Paleontology Commons^™

Shallow-Marine Crinoid Genera May Have Been More Resilient Across Permian Extinction Events In Terms Of Richness Than Deep-Marine Crinoid Genera, Amelia C. Kolstad

University Honors Theses

Throughout the Phanerozoic, one of the most successful phyla of marine invertebrates has been Echinodermata. Echinodermata was particularly successful throughout the Paleozoic, with class Crinoidea making up a majority of occurrences. The proportion of echinoderm occurrences that are crinoids declined massively at the end of the Permian and continued to decrease into modern day, never fully recovering. One of the factors thought to have been so important for crinoid success in the Paleozoic and their decreasing success throughout geologic time has been their propensity for success in shallow-marine environments, though crinoids also exist within deep-marine environments. This study seeks to …

Ontogeny And The Effects Of Perinatal Stress In The First Immature Skeleton Of Teleoceras Aepysoma (Perissodactyla, Rhinocerotidae), Joe Gajewski

Electronic Theses and Dissertations

Recovery of an immature fossil rhinoceros (Teleoceras aepysoma) skeleton (ETMNH 32999) from the Gray Fossil Site in Tennessee offers the first insights into the ontogeny of the species. ETMNH 32999 is interpreted to be a subadult male in an age class equivalent to ~17-20% of potential lifespan. Comparative analyses support the hypothesis that subadults of related taxa are typically ~91% of average adult size in most dimensions. However, certain cranial dimensions are substantially lower than expected in ETMNH 32999, averaging only ~74% of the average conspecific adult. Such departure from typical growth rates is suggested to be attributable …

A Multi-Isotope Investigation Of Mammoth Diet, Habitat, And Movement Patterns In The Central Great Plains, U.S.A., Jacob Mccloskey

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

During the Last Glacial Maximum (LGM), the Great Plains in Nebraska consisted of wooded parklands and grasslands, whereas spruce (Picea) forests, parklands, and steppe grasslands dominated east of the Missouri River. I analyzed bulk carbon (δ¹³C), oxygen (δ¹⁸O), and strontium (⁸⁷Sr/⁸⁶Sr) isotopes from tooth enamel in isolated mammoth (Mammuthus) molars to compare diet and movement patterns in two late Pleistocene assemblages: one in Nebraska (n = 24), and one in western Iowa and Missouri (n = 17). Furthermore, I identified a morphometric subset of molars as …

A System And Method For Measuring Spatially Varying Surface Appearances With A Study Of Feathers, Jessica Baron-Lis

All Dissertations

Real-world materials, particularly biological structures such as feathers exhibit complex appearances that vary spatially across their surfaces. The field of computer graphics provides a means of understanding such surfaces through material modeling which uses both analytical models and data acquired from light-surface interactions. There are many efforts within the past decade in measuring materials for graphics, but common limitations in these works include not accounting for spatially varying properties and reliance on neural networks and synthetic datasets.

Feathers from modern birds present diverse appearances due to how light interacts with their unique hierarchical microstructures. Variations in those structures lead to …

Microbial Mats In Dinosaur Ichnocoenoses, Nora Noffke, Paul C. Murphey, Roger C. Buntin, Martin G. Lockley

OES Faculty Publications

Until now, the significance of microbial mats in preservation of dinosaur tracks and in reconstructing the palaeoenvironment in which dinosaurs roamed was rarely studied. Dinosaur tracks are commonly found close to ancient aquatic bodies where moist sediment had once allowed footstep registration. At such sites, epibenthic or endobenthic microbial mats may have developed as well and may have led to exquisite track preservation. Microbial mats in coastal, lacustrine or flood plain environments cause ‘microbially induced sedimentary structures—MISS’, which are micobialites found in sandy or evaporitic settings. The microbial structures record environmental parameters including meteorological conditions that existed at the time …

Biostratigraphic Revision Of Esthonyx (Tillidontia, Mammalia) In The Contect Of Climate Change In The Lower Eocene Of The Bighorn Basin, Wyoming, John Colter Johnson

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Esthonyx is an early Eocene tillodont found in North America, Europe, and India. The genus was named by Edward Drinker (E. D.) Cope in 1874 based on specimens from the San Jose Formation in New Mexico. Since then, several species of Esthonyx have been described from North America alone. The best record of Esthonyx comes from the central Bighorn Basin (BHB) in Wyoming, where it appears in 220 localities in a 640 m-thick stratigraphic succession. Since the last summary of Esthonyx in the BHB, several hundred new specimens have been collected and curated at the Denver Museum of Nature and …

Ecological Dynamics Of Cenozoic North American Mammals, Alexandria Brandis Shupinski

Dissertations and Doctoral Documents, University of Nebraska-Lincoln, 2023–

Understanding how community structure is affected by various ecological, climatic, and environmental changes is a long-standing goal in ecology. However, disentangling human impacts from these other variables is difficult when using short modern timescales and anthropogenically altered communities. The fossil record provides a means to investigate community structure dynamics across critical intervals, excluding humans as a variable. Here, in three studies, I assess changes in North American mammal paleocommunity structure across the last 66 million years (Ma) at multiple temporal and spatial scales. In Chapter 1, I identify changes in mammal functional diversity (FD) locally and continentally, using three independent …

Pathology And Osteological Observations Of Early Pliocene Rhinoceros, Teleoceras Aepysoma (Perissodactyla, Rhinocerotidae) From Gray Fossil Site, Tennessee, Thomas Scaife

Electronic Theses and Dissertations

Rhinoceroses were an important part of North America’s Paleogene and Neogene ecosystems, with Teleoceras aepysoma being one of the last representatives of this family. Specimens of T. aepysoma from the Gray Fossil Site (GFS) possess distinct/peculiar pathologies: including a pair of fused ribs and ankylosed phalanges. A qualitative description of the pathologies in the GFS T. aepysoma, including new material, was conducted to accurately identify pathologies and make interpretations about the life history of the GFS rhinos. Analysis suggests that rheumatoid arthritis is common in the lower limb bones of GFS rhinos. Additionally, the rib and toe pathologies are more …

Hunting Types In North American Eocene And Oligocene Carnivores And Implications For Nimravid Extinction, Miguel Andres Castellanos

Graduate Research Theses & Dissertations

The late Eocene and Oligocene is traditionally seen as a time of great climactic shift, with open habitats beginning to appear and forests receding. This time period saw both the diversification and subsequent extinction of North American nimravids. The causes of the extinction of this early sabertooth group in the Arikareean remain unclear. One possibility for their decline is that nimravids were competitively disadvantaged in open habitats relative to contemporary, possibly more cursorial caniform predators. Here I analyze and compare the hunting styles of Eocene and Oligocene predators in correlation with their ecological implications via discriminant function analysis of the …

New Species Of Dryolestoid From The Late Cretaceous Allen Formation And Implications For South American Faunal Diversity., Brigid Erin Connelly

Electronic Theses and Dissertations

Dryolestoids are extinct cladotherians mammals from the Jurassic and Cretaceous periods. I describe a collection of dryolestoid specimens from the Late Cretaceous localities of Cerro Tortuga (Allen Formation), Anfiteatro 1, and Shining (both La Colonia Formation) from Patagonia, Argentina. Using comparative morphology, I identify a new species of meridiolestidan dryolestoid based on eleven specimens across both formations. The new species’ recovery from La Colonia Formation represents the first dryolestoid connection between the two approximately contemporaneous formations. The species’ morphology may represent an ecological shift within Meridiolestida from insectivory to herbivory, showing a transition in characters between the plesiomorphic sharp-toothed meridiolestidans …

New Reports Of Smilodon And Panthera From North American Cave Sites With Reviews Of Taxonomy, Biogeography, And History, Matthew Bushell

Electronic Theses and Dissertations

In this study, three new reports of large felids from the Pleistocene are presented and discussed. A single partial left dentary from Jawbone Cave represents the first confirmed presence of S. fatalis in east Tennessee and was sampled for radiocarbon dating. Wind Tunnel and Chilly Bowl Cave are among the first records of P. onca to be identified confidently from Arkansas. Chilly Bowl Cave also produced two trackways attributable to Panthera sp., one set potentially belonging to P. atrox. In addition to the description of these remains and the caves they were found in, taxonomic histories for both S. fatalis …

Rapid Growth In Late Cretaceous Sea Turtles Reveals Life History Strategies Similar To Extant Leatherbacks, Laura E. Wilson

Geosciences Faculty Publications

Modern sea turtle long bone osteohistology has been surprisingly well-studied, as it is used to understand sea turtle growth and the timing of life history events, thus informing conservation decisions. Previous histologic studies reveal two distinct bone growth patterns in extant sea turtle taxa, with Dermochelys (leatherbacks) growing faster than the cheloniids (all other living sea turtles). Dermochelys also has a unique life history compared to other sea turtles (large size, elevated metabolism, broad biogeographic distribution, etc.) that is likely linked to bone growth strategies. Despite the abundance of data on modern sea turtle bone growth, extinct sea turtle osteohistology …

The Anatomy And Phylogeny Of A New Large Plioplatecarpine Mosasaur From The Campanian Bearpaw Shale Of Montana (Usa), Richard A. Carr

Master's Theses or Doctor of Nursing Practice

In 2018, a large and associated plioplatecarpine mosasaur skull, pectoral girdle, and rib cage, whose total body length may have exceeded five meters, was uncovered in the Late Campanian Bearpaw Shale of Northeast Montana (USA). Phylogenetic analysis of this specimen, MOR 10855, recovers this individual as a basal member of the genus Plioplatecarpus. This specimen, is unique in that it is estimated to be nearly twice the size of any of the other species of Plioplatecarpus found in the Western Interior Seaway during this part of the Cretaceous. While the included phylogenetic study suggests MOR 10855 represents a new …

Timing Of Diversification, Dispersal, And Biogeography Of Parrots In The Genus Amazona (Psittaciformes: Psittacidae) Throughout The Caribbean, Visualized In Gis, Christopher Kingwill

Master's Theses or Doctor of Nursing Practice

Avian fossil records from across the Caribbean (Greater and Lesser Antilles) demonstrate higher avian diversity prior to extinction events due to climate change at the end of the Pleistocene and human impact across the Caribbean throughout the Holocene. Amazon parrots (Amazona) are a diverse genus of New World parrots found throughout Central and South America, as well as the Caribbean. Their phylogeny and evolutionary history, specifically for Caribbean species, has been debated in terms of source areas in Central and South America and the timing of and number of colonization events to different islands that preceded diversification into …

Two New Late Quaternary Avifaunas From The East-Central Great Basin With The Description Of A New Species Of Falco, Steven D. Emslie, Jim I. Mead

Western North American Naturalist

We report 2 new late Quaternary avifaunas from cave deposits in Snake Valley, White Pine County, eastern Nevada, that provide new records of extinct and extant species in the east-central Great Basin. These avifaunas collectively represent at least 24 species, including the most northern inland record of California Condor (Gymnogyps californianus) in western North America and a new species of kestrel (Falco sp.) described herein. Two extralimital living species also were identified—the Harlequin Duck (Histrionicus histrionicus) and Snowy Owl (Bubo scandiacus)—and are the first fossil records of these species in the Great Basin. …

Reconstructing The Ecological Relationships Of Late Cretaceous Antarctic Dinosaurs And How Functional Tooth Morphology Influenced These Relationships, Ian D. Broxson

2022 Symposium

The Sandwich Bluff Formation of the James Ross Basin of Antarctica has recently yielded a group of five late Cretaceous dinosaurs that lived contemporaneously with each other, a first for Antarctica. These five dinosaurs include fragmentary remains of two differently sized elasmarian ornithopods, a possible megaraptor, a hadrosaur, and a nodosaur. In this study we will construct a model of the ecological relationships of late Cretaceous Antarctica. Additionally, we will look at what specific factors allowed this group of four herbivores and a carnivore to coexist in a restricted locality and what niches were filled by each species. Methods to …

Terrestrial Soldier Crab (Coenobita Clypeatus, Fabricius 1787) And Cerion Spp. (Röding 1798) Shell Relationship On San Salvador Island, Bahamas, Harley Hunt

Biology Theses

The Caribbean terrestrial soldier crab, Coenobita clypeatus(Fabricius 1787), coexist and utilize the shells of numerous species of land and marine gastropods. Soldier crabs rely on gastropod shells for protection as the crabs have a soft abdomen, leaving them vulnerable for predation and desiccation, threatening their survival. This creates a strong pressure to obtain well-fitting shells that provide adequate protection against water loss. Cerion of Röding (1798) shells are one of the most commonly used shells among living colonies of C. clypeatuson San Salvador Island. This study is interested in the frequency of shell use by C. clypeatus crabs …

Holocene Records Of Nebraska Mammals, Hugh H. Genoways

Zea E-Books Collection

A survey of the archeological and paleontological literature allowed a compilation of Holocene records of mammals in Nebraska. This survey identified Holocene records from 338 sites in 62 of the 93 Nebraska counties. These counties were located throughout state, but there was a concentration of sites in southwestern Nebraska where there were 27 fossil sites in Frontier County and 22 in Harlan County. Fossils sites were underrepresented in the Sand Hills region. Records of fossil mammals covered the entire Holocene period from 13,000 years ago until AD 1850. A minimum of 57 species (with eight additional species potentially present) representing …

An Exceptionally Small New Polycotylid Plesiosaur (Reptilia: Sauropterygia) With Raptorial Eyes From The Western Interior Seaway Of North America, Robert O’Brien Clark

Theses, Dissertations and Capstones

Polycotylidae is a family of plesiosaurian marine reptiles that evolved during the Early Cretaceous and radiated into multiple genera during the Late Cretaceous, achieving a worldwide distribution. Derived polycotylids of the subclade Polycotylinae have a gracile and elongated rostrum, homodont dentition, an extended mandibular symphysis, and foreshortened temporal fenestrae. In this thesis, I describe a small and highly derived new polycotylid taxon based on three specimens from the Campanian of the Western Interior Seaway in North America. A high number of maxillary teeth, fused neural arches, propodials with well-defined facets, and heavily remodeled cortical bone indicate the specimens are adults, …

Ontogenetic And Adult Shape Variation In The Endocast Of Tapirus: Implications For T. Polkensis From The Gray Fossil Site, Thomas M. Gaetano

Electronic Theses and Dissertations

Endocranial morphology provides evidence of sensory ecology and sociality of extinct vertebrates. The Earliest Pliocene Gray Fossil Site (GFS) of NE Tennessee features a conspicuous dominance of skeletal elements belonging to the dwarf tapir, Tapirus polkensis. Numerous individuals in one fossil locality often suggests gregarious behavior, but sociality in T. polkensis contradicts behavior documented for extant Tapirus species. I test T. polkensis for variation in sensory and social ecology using computed tomography and 3D digital endocasts from an ontogenetic sequence. I compare the T. polkensis endocasts with extant Tapirus species using Encephalization Quotients (EQs) and 3D geometric morphometrics. Results …

Encephalization In Commensal Raccoons: A Unique Test Of The Cognitive Buffer Hypothesis, Peter M. Anderson

Electronic Theses and Dissertations

This study investigated selective pressures associated with encephalization in mammals and discussed broader implications. Relative brain size as measured by EQ (Encephalization Quotient) was compared between ecological categories. Omnivores had higher average EQ than ecological specialists. Since specialists are disproportionately affected by extinction events, selection for ecological generalism is proposed as encephalization mechanism. This mechanism may reinforce the more widely known Cognitive Buffer Hypothesis (CBH)—the idea that possessing relatively large brains has buffered lineages against environmental change. CBH is tested here by comparing EQs in Procyon lotor (raccoon) in urban and rural environments. CBH predicts that raccoons in the most …

An Assessment Of Convergence In The Feeding Morphology Of Xiphactinus Audax And Megalops Atlanticus Using Landmark-Based Geometric Morphometrics, Edward Chase Shelburne

Master's Theses or Doctor of Nursing Practice

Convergence is an evolutionary phenomenon wherein distantly related organisms independently develop features or functional adaptations to overcome similar environmental constraints. Historically, convergence among organisms has been speculated or asserted with little rigorous or quantitative investigation. More recent advancements in systematics has allowed for the detection and study of convergence in a phylogenetic context, but this does little to elucidate convergent anatomical features in extinct taxa with poorly understood evolutionary histories. The purpose of this study is to investigate one potentially convergent system—the feeding structure of Xiphactinus audax (Teleostei: Ichthyodectiformes) and Megalops atlanticus (Teleostei: Elopiformes)—using a comparative anatomical approach to assess …

Reorganization Of Surviving Mammal Communities After The End-Pleistocene Megafaunal Extinction, Anikó B. Tóth, S. Kathleen Lyons, W. Andrew Barr, Anna K. Behrensmeyer, Jessica L. Blois, René Bobe, Matt Davis, Andrew Du, Jussi T. Eronen, J. Tyler Faith, Danielle Fraser, Nicholas J. Gotelli, Gary R. Graves, Advait M. Jukar, Joshua H. Miller, Silvia Pineda-Munoz, Laura C. Soul, Amelia Villaseñor, John Alroy

School of Biological Sciences: Faculty Publications

Large mammals are at high risk of extinction globally. To understand the consequences of their demise for community assembly, we tracked community structure through the end- Pleistocene megafaunal extinction in North America.We decomposed the effects of biotic and abiotic factors by analyzing co-occurrence within the mutual ranges of species pairs. Although shifting climate drove an increase in niche overlap, co-occurrence decreased, signaling shifts in biotic interactions. Furthermore, the effect of abiotic factors on cooccurrence remained constant over time while the effect of biotic factors decreased. Biotic factors apparently played a key role in continental-scale community assembly before the extinctions. Specifically, …

Body Size Downgrading Of Mammals Over The Late Quaternary, Felisa A. Smith, Rosemary E. Elliott Smith, S. Kathleen Lyons, Jonathan L. Payne

School of Biological Sciences: Faculty Publications

Since the late Pleistocene, large-bodied mammals have been extirpated from much of Earth. Although all habitable continents once harbored giant mammals, the few remaining species are largely confined to Africa. This decline is coincident with the global expansion of hominins over the late Quaternary. Here, we quantify mammalian extinction selectivity, continental body size distributions, and taxonomic diversity over five time periods spanning the past 125,000 years and stretching approximately 200 years into the future. We demonstrate that size-selective extinction was already under way in the oldest interval and occurred on all continents, within all trophic modes, and across all time …

Late Quaternary Dragon Lizards (Agamidae: Squamata) From Western Australia, Julie Rej

Electronic Theses and Dissertations

Fossil Agamidae from Western Australia have been the subject of limited study. To aid in fossil agamid identification, Hocknull (2002) examined the maxilla and dentary of several extant species from Australia and determined diagnostic characters for various species groups. In the study here, fossil agamids from two localities in Western Australia, Hastings Cave and Horseshoe Cave, were examined, grouped, and identified to the lowest unambiguous taxonomic level. Morphometric analyses were conducted to compare morphotypes, and find additional diagnostic characters. From Hastings Cave there were two maxilla morphotypes and three dentary morphotypes. Based on identifications, taxa present at this locality were …

Description Of Cranial Elements And Ontogenetic Change Within Tropidolaemus Wagleri (Serpentes: Crotalinae)., Nicolette L. Hill

Electronic Theses and Dissertations

Tropidolaemus wagleri is a species of Asian pitviper with a geographic range including Thailand, Vietnam, Malaysia, Singapore, Bruniei, parts of Indonesia, and the hilippines. Tropidolaemus is a member of the Crotalinae subfamily, within Viperidae. The genus Tropidolaemus includes five species, and was once included within the genus Trimeresurus. While some osteologic characteristics have been noted a comprehensive description of cranial elements has not been produced for T. wagleri. An in-depth description of the cranial skeleton of Tropidolaemus wagleri lays the foundation for future projects to compare and contrast other taxa within Crotalinae and Viperidae. The chosen reference specimen …

Analyzing Pterosaur Ontogeny And Sexual Dimorphism With Multivariate Allometry, Erick Charles Anderson

Theses, Dissertations and Capstones

The relationships of pterosaurs have been previously inferred from observed traits, depositional environments, and phylogenetic associations. A great deal of research has begun to analyze pterosaur ontogeny, mass estimates, wing dynamics, and sexual dimorphism in the last two decades. The latter has received the least attention because of the large data set required for statistical analyses. Analyzing pterosaurs using osteological measurements will reveal different aspects of size and shape variation in Pterosauria (in place of character states) and sexual dimorphism when present. Some of these variations, not easily recognized visually, will be observed using multivariate allometry methods including Principle Component …

The Cova Des Pas De Vallgornera (Llucmajor, Mallorca): A Singular Deposit Bearing An Exceptional Well Preserved Early Pleistocene Vertebrate Fauna, Pere Bover, Alex Valenzuela, Carmen Guerra, Juan Rofes, Josep A. Alcover, Joaquin Gines, Joan J. Fornos, Gloria Cuenca-Bescós, Antoni Merino

International Journal of Speleology

The Cova des Pas de Vallgornera is the longest cave of Mallorca (Balearic Islands, Western Mediterranean) and one of the 30 longest caves in the world. The exploration of one of the galleries allowed the discovery of a fossiliferous deposit of vertebrate remains in a remarkable preservation state. The fossil faunal complex found in this gallery is composed of up to 5 mammalian species (Myotragus aff. kopperi, Hypnomys onicensis, Nesiotites aff. ponsi, Rhinolophus aff. mehelyi and Pipistrellus sp.), at least 14 bird species (among them two Mallorcan endemic taxa: Pica mourerae and Athene vallgornerensis), one …

Shape Evolution And Sexual Dimorphism In The Mandible Of The Dire Wolf, Canis Dirus, At Rancho La Brea, Alexandria L. Brannick

Theses, Dissertations and Capstones

The Rancho La Brea tar pits are a classic fossil lagerstätten, famous for the deposition of episodic accumulations of large numbers of fossils over short intervals in the late Pleistocene. I analyzed 157 Canis dirus (dire wolf) hemi-mandibles from Rancho La Brea through a 2D landmark-based morphometric analysis to test for size and shape changes through time. I scored 16 landmarks on each mandible gathered from four pits of different ages: 61/67 (~13-14 thousand years ago [ka]), 13 (~17-18 ka), 2051 (~26 ka), and 91 (~28 ka). Analyses indicate size does change through time, and shares a broad correlation with …

110 Years Of Avipoxvirus In The Galapagos Islands, Patricia Parker, Elizabeth Buckles, Heather Farrington, Kenneth Petren, Noah Whiteman, Robert Ricklefs, Jennifer Bollmer, Gustavo Jiménez-Uzcátegui

Biology Department Faculty Works

The role of disease in regulating populations is controversial, partly owing to the absence of good disease records in historic wildlife populations. We examined birds collected in the Galapagos Islands between 1891 and 1906 that are currently held at the California Academy of Sciences and the Zoologisches Staatssammlung Muenchen, including 3973 specimens representing species from two well-studied families of endemic passerine birds: finches and mockingbirds. Beginning with samples collected in 1899, we observed cutaneous lesions consistent with Avipoxvirus on 226 (6.3%) specimens. Histopathology and viral genotyping of 59 candidate tissue samples from six islands showed that 21 (35.6%) were positive …