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Articles 1 - 30 of 58
Full-Text Articles in Paleontology
A Large And Robust Legless Lizard (Squamata, Anguinae) From The Gray Fossil Site Of Eastern Tennessee, Justice D. Lamer
A Large And Robust Legless Lizard (Squamata, Anguinae) From The Gray Fossil Site Of Eastern Tennessee, Justice D. Lamer
Electronic Theses and Dissertations
The early Pliocene Gray Fossil Site in Tennessee preserved an abundance of microfossils, including numerous anguid lizard specimens. This includes the following elements: dentaries, a pterygoid fragment, a jugal, frontals, parietal fragments, dorsal and caudal vertebrae, and many osteoderms. Because these specimens had not been placed taxonomically beyond the subfamily level, comparative anatomy of similar elements from extant anguids and some quantitative data were used to provide more refined identifications. Using these methods, it was determined that a new, robust type of Ophisaurus is represented at the Gray Fossil Site. Important novel features include relatively robust dentaries, fewer but broader …
Estimating Body Size And Biogeography In The Genus Alligator, Madelyn M. Turala
Estimating Body Size And Biogeography In The Genus Alligator, Madelyn M. Turala
Electronic Theses and Dissertations
Several studies have examined correlations between total body length (TL) and skeletal measurements in the American alligator (Alligator mississippiensis), with some measurements serving as stronger predictors of TL than others. Femur length (FL) has been found to be tightly correlated with TL in A. mississippiensis, and this relationship has been used to estimate body size of extinct crocodilians. This thesis demonstrates that the humerus, radius, ulna, and tibia are also strongly correlated with TL and useful for body size estimation in A. mississippiensis, and thus, can be applied to related extinct crocodilians. Additionally, species distribution modeling …
On The Paleontology Of Belize, Alson Ovando
On The Paleontology Of Belize, Alson Ovando
Electronic Theses and Dissertations
Paleontology can be considered an underrepresented and emerging area of science in Belize. A primary aim of this thesis is to provide a background of paleontology in Belize and present the first comprehensive study of Quaternary vertebrate fossils known from the country. Quaternary fossil sites across Belize include caves, rivers, and sinkholes. In addition to reviewing and assessing material from known sites, this thesis presents new information and discoveries that were made while investigating Belize’s Quaternary fossil record. This thesis also includes a geospatial model depicting ecoregions in Belize during the last glacial maximum. Overall, the results presented here demonstrate …
Ontogeny And The Effects Of Perinatal Stress In The First Immature Skeleton Of Teleoceras Aepysoma (Perissodactyla, Rhinocerotidae), Joe Gajewski
Electronic Theses and Dissertations
Recovery of an immature fossil rhinoceros (Teleoceras aepysoma) skeleton (ETMNH 32999) from the Gray Fossil Site in Tennessee offers the first insights into the ontogeny of the species. ETMNH 32999 is interpreted to be a subadult male in an age class equivalent to ~17-20% of potential lifespan. Comparative analyses support the hypothesis that subadults of related taxa are typically ~91% of average adult size in most dimensions. However, certain cranial dimensions are substantially lower than expected in ETMNH 32999, averaging only ~74% of the average conspecific adult. Such departure from typical growth rates is suggested to be attributable …
Inferring Feeding Biology In Extinct Shrews (Soricidae: Mammalia): Application Of A Linear Morphometric Approach, Derek J. Den Ouden
Inferring Feeding Biology In Extinct Shrews (Soricidae: Mammalia): Application Of A Linear Morphometric Approach, Derek J. Den Ouden
Electronic Theses and Dissertations
Linear morphometrics have been used extensively to infer the ecology of fossil organisms, but shrews have not yet been thoroughly explored using this technique. Often considered to have a homogenous insectivorous diet, direct dietary observation suggests shrew dietary diversity is substantial. Diversity of this nature is well established to have influenced skull morphology in other mammal clades, so similar patterns are hypothesized to be detectable in shrews. To test this, I examined the link between known diet and morphological disparity in extant shrews using linear morphometrics and then utilized that framework to infer the dietary ecology of several fossil taxa. …
Cranial Ontogeny Of The Extinct Dwarf Tapir Tapirus Polkensis; With Comparison To That Of The Extant Tapirus Bairdii (Tapiridae, Perissodactyla), Johannah Orendorff
Cranial Ontogeny Of The Extinct Dwarf Tapir Tapirus Polkensis; With Comparison To That Of The Extant Tapirus Bairdii (Tapiridae, Perissodactyla), Johannah Orendorff
Electronic Theses and Dissertations
Tapirus polkensis is the smallest known species of tapir to ever live, and the largest deposit of their fossils is currently located at the GFS in Gray, TN. Phylogenetics suggest that the extant T. bairdii may be the sister taxon to T. polkensis, although they differ greatly in their morphology. Variation exists among adult T. polkensis specimens, with patterns not present in T. bairdii. For this study, as series of 2D GMAs, with TPSs, were rendered on 10 skulls of T. polkensis and 48 skulls of T. bairdii, across both species’ ontogenetic series. Analyses confirmed that growth …
Pathology And Osteological Observations Of Early Pliocene Rhinoceros, Teleoceras Aepysoma (Perissodactyla, Rhinocerotidae) From Gray Fossil Site, Tennessee, Thomas Scaife
Electronic Theses and Dissertations
Rhinoceroses were an important part of North America’s Paleogene and Neogene ecosystems, with Teleoceras aepysoma being one of the last representatives of this family. Specimens of T. aepysoma from the Gray Fossil Site (GFS) possess distinct/peculiar pathologies: including a pair of fused ribs and ankylosed phalanges. A qualitative description of the pathologies in the GFS T. aepysoma, including new material, was conducted to accurately identify pathologies and make interpretations about the life history of the GFS rhinos. Analysis suggests that rheumatoid arthritis is common in the lower limb bones of GFS rhinos. Additionally, the rib and toe pathologies are more …
Descriptions Of Interglacial Mastodons From Snowmass, Colorado, Connor White
Descriptions Of Interglacial Mastodons From Snowmass, Colorado, Connor White
Electronic Theses and Dissertations
The Ziegler Reservoir fossil site (ZRFS) in Colorado contains over 4000 mastodon bones that date from 140,000 to 100,000 years ago. At an elevation of ~2705 meters above sea level, ZRFS represents an alpine ecosystem dated to Marine Isotope Stage (MIS) 5. Formal descriptions of cheek teeth, mandibles, crania, and femora were completed. Statistical analyses of the upper and lower third molars, including a novel measurement of interloph(id) distances, indicate significant differences between ZRFS mastodons and Mammut pacificus, while falling within the ranges for Mammut americanum. This study agrees with the taxonomic assignment of ZRFS mastodons to Mammut …
Establishing “The Fossil Record”: A Database Of Vertebrate Paleontological Sites Across The State Of Tennessee, Sarah Mclaurine
Establishing “The Fossil Record”: A Database Of Vertebrate Paleontological Sites Across The State Of Tennessee, Sarah Mclaurine
Electronic Theses and Dissertations
Fossil localities across the state of Tennessee and the data related to those sites were compiled from Tennessee Division of Geology Bulletin 84, titled “Tennessee’s Prehistoric Vertebrates,” and stored in a Microsoft Access geodatabase housed by the Department of Collections at the East Tennessee State University Museum of Natural History located at the Gray Fossil Site. Included in the database are forms to enter new site localities, view information about those already entered, view and add data to a master faunal list for the state, view sites repository information and store and add documents that are key-word searchable from the …
Isotopic Ecology Of Bison And Bootherium At Big Bone Lick, Kentucky, Frances Stephenson
Isotopic Ecology Of Bison And Bootherium At Big Bone Lick, Kentucky, Frances Stephenson
Electronic Theses and Dissertations
Big Bone Lick (BBL) is a late Pleistocene and Holocene fossil locality in the Ohio River valley of Kentucky. This study utilized stable isotopes (δ13C, δ18O, and 87Sr/86Sr) in tooth enamel to reconstruct mobility and diet in Bison, as well as diet in Bootherium bombifrons. Isotopic results from seven Bison and two B. bombifrons are reported. Results suggest Pleistocene Bison and Bootherium occupied different dietary niches. Although both had C3-dominated diets, Bison occupied more open environments than B. bombifrons. Two bison from the late Holocene deposit had diets that consisted …
Early Pliocene Mice And Rats From The Gray Fossil Site Of Eastern Tennessee: Implications For The Evolution Of Cricetidae And Understanding Of The Past Ecosystem, Ziqi Xu
Electronic Theses and Dissertations
Cricetidae ranks as the second-most species-rich and abundant mammalian family, with limited studies on eastern North American records prior to the Pleistocene. While cricetids has been previously noted at the early Pliocene Gray Fossil Site (GFS), this study provides a detailed description of eight taxa: Postcopemys (two species), Symmetrodontomys, Oryzomyini, Peromyscus, Neotoma, Neotomodon, and Xenomys. Postcopemys is the most common cricetid taxon at GFS, followed by Peromyscus and Neotoma. These records expand the stratigraphic and geographic range of multiple genera. Distinctive morphological features of GFS taxa suggest presence of several new species. The …
Geometric Morphometric Analysis Of Modern Viperid Vertebrae Facilitates Identification Of Fossil Specimens, Lance D. Jessee
Geometric Morphometric Analysis Of Modern Viperid Vertebrae Facilitates Identification Of Fossil Specimens, Lance D. Jessee
Electronic Theses and Dissertations
Snake vertebrae are common in the fossil record, whereas cranial remains are generally fragile and rare. Consequently, vertebrae are the most commonly studied fossil element of snakes. However, identification of snake vertebrae can be problematic due to extensive variation. This study utilizes 2-D geometric morphometrics and canonical variates analysis to 1) reveal variation between genera and species and 2) classify vertebrae of modern and fossil eastern North American Agkistrodon and Crotalus. The results show that vertebrae of Agkistrodon and Crotalus can reliably be classified to genus and species using these methods. Based on the statistical analyses, four of the …
Age And Paleontology Of The Turin Pit Locality, Monona County, Iowa, Samantha Wright
Age And Paleontology Of The Turin Pit Locality, Monona County, Iowa, Samantha Wright
Electronic Theses and Dissertations
The Turin Pit locality (Monona county, Iowa) has been known to paleontologists since 1908, yet the age of the fauna has been unclear. Early paleontologists considered Turin Pit to date to a pre-Illinoian interglacial (the “Aftonian).” Subsequent researchers suggested it dated to the last glaciation. This study provides a partial list of mammals in the Turin Pit fauna, and together with stratigraphic information, uses the known age ranges of taxa to estimate an age for the assemblage. The presence of Mammuthus, Aenocyon, and Castoroides combined with a magnetically-reversed till located stratigraphically above fossil-bearing deposits, suggest the Turin Pit …
First Skulls Of Arctomeles Dimolodontus (Mustelidae: Melinae) From The Gray Fossil Site (Early Pliocene, Tn) Reveal Extreme Intraspecific Variation, Charles Bruce
Electronic Theses and Dissertations
Here, a morphological description of the extinct meline badger Arctomeles dimolodontus from the Gray Fossil Site (GFS), Tennessee is provided with comparison to extant Meles meles. Originally described solely on upper teeth of a single individual, the recovery of several crania and jaws provides a sample of at least five individuals, affording the largest known fossil sample in North America of the enigmatic Melinae. Examination of A. dimolodontus skull material reveals extreme intraspecific variation in previously identified diagnostic characters, highlighting the need for caution when identifying fossil taxa based on similarly fragmentary remains. Dental variation is likely a result …
Identifying Taphonomic Distribution Patterns At The Gray Fossil Site, Shawn Haugrud
Identifying Taphonomic Distribution Patterns At The Gray Fossil Site, Shawn Haugrud
Electronic Theses and Dissertations
Since the early days of the discovery of the Gray Fossil Site (GFS), meticulous efforts to preserve the spatial data were a priority. Direct surveying of fossils prior to recovery, as well as grid mapping the site, provided relative spatial data within a square meter. Such efforts meant that even fragments and microfossils, recovered during the screening operations and eventual concentrate picking, maintained some spatial data. Available spatial data are used to identify smaller deposits within the greater system, as well as non-random distribution patterns among a number of GFS taxa. Patterns are particularly pronounced in the large-bodied taxa, Teleoceras …
New Reports Of Smilodon And Panthera From North American Cave Sites With Reviews Of Taxonomy, Biogeography, And History, Matthew Bushell
New Reports Of Smilodon And Panthera From North American Cave Sites With Reviews Of Taxonomy, Biogeography, And History, Matthew Bushell
Electronic Theses and Dissertations
In this study, three new reports of large felids from the Pleistocene are presented and discussed. A single partial left dentary from Jawbone Cave represents the first confirmed presence of S. fatalis in east Tennessee and was sampled for radiocarbon dating. Wind Tunnel and Chilly Bowl Cave are among the first records of P. onca to be identified confidently from Arkansas. Chilly Bowl Cave also produced two trackways attributable to Panthera sp., one set potentially belonging to P. atrox. In addition to the description of these remains and the caves they were found in, taxonomic histories for both S. fatalis …
Fossil Mammals From Hickory Tree Cave, Sullivan County, Tennessee, Alexis Rivera
Fossil Mammals From Hickory Tree Cave, Sullivan County, Tennessee, Alexis Rivera
Electronic Theses and Dissertations
Hickory Tree Cave, also known as Big Spring Cave, is located in the southern Appalachians and is known for fossils that are considered to be Quaternary in age. Fossil mammals were identified and assigned to the lowest taxonomical level possible. Most remains are fragmented or digested and it seems likely that various taphonomic processes are responsible for the resulting assemblage. The site lacks the extreme boreal component of Pleistocene cave faunas in the region (e.g. Baker Bluff Cave), with most reported taxa inhabiting Appalachian deciduous forest environments in North America today. While the presence of tapir (Tapirus sp.) …
Community Structure Analysis Of Turtles With Application To The Early Pliocene Gray Fossil Site, Julian Conley
Community Structure Analysis Of Turtles With Application To The Early Pliocene Gray Fossil Site, Julian Conley
Electronic Theses and Dissertations
Turtles are important components of ecosystems around the world, with diverse ecological niches and adaptations. However, there are few detailed studies of how turtle community structure reflects local environments. This project applied techniques of community structure analysis to sites across the United States to infer past ecosystem and environmental conditions of the early Pliocene Gray Fossil Site (GFS) in northeastern Tennessee based on the ancient turtle community. Results indicate extant turtle community structure closely reflects environmental conditions, and that ancient turtle communities can be used to infer climate and habitat conditions of past ecosystems. Application to the GFS turtle community …
The Paleoecology Of High-Elevation Bison In The Greater Yellowstone Ecosystem And Implications For Modern Bison Conservation, Darian Bouvier
The Paleoecology Of High-Elevation Bison In The Greater Yellowstone Ecosystem And Implications For Modern Bison Conservation, Darian Bouvier
Electronic Theses and Dissertations
The national mammal of the United States, the American Bison (Bison bison) was once nearly extinct. Populations have recovered to the degree that thousands roam the Great Plains today. Due to their large numbers and body size, this species has an oversized impact on the ecological communities where it lives and is considered a keystone herbivore in modern North American grasslands. This study explores the detailed, seasonally resolved, paleoecology of seven bison from the Greater Yellowstone Ecosystem during the Late Holocene through stable isotope analyses and species niche modeling. Isotopic analyses of δ13C, δ15N, …
An Evaluation Of Castor Californicus And Implications For The Evolution And Distribution Of The Genus Castor (Rodentia: Castoridae) In North America, Kelly Lubbers
Electronic Theses and Dissertations
The genus Castor is represented in Eurasia by Castor fiber, North America by C. canadensis, and has been in North America since the late Miocene. This study aims to assess whether morphology of Miocene-Pliocene C. californicus and extant C. canadensis are distinctly different. Specimens of Castor were compared using geometric morphometrics on cranial material and linear measurements of postcranial material. Species occurrence data were compared with past and future climate data to assess Castor distribution in North America through time. Results show that C. canadensis is highly variable in both cranial and postcranial morphology and C. californicus falls …
Community Structure Analysis Of Mammals Found At The Gray Fossil Site, Tn, Sarah Clark
Community Structure Analysis Of Mammals Found At The Gray Fossil Site, Tn, Sarah Clark
Electronic Theses and Dissertations
The early Pliocene Gray Fossil Site (GFS) is a biodiverse site with a unique faunal assemblage that represents one of few sites of its age in eastern North America. A community structure analysis of the mammals at GFS was done to characterize species and better understand the paleoenvironment. Data and was gathered from twenty modern communities and five late Neogene sites to compare with GFS. Species from these 26 sites were categorized by body size, locomotor mode, cheek tooth crown height, and diet to characterize niches occupied. Descriptive statistics contrasted proportions of species within categories across communities. Discriminant function analyses …
Distinguishing Mustela From Neogale (Mustelidae) Through Both A Qualitative And Quantitative Analysis Of Skull And Tooth Morphology, Ronald W. Peery
Distinguishing Mustela From Neogale (Mustelidae) Through Both A Qualitative And Quantitative Analysis Of Skull And Tooth Morphology, Ronald W. Peery
Electronic Theses and Dissertations
Weasels and mink (Mustela and Neogale) can be difficult to distinguish osteologically due to similarities in morphology, thus suggesting the need for an accurate tool in distinguishing among taxa. This study utilized a combination of character state and stepwise discriminant function (DFA) analyses to examine potential distinguishing features of skull and tooth morphology. Measurements and ratios were collected from all 18 extant musteline species, as well as the extinct Neovison macrodon, Mustela rexroadensis, Mustela meltoni, Mustela gazini, and Mustela jacksoni. Unidentified musteline specimens from the Gray Fossil Site were also included. Results of …
Spatial Analyses Of Gray Fossil Site Vertebrate Remains: Implications For Depositional Setting And Site Formation Processes, David Carney
Spatial Analyses Of Gray Fossil Site Vertebrate Remains: Implications For Depositional Setting And Site Formation Processes, David Carney
Electronic Theses and Dissertations
This project uses exploratory 3D geospatial analyses to assess the taphonomy of the Gray Fossil Site (GFS). During the Pliocene, the GFS was a forested, inundated sinkhole that accumulated biological materials between 4.9-4.5 mya. This deposit contains fossils exhibiting different preservation modes: from low energy lacustrine settings to high energy colluvial deposits. All macro-paleontological materials have been mapped in situ using survey-grade instrumentation. Vertebrate skeletal material from the site is well-preserved, but the degree of skeletal articulation varies spatially within the deposit. This analysis uses geographic information systems (GIS) to analyze the distribution of mapped specimens at different spatial scales. …
Isotopic Analysis And Mobility Mapping Of Mammuthus Columbi From The Mammoth Site In South Dakota, Matthew Harrington
Isotopic Analysis And Mobility Mapping Of Mammuthus Columbi From The Mammoth Site In South Dakota, Matthew Harrington
Electronic Theses and Dissertations
The Mammoth Site in Hot Springs, South Dakota preserves a unique death assemblage of sub-adult and adult male Columbian mammoths (Mammuthus columbi). Extensive work on the site has led to a detailed understanding of the taphonomy of the assemblage; yet the life histories and ecology of these mammoths remain relatively unknown. Tooth enamel from four Mammoth Site mammoth individuals were bulk sampled with one of the individuals (MSL 742) also serially micro-sampled for 𝛿13C, 𝛿18O, and 87Sr/86Sr. Isotopic results indicate that MSL 742 remained within the southern and western Black Hills …
Morphology And Paleoecology Of Nimravides Galiani (Felidae) And Barbourofelis Loveorum (Barbourofelidae) From The Late Miocene Of Florida, Christianne Ormsby
Morphology And Paleoecology Of Nimravides Galiani (Felidae) And Barbourofelis Loveorum (Barbourofelidae) From The Late Miocene Of Florida, Christianne Ormsby
Electronic Theses and Dissertations
Saber-toothed remains have been found worldwide throughout the Cenozoic, until the end of the Pleistocene. One site from Alachua County, Florida preserves a diverse Miocene fauna, including the machairodontine Nimravides galiani (Felidae) and the saber-toothed Barbourofelis loveorum (Barbourofelidae). Both taxa roamed what would become the Love Bone Bed site during the Late Miocene (Late Clarendonian NALMA), ~ 9.5 Mya. Previous descriptions focused on crania; yet the large sample of postcrania remained undescribed. Hence, this project includes a detailed postcranial description of both taxa. Results show that N. galiani resembles extant felids, whereas B. loveorum resembles Smilodon fatalis, as well …
Gravel Geology And Muskoxen Paleontology Of A Late Pleistocene Fossil Site In Saltville, Virginia, Nickolas A. Brand
Gravel Geology And Muskoxen Paleontology Of A Late Pleistocene Fossil Site In Saltville, Virginia, Nickolas A. Brand
Electronic Theses and Dissertations
Two distinct studies within the Saltville Valley of southwestern Virginia revealed insights into local Pleistocene geology and paleontology. A variety of analytical techniques were applied to gravel deposits within the paleontological site of SV-5/7 that revealed this unit is very poorly sorted, has a subangular matrix, and contains significant components of silt and sand in addition to rounded cobbles. These results suggest that rather than being deposited by fluvial processes as previously suggested, these gravels were likely the result of one or many debris flows. Additionally, the identity of fossil muskoxen from Saltville was reassessed using cranial and dental material. …
Paleoenvironmental Reconstruction Of Quaternary Saltville, Virginia, Using Ostracode Autecology, Austin Gause
Paleoenvironmental Reconstruction Of Quaternary Saltville, Virginia, Using Ostracode Autecology, Austin Gause
Electronic Theses and Dissertations
The Saltville valley in southwestern Virginia is home to Quaternary localities containing paleontological and archaeological remains. Historically the valley has been mined for salt and the small lakes, ponds and springs along the valley floor have a brackish signature. A preliminary report on the site’s ostracode fauna suggested that the site’s water was not always saline. This study analyzed modern and Quaternary ostracodes to understand the valley’s hydrologic and chemical evolution. Sediments contained primarily freshwater species, including the environmentally sensitive Candona crogmaniana. The presence of Pelocypris tuberculatum and a new Fabaeformiscandona species throughout a vertical section spanning the latest Pleistocene …
An Unusual Mastodon Revisited: Providing A Regional Focus For Mammut Americanum (Proboscidea, Mammutidae) In The Southeast, Matthew Inabinett
An Unusual Mastodon Revisited: Providing A Regional Focus For Mammut Americanum (Proboscidea, Mammutidae) In The Southeast, Matthew Inabinett
Electronic Theses and Dissertations
A century ago, two large American mastodon Mammut americanum specimens from coastal South Carolina were mounted for display at Amherst College. Their robust build, broad and pentalophodont third molars, and well-developed mandibular tusks were noted as unusual. Here, these specimens are redescribed; three additional mastodon mandibles from the same region of South Carolina are also described. Though collection information on these specimens is sparse, they appear to span >150 ka in the Middle to Late Pleistocene, and probably include the first report of Mammut from the earliest-Rancholabrean Ten Mile Hill Formation. Some “unusual” features — moderately to extremely broad third …
Ontogenetic And Adult Shape Variation In The Endocast Of Tapirus: Implications For T. Polkensis From The Gray Fossil Site, Thomas M. Gaetano
Ontogenetic And Adult Shape Variation In The Endocast Of Tapirus: Implications For T. Polkensis From The Gray Fossil Site, Thomas M. Gaetano
Electronic Theses and Dissertations
Endocranial morphology provides evidence of sensory ecology and sociality of extinct vertebrates. The Earliest Pliocene Gray Fossil Site (GFS) of NE Tennessee features a conspicuous dominance of skeletal elements belonging to the dwarf tapir, Tapirus polkensis. Numerous individuals in one fossil locality often suggests gregarious behavior, but sociality in T. polkensis contradicts behavior documented for extant Tapirus species. I test T. polkensis for variation in sensory and social ecology using computed tomography and 3D digital endocasts from an ontogenetic sequence. I compare the T. polkensis endocasts with extant Tapirus species using Encephalization Quotients (EQs) and 3D geometric morphometrics. Results …
Encephalization In Commensal Raccoons: A Unique Test Of The Cognitive Buffer Hypothesis, Peter M. Anderson
Encephalization In Commensal Raccoons: A Unique Test Of The Cognitive Buffer Hypothesis, Peter M. Anderson
Electronic Theses and Dissertations
This study investigated selective pressures associated with encephalization in mammals and discussed broader implications. Relative brain size as measured by EQ (Encephalization Quotient) was compared between ecological categories. Omnivores had higher average EQ than ecological specialists. Since specialists are disproportionately affected by extinction events, selection for ecological generalism is proposed as encephalization mechanism. This mechanism may reinforce the more widely known Cognitive Buffer Hypothesis (CBH)—the idea that possessing relatively large brains has buffered lineages against environmental change. CBH is tested here by comparing EQs in Procyon lotor (raccoon) in urban and rural environments. CBH predicts that raccoons in the most …