykhenova, Toshiaki Tsurumaru, Masao Ambiru, Jun Takakura, Yoshiaki Otsuka, Shigeo Iida, Natalya Schepina, Batmunkh Tsogtbaatar

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

During excavation of the Bayan-gol I Palaeolithic site in the Bulgan Aimak, Mongolia, some faunistic remains were recovered attributable to Aves (two species), Anura (two species), and mammals (three species). The fossil remains bear a considerable similarity to the corresponding species of today in Mongolia.

Ochotona Daurica Pallas, 1776: Modern And Past Distribution Area In Mongolia And The Transbaikal Region, M. A. Erbjeva, N. V. Alexeeva, T. V. Kisloschaeva

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Ochotona daurica Pallas, 1776 is one of the ancient species among modern taxa having an origination in Transbaikalia possible at the end of Pleistocene. Review on the Late Pleistocene- Holocene-Recent area of distribution of this species is given on the base of new evidences and detail analysis of the previous data.

Ecology And Behaviour Of The Black-Billed Capercaillie (Tetrao Urogalloides Stegmanni) In The Khentej Mountains, Mongolia, Siegfried Klaus, Karl-Heniz Schindlatz, Alexander V. Andreev, Hans-Heiner Bergmann

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Introduction

A common inhabitant of East Siberian larch forests, the Siberian or black-billed capercaillie (Tetrao urogalloides) ranges south to the limits of the boreal forests in the northern Mongolian mountains and east to the very coast of the Asian continent (KLAUS et al. 1989). Tetrao urogalloides MIDDENDORF (1851) has priority over T. parvirostris BONAPARTE (1856). Therefore, we prefer the T. urogalloides.

The subspecies T.u. stegmanni was first described on the basis of morphological differences by POTAPOV (1985) using specimens collected during Russian expeditions by KOZLOVA (1930). This description was based on 18 males in the collection at …

Recruitment Of Larix Sibirica Ledeb. In Closed Forest Stands, On Clear-Felling Sites And At Fire-Sites In The Forests Of Mongolia, Vasiliy T. Yarmishko, Nikolay N. Slemnev

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The paper deals with recruitment patterns in larch forests of Mongolia following anthropogenic impacts (felling, fires), and describes successional trends in highland forest communities. It is established that mass seed recruitment of Larix sibirica Ledeb. took place during anomalous combinations of hydrothermal conditions supposedly occurring at periods of about 100 years. During the last decades, frequent fires of various intensities put serious constraints on reforestation of the larch, and induced successional trends in disturbed forests.

Osnabrück Botanical Expeditions To Mongolia, Barbara Neuffer, Nikolai Friesen, Batlai Oyuntsetseg, Tseden Jamsran, Herbert Hurka

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Three botanical expeditions to Mongolia have been undertaken by the Botany Department of the University of Osnabrück in cooperation with the Botany Department of the National Mongolian University of Ulaanbaatar. The first expedition in 2000 took us to the Mongolian Altay, the second in 2001 to the Gobi and Gobi Altay, and the third in 2010 to East Mongolia including the Khingan Mts. (Numrug). In 2000, we were the first botanists being allowed to enter the border area between China and Russia in the Mongolian Altay since a long time, because this had been a prohibited area for decades. We …

Biodiversity And Evolutionary Development Of Oligocene-Pliocene Lagomorphs (Lagomorpha, Mammalia) Of Mongolia, M. A. Erbajeva

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Lagomorphs (pikas and hares) are an ancient group of small mammals originated in Asia in the Paleocene-Eocene. The earliest evidence of their presence in Mongolia is dated to the Early Oligocene. The taxa flourished during the Late Oligocene, Miocene and Pliocene; at the Late Pliocene they were reduced both in their diversity and in abundance. No data on lagomorph are known from the Pleistocene and Holocene, though there are 7 taxa of lagomorphs present in the modern fauna. Altogether, more than 50 lagomorph species, extinct at present, are known to have existed in Mongolia since the Oligocene through the Late …

Diversity And Community Pattern Of Darkling Beetles (Coleoptera: Tenebrionidae) Along An Ecological Gradient In Arid Mongolia, M. Pfeiffer, E. Bayannasan

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

One of the most conspicuous detrito- and phytodetritophagous groups of beetles in the Asian steppes and deserts is the family Tenebrionidae (Coleoptera, Polyphaga) (KONSTANTINOV et al. 2009). Mongolia harbors a rich diversity of these beetles with 215 species and 50 genera of Tenebrionidae listed for the country (MEDVEDEV 1990), many of them restricted to its arid parts, where they obtain high abundances and dominate – together with ants – the insect soil fauna (PFEIFFER et al. 2003). Their dark color may be a means to withstand the high radiation at daytime; however, a large number of species is night active …

“Agrionemys Kazachstanica Terbishi” Or The Two-Faced Mongolian Steppe Tortoise, Hermann Ansorge, Uwe Fritz, Khayankhyarvaa Terbish, Setev Shar

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Although no extant native turtle and tortoise species is known to occur in Mongolia, a new subspecies of the Central Asian tortoise was described by Chkhikvadze under the name Agrionemys kazachstanica terbishi in 2009. The description was based on a mummified tortoise kept in a museum collection. Since then the Mongolian steppe tortoise has been considered as an endemic taxon Testudo horsfieldii terbishi (Chkhikvadze, 2009) for Mongolia.

However, there is no evidence for the occurrence of any wild tortoise species in Mongolia, even in the putative area of origin of the type specimen. The closest confirmed occurrence of Central Asian …

New Species Of Eimeria (Apicomplexa: Eimeriidae) From Ochotona Hyperborea And Ochotona Pallasi (Lagomorpha, Ochotonidae) In Mongolia, David S. Tinnin, Ethan T. Jensen, Nyamsuren Batsaikhan, Sumiya Ganzorig, Scott Lyell Gardner

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

An examination of the feces from 8 pikas collected in 1999 and from 17 pikas collected in 2009 in Mongolia revealed the presence of 3 new eimerian species. Four of the 5 species of pikas present in Mongolia were studied including: Ochotona alpina, O. dauurica, O. pallasi, and O. hyperborea. Oocysts of Eimeria dunnumi n. sp. from O. hyperborea collected in 1999 are ellipsoid, average length and width of 31.4 x 20.8 μm, with a 1.4 μm thick double layered wall, lacking a micropyle, oocyst residuum, and polar granule. Sporocysts of this species are ellipsoid, 12.8 …

Herpetological Diversity Of Mongolia And Its Conservation Issues, Khorloo Munkhbayar, M, Munkhbaatar

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

From the viewpoint of evolution, the classes of amphibians and reptiles could be considered as relicts, and because they are poikilothermic animals, it’s very difficult for these species to live under the dry and cold climatic conditions in Mongolia. Even species diversity is poor, Mongolian herpetological composition is unique, highly adopted to the country’s harsh climate and originated a long time ago.

In Mongolia, six species of amphibians belong to four genera, four families and two orders and the recorded 21 species of reptiles belong to 13 genera in six families of two suborders.

Morphology And Variation Of The Asiatic Wild Ass (Equus Hemionus Hemionus), Kati Schöpke, Annegret Stubbe, Michael Stubbe, Nyamsuren Batsaikhan, Renate Schafberg

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

A collection of recent skeletal remains from the Asiatic wild ass (Equus hemionus hemionus PALLAS) was prepared and stored in the Museum of Domesticated Animals “JULIUS KÜHN” in Halle, Germany.

The collection is based on carcass remains sampled between 2001 and 2006 under the leadership of Michael Stubbe during joint Mongolian-German Biological Expeditions

Skeletal remains of 43 individual specimens were studied (18 mares, 15 stallions, and 10 juvenile asses). The determination of sex and age was initially based on carcass information from observations made in situ in the field. Subsequently, skull traits, including dentition and development of canine teeth …

Long-Term Ecology Of Asiatic Wild Ass (Equus H. Hemionus Pallas) In Central Asia, Annegret Stubbe, Michael Stubbe, Nyamsuren Batsaikhan, Ravčigijn Samjaa

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The main population of the nominate form Equus hemionus hemionus Pallas, 1775 occurs in the south-east Gobi desert of Mongolia. From 2003 to 2012 we studied the reproductive ecology and mortality of dschiggetajs or khulans (English also ‘hulan’). During these years, the rate of reproduction was variable, ranging from 6.4% to 23.0% depending on climate, feeding conditions, and fitness of the mares. Our research shows that there appears to be a relationship between minimum temperatures in June (main foaling time) and rate of successful reproduction. The primary cause of decline in the numerical density of the population of E. hemionus …

Bacterial Diversity In Biological Soil Crusts From Extrazonal Mountain Dry Steppes In Northern Mongolia, Anne Kemmling, Birgit Pfeiffer, Rolf Daniel, Michael Hoppert

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Biological Soil Crusts (BSCs), consisting of prokaryotes, microalgae, lichens, mosses and eventually small vascular plants, cover wide areas in arid and semi-arid environments. In the present study, the microbial diversity of these crusts was explored at extrazonal mountain steppe sites in the western Khentej (Northern Mongolia). At the study site the Siberian taiga borders on the Mongolian-Daurian forest steppe, resulting in a unique intermixture of the dark taiga, the light taiga, and forest steppe (DULAMSUREN 2004). Due to the presence of boreal, temperate and dauric elements the forest steppe is eminently rich in species (MÜHLENBERG et al. 2004).

BSCs in …

Indicators Of Pasture Digression In Steppe Ecosystems Of Mongolia, Elena V. Danzhalova, S. N. Bazha, P. D. Gunin, Yu. I. Drobyshev, T. I. Kazantseva, A. V. Prischepa, N. N. Slemnev, E. Ariunbold

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The research shows that widely used key measures of vegetation structure (species diversity, projected cover and above-ground phytomass) are not always suitable as indicators of pasture degradation. Based on an analysis above-ground phytomass composition, new quantitative indices are offered that give a more realistic picture of rangeland condition in Mongolia.

Some Taxonomic Records Of Aquatic Insects In The Eroo River Basin (West Khentii, Northern Mongolia), Purevdorj Surenkhorloo, Ravchig Samiya, Jolanta Slowik, Michael Mühlenberg

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In the Eroo basin of Mongolia, many species of aquatic insects have been recorded including five species of Diptera in the families Psychodidae, Dixidae, and Blephaceridae and 25 species across eight families of stoneflies. One species of Psychodidae, Bazarella baikalensis Wagner is reported for the first time in Mongolia while the other families are discussed and a species list is provided for the country.

First Stable Isotope Analysis Of Asiatic Wild Ass Tail Hair From The Mongolian Gobi, Micha Horacek, Petra Kaczensky, Merina Burnik Sturm

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Stable isotope analysis has become a powerful tool to study feeding ecology, water use or movement pattern in contemporary, historic and ancient species. Certain hair and teeth grow continuously, and when sampled longitudinally can provide temporally explicit information on dietary regime and movement pattern. In an initial trial, we analysed a tail sample of an Asiatic wild ass (Equus hemionus) from the Mongolian Gobi. We found seasonal variations in H, C, and N isotope patterns, likely being the result of temporal variations in available feeds, water supply and possibly physiological status. Thus stable isotope analysis shows promise to …

Ectoparasites Of Bats In Mongolia, Part 2 (Ischnopsyllidae, Nycteribiidae, Cimicidae And Acari), Ingo Scheffler, Dietrich Dolch, Jargalsaikhan Ariunbold, Annegret Stubbe, Andreas Abraham, Klaus Thiele

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

This study analyses ectoparasites found on Mongolian bats between 2008 and 2011. We examined 12 different bat species, with a total of 23 ectoparasite species present. Apart from reporting distributions, we also discuss specific host-parasite relationships. Owing to recent taxonomic changes splitting the Myotis mystacinus-group into several new taxa, their corresponding ectoparasite fauna could also be addressed in detail. Introducing ectoparasitic insects at length elsewhere (Scheffler et al. 2010), this paper focuses on the analysis of parasitic Acari. Additional findings for Spinturnicidae (wing mites) and Macronyssidae broadened the spectrum of known parasites. Altogether, the knowledge of bat ectoparasites from Mongolia …

Siphonaptera Of Mongolia And Tuva: Results Of The Mongolian- German Biological Expeditions Since 1962 – Years 1999-2003, Daniel Kiefer, Michael Stubbe, Annegret Stubbe, Scott Lyell Gardner, D. Tserenorov, R. Samiya, D. Otgonbaatar, D. Sumiya, Matthias S. Kiefer

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

This report provides an overview of flea species and the corresponding hosts in Mongolia and Tuva during 1999-2003. The taxonomical development of this vector group of great medical importance covers more than a century of flea research in Mongolia, resulting in the current number of 162 species known from this area.

Creating A Habitat Suitability Index Model For Argali Sheep (Ovis Ammon), And Siberian Ibex (Capra Sibirica) In Ikh Nartin Reserve, Mongolia, Nanette Bragin

Geography and the Environment: Graduate Student Capstones

Mongolia is a country with a park system of protected areas, preserving native flora and fauna. In order to inform decision makers about the status of endangered species and park land health, two species of concern, the Argali sheep (Ovis ammon) and Siberian ibex Capra siberica) in the Ikh Nart Reserve in Mongolia were studied. A Habitat Suitability Index Model was developed to study the Argali and ibex habitat preference and discern if the reserve and core zone offer optimal habitat. A total of 57 animals were examined, creating 95% and 50% home range kernels, and minimum convex polygons. The …

Negotiating Everyday Islam After Socialism: A Study Of The Kazakhs Of Bayan-Ulgii, Mongolia, Namara Brede

Geography Honors Projects

Using ethnographic interviews and participant observations from the Kazakh community of Bayan-Ulgii, Mongolia in June 2009, this study examines how Islamic discourses, practices, experiences, and scales of influence are negotiated in post-socialist Central Asia. To do this, local, national, and transnational dynamics of Mongolian Kazakh religious practice are considered alongside the individual-scale mediating roles of personal preference, social position, life course, power, and social networks. Islam in Bayan-Ulgii is shown to be integral to community and ethnic identity but also multifaceted, dynamic, and multi-scalar, militating against essentialist portrayals of Islam as monolithic or dichotomously split between “high” and “low” forms.

Comparative Morphology Of Two Sympatric Species Of Hedgehog In Ikh Nart Nature Reserve, Mongolia, Richard P. Reading, David Kenny, Sodnomphil Batdorj, James Murdoch

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Daurian hedgehog (Mesechinus dauuricus) and long-eared hedgehog (Hemiechinus auritus) inhabit Ikh Nart Nature Reserve, Mongolia and both species appear to occupy similar niches. We gathered morphological measurements to test for differences between species and sexes. We hoped to gain insight into the mechanisms that allow the species to coexist. We collected morphometric data from 10 long-eared hedgehogs (6 male, 4 female) and 18 Daurian hedgehogs (10 male, 7 female, and 1 unknown sex). Only total body length in Daurian hedgehogs differed significantly between males and females. Otherwise, we found no significant differences between morphometric measures of …

Estimates Of Toad Headed Agama Density In Three Steppe Habitats Of Mongolia, James Murdoch, Buyandelger Suuri, Richard P. Reading

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The toad headed agama (Phrynocephalus versicolor) ranges across the arid steppe regions of southern and western Mongolia and represents one of the most common reptiles in the country. However, few details of the species’ population characteristics exist, which may be important for evaluating population trends and assessing its conservation status. We estimated toad headed agama density in Ikh Nart Nature Reserve, Mongolia based on surveys conducted in the summers (June-September) of 2005, 2006, 2008, and 2009. We conducted surveys in three steppe habitats including tall grassland (n = 7), shrub-steppe (n = 7), and semi-shrub steppe …

Temporal Dynamics Of Group Size And Sexual Segregation In Ibex, N. J. Singh, S. Amgalanbaatar, Richard P. Reading

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Group size is an important variable describing behavioral ecology of animals. A variety of factors such as habitat characteristics, life history, spatio-temporal resource dynamics, population density, predation risk, competition with kin, and social learning often determine group size in large mammals. We studied temporal dynamics of group size in Siberian ibex (Capra sibirica) in a protected area in Mongolia. We measured monthly and yearly variations in typical group size and used the sexual segregation and aggregation statistic to assess sexual segregation. Ibex formed the largest groups in November and smallest groups in July. However, group sizes did not …

Waterbirds Of Uvs-Nuur Depression, T. P. Archimaeva-Ozerskaya, V. I. Zabelin

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

This report provides an overview of water birds (Non-Passeriformes) inhabiting the lakes of Uvs- Nuur Depression. The article includeds material obtained during field work in the Uvs-Nuur Depression during 1999-2006, unpublished observations of rare vagrant species of birds, and data from the literature. Altogether we and others recorded 81 bird species from ten families on the lakes within the basin. The list we provide does not include sparrows or birds from habitats closely associated with the basin. The following non-Passeriformes bird families were noted (with the number of species in parentheses): Gaviidae (1), Podicipedidae (2), Phalacrocoracidae (1), Ardeidae (5), Anatidae …

In Memoriam, Academician Prof. Dr. Osor Shagdarsuren (1929-2010), Sundev Gombobaatar, D. Sumiya, R. Samiya, B. Bayartogtokh, S. Shar, Nyamsuren Batsajchan, M. Altantsetseg, Ch. Uuganbayar, D. Usukhjargal, P. Amartuvshin, P. Jargalsaikhan, Sh. Boldbaatar, N. Tseveenmyadag, G. Mainjargal, B. Nyambayar, E. R. Potapov, Michael Stubbe, Annegret Stubbe, Andrew Dixon

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Academician, Professor Osor Shagdarsuren passed away due to apoplexy on Tuesday, February 2, 2010, at the age of 81. He was one of the most respected Mongolian ornithologists, biologists, and educators. The Mongolian scientific community has lost one of its greatest members, the premier Mongolian ornithologist.

Ecology Of Eurasian Black Vultures (Aegypius Monachus) In Ikh Nart Nature Reserve, Mongolia, Richard P. Reading, David Kenny, John Azua, Travis Garrett, Mary Jo Willis, Tsolmonjav Purevsuren

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

We studied the nesting ecology and movement patterns of Eurasian black vultures (Aegypius monachus) nesting in and near Ikh Nart Nature Reserve, Dornogobi Aimag, Mongolia. From 2003 till 2009, we monitored nesting pairs from incubation to fledging and compared nesting success for pairs nesting on rocks and trees. We captured chicks just prior to fledging to apply leg bands and wing tags. We also captured adults in June and attached solar-powered global positioning system (GPS)/satellite telemetry units, leg bands, and wing tags. We collected resighting data on marked birds opportunistically and telemetry data on tagged birds. We monitored …

Brutareale Und Brutbiologie Der Greifvogelarten Der Mongolei = Grid Mapping And Breeding Ecology Of Raptors In Mongolia, Michael Stubbe, Annegret Stubbe, Nayamsuren Batsaikhan, Sundev Gombobaatar, T. Stenzel, H. Von Wehrden, Sh. Boldbaatar, B. Nayambayar, D. Sumjaa, Ravčigijn Samjaa, N. Ceveenmjadag, A. Bold

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

This work summarizes the longtime ecological research of the German-Mongolian scientific cooperation regarding biodiversity studies in Central Asia, focusing on native raptor species (Aves: Falconiformes). There is included a short overview on the history of raptor research in Mongolia. One of the primary goals was the creation of distribution maps of breeding records based on definitive time and space coordinates. Additional data on the breeding biology amend the distribution data. Currently 43 raptor species are recorded for Mongolia. Whenever possible were also incorporated data from adjacent regions of Mongolia (China, Tuva, Burjatia, Pribaikalia), in order to embed the avifauna of …

Einige Bemerkungen Zur Biologie Der Wüstenkröte (Bufo Raddei), Ellen Driechciarz, René Driechciarz

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Some localities in the Taatsyn river in the valley of the Great Lakes in the South of Changaj in Mongolia were found to harbor Bufo raddei. The authors collected data on population density and tadpole holes of the larvae, collected for the first time from this Asian amphibian species.