Paleobiology Commons^™

Minerals Rock!: Improving Audience Engagement And Interest In Stem Through Enhancement Of The In-Person And Online Mineral Collection At Ursinus College, Juliette Lyons

Environment and Sustainability Summer Fellows

Museums are a vital source of educational resources for the general public. They provide information on a variety of topics and disciplines and encourage visitors to be excited about and engaged in learning something new. As we work to comprehend key topics such as climate change, it is especially important for the broader public to better understand science, including geology. While the geological collection at Ursinus, Pieces of Earth’s History, is not in a museum, implementing methods that museums use can improve the reception of the collection. Through the application of research gathered through articles, journals, books, and talking with …

Updates To The Caniform Fauna (Carnivora, Caniformia) Of The John Day Formation, Oregon, Usa, John D. Orcutt, Kevin C. Talbott, Michael J. Monroe, Nicholas A. Famoso

Biology Faculty Scholarship

The John Day Formation of central Oregon preserves a diversity of Oligo-Miocene caniform carnivorans, many of which were first described from the formation. However, many historically important specimens lack detailed locality or stratigraphic information, complicating any analysis of faunal change through time. Here, we describe 16 new specimens of caniforms from the John Day Formation, most with precise stratigraphic information. We report multiple specimens of the amphicyonids Daphoenus Leidy, 1853, Paradaphoenus cuspigerus (Cope, 1878), and Temnocyon altigenis (Cope, 1878), all previously known from the formation. We also report the first definitive occurrence of the hesperocyonine canid Osbornodon Wang, 1994 from …

Shallow-Marine Crinoid Genera May Have Been More Resilient Across Permian Extinction Events In Terms Of Richness Than Deep-Marine Crinoid Genera, Amelia C. Kolstad

University Honors Theses

Throughout the Phanerozoic, one of the most successful phyla of marine invertebrates has been Echinodermata. Echinodermata was particularly successful throughout the Paleozoic, with class Crinoidea making up a majority of occurrences. The proportion of echinoderm occurrences that are crinoids declined massively at the end of the Permian and continued to decrease into modern day, never fully recovering. One of the factors thought to have been so important for crinoid success in the Paleozoic and their decreasing success throughout geologic time has been their propensity for success in shallow-marine environments, though crinoids also exist within deep-marine environments. This study seeks to …

Segmented Inner Skull Cavities Of Eothoracosaurus Mississippiensis (Msu 3293), Amy Moe-Hoffman, Renee Clary

Research Data

Dunn-Seiler Museum specimen data:

• Catalog Number: DSM/MSU#3293 HOLOTYPE 
  
• Geographic Locality: Oktibbeha County, MS Rock Hill vicinity 
  
• Geologic Locality: Ripley Formation 
  
• Age: Cretaceous (Maastrichtian)   

Specimen includes Cranium, Mandible, loose teeth, and post-cranial elements.  

1. CT Scan description: The specimen MSU 3293 was scanned at the Mississippi State University College of Veterinary Medicine using a GE LightSpeed VCT (GE Healthcare, Chicago, Illinois, USA; www.gehealthcare.com ) on May 4th 2022, with scan energies of 120kV and 200mA. This produced 1593 images with a slice thickness of 625µm. 
2. 3D models: 3D models of segmented inner skull cavities based on segrmetation of the CT scanned …

On The Paleontology Of Belize, Alson Ovando

Electronic Theses and Dissertations

Paleontology can be considered an underrepresented and emerging area of science in Belize. A primary aim of this thesis is to provide a background of paleontology in Belize and present the first comprehensive study of Quaternary vertebrate fossils known from the country. Quaternary fossil sites across Belize include caves, rivers, and sinkholes. In addition to reviewing and assessing material from known sites, this thesis presents new information and discoveries that were made while investigating Belize’s Quaternary fossil record. This thesis also includes a geospatial model depicting ecoregions in Belize during the last glacial maximum. Overall, the results presented here demonstrate …

Ontogeny And The Effects Of Perinatal Stress In The First Immature Skeleton Of Teleoceras Aepysoma (Perissodactyla, Rhinocerotidae), Joe Gajewski

Electronic Theses and Dissertations

Recovery of an immature fossil rhinoceros (Teleoceras aepysoma) skeleton (ETMNH 32999) from the Gray Fossil Site in Tennessee offers the first insights into the ontogeny of the species. ETMNH 32999 is interpreted to be a subadult male in an age class equivalent to ~17-20% of potential lifespan. Comparative analyses support the hypothesis that subadults of related taxa are typically ~91% of average adult size in most dimensions. However, certain cranial dimensions are substantially lower than expected in ETMNH 32999, averaging only ~74% of the average conspecific adult. Such departure from typical growth rates is suggested to be attributable …

Mumm Vs. Minium: A Tale Of Two Quarries, Kale Link, Amanda Peng

SACAD: Scholarly Activities

In 2025, a discrepancy was discovered in the field numbers of a selection of mammal fossils, which had been mistakenly catalogued as originating from the Minium Quarry in Graham County, KS instead of the Mumm Quarry in Rawlins County, KS. While both quarries belong to the Ogallala Formation, they demonstrate differing biostratigraphic ages, faunal assemblages, and taphonomic settings. Mumm is Clarendonian aged, while Minium is Hemphillian. To correct this issue, all the Mumm specimens were assigned to an updated locality with more precise geographic and biostratigraphic data.

Mandibular Ecomorphology Of Tylosaurus (Reptilia: Mosasauridae), Andrew I. Price, Amanda Peng

SACAD: Scholarly Activities

The large mosasaur Tylosaurus is known to have been a predator of large marine animals. Performing beam theory analysis, a computational technique that uses linear dimensions of a structure to calculate its resistance to bending, on the mandibles of six Tylosaurus specimens from the Smoky Hill Chalk of Kansas indicates dorsoventral and mediolateral bending strength both increase posteriorly along the mandible. Comparisons with Varanus komodoensis indicate relatively greater mediolateral bending strength mid-dentary in Tylosaurus, suggesting that resisting torsional forces of struggling prey was more important at this landmark in Tylosaurus and that more of its jaw length was used …

Overview And Methods Of Molecular Paleontology: A Review, Shelby Touchstone

Williams Honors College, Honors Research Projects

Molecular paleontology—the study of ancient biomolecules preserved in fossils and sediments, including ancient DNA, lipids, and proteins—has motivated researchers since the 1950s, inspiring both scientific inquiry and broader cultural imagination. Think Jurassic Park. This honors thesis presents a comprehensive review of the history, current state, and methods of the field of molecular paleontology. Drawing on peer-reviewed literature and scholarly resources accessed through databases such as EBSCO and Bierce library, this review examines key techniques such as Next Generation Sequencing and amino acid racemization and will go over limitations and controversies in the field. It will also explore potential new …

A Multi-Isotope Investigation Of Mammoth Diet, Habitat, And Movement Patterns In The Central Great Plains, U.S.A., Jacob Mccloskey

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

During the Last Glacial Maximum (LGM), the Great Plains in Nebraska consisted of wooded parklands and grasslands, whereas spruce (Picea) forests, parklands, and steppe grasslands dominated east of the Missouri River. I analyzed bulk carbon (δ¹³C), oxygen (δ¹⁸O), and strontium (⁸⁷Sr/⁸⁶Sr) isotopes from tooth enamel in isolated mammoth (Mammuthus) molars to compare diet and movement patterns in two late Pleistocene assemblages: one in Nebraska (n = 24), and one in western Iowa and Missouri (n = 17). Furthermore, I identified a morphometric subset of molars as …

The Petm: A Window Into Earth's Future Climate, Jacalyn M. Wittmer Malinowski

Climate Change and the Individual, 2025-26

The Paleocene-Eocene Thermal Maximum (PETM) is not just an event that occurred in Earth’s distant past (~55 million years ago) but is a crucial lesson for today’s meteoric climate change. The PETM offers a historical parallel of rapid flux of greenhouse gases into the atmosphere that caused global warming, ocean acidification, dramatic changes in weather, and extinctions. This global warming event was the result of 20,000 years of “rapid” warming (in geological terms) however, we are experiencing rates of warming that are 5-10x higher than PETM levels. We are not experiencing rapid warming today, in comparison to the past, we …

The Human Legacy: Agriculture, Civilizations, Strife, And Pandemics Changing Our Climate, Jacalyn M. Wittmer Malinowski

Climate Change and the Individual, 2025-26

For most of human history, climate has shaped our lives, while our influence on it remained minimal. However, the notion that early humans had no measurable impact on the climate before industrialization is inaccurate. Early in human history, activities – such as the rise of agriculture, the growth of civilizations, periods of conflict, and pandemics – altered greenhouse gas levels in the atmosphere. These changes contributed to significant periods of climate change long before the burning of fossil fuels set us on the path to today’s global climate crisis.

Primates From Senèze, Eric Delson

Publications and Research

Senèze has yielded two primate fossils, neither from the recent field work. A partial ulna can be identified as cf. Macaca sylvanus. A nearly complete female skull is the holotype of Dolichopithecus arvernensis Depéret, 1928, now included in Paradolichopithecus. Emended and differential diagnoses are provided for this genus and species for the first time, and the Senèze skull is described and briefly compared to those of other species of the genus. Dental and cranial measurements are provided, along with comparative data. The macaque and Paradolichopithecus are moderately to highly terrestrial, respectively, and they suggest a woodland to more open …

Sedimentary Archives Of Late Quaternary Limnological, Ecological, And Environmental Change In The Eastern Sierra Nevada (California, Usa), Bailee Hodelka

Theses and Dissertations--Earth and Environmental Sciences

The Sierra Nevada essentially serve as the headwaters of California, providing crucial freshwater to the state through seasonal snowmelt. This water supply is threatened by climate change in complex ways that are not fully understood. Geological records provide a valuable source of information on how the hydrology, limnology, and ecology of the Sierra Nevada region has responded to climate changes of the past. This dissertation presents new sedimentary and paleoecological research from Mono Lake and Convict Lake (Mono County, California) and aims to shed light on environmental changes in the eastern Sierra Nevada from ~16 ka to present.

At Mono …

An Understanding Of The Feeding Ecology Of Teleoceras Fossiger And Teleoceras Aepysoma Using Dental Microwear And Mesowear Analysis, Gabriel S. Goncalves

Master's Theses or Doctor of Nursing Practice

Teleoceras is a genus of rhinocerotids that belongs to the Tribe Teleoceratines and was widespread across North America from the late Early Miocene into the Early Pliocene. This genus was very successful, spreading across the continental United States from Florida and Tennessee to California and Washington state down to central Mexico and Honduras. Teleoceras is characterized by having a large, rounded chest cavity and short limbs with relatively high crowned teeth and lower incisors modified into tusk-like teeth. Initial ecological comparisons were made to the modern hippopotamus based on proportions of the chest cavity and its hypsodont dentition. Stable carbon …

Distribution And Tooth Morphoguilds Of Mosasaurids (Squamata) Of The Campanian Western Interior Seaway With Implications For Controls On Mosasaur Paleobiogeography, Alec Zaborniak

Master's Theses or Doctor of Nursing Practice

The Western Interior Seaway (WIS) of North America is well known for its mosasaurid squamate diversity, particularly during the Campanian of the Late Cretaceous. Mosasaurid diversity has historically been examined at a fine scale, with many studies investigating the faunal composition of specific assemblages; however, studies focusing on mosasaur communities within the Western Interior Seaway during specific Ages are currently lacking. Mosasaur tooth morphology has also been extensively studied, with tooth characteristics commonly being used as phylogenetic characters. However, many studies examining mosasaur communities and tooth morphology have been focused on broader spatiotemporal or phylogenetic scales. This study investigates the …

A Histological Survey Of A Large, Sub-Adult Tylosaurus Nepaeolicus With Implications On The Validity Of Tylosaurus Kansasensis, Carson Cope

Master's Theses or Doctor of Nursing Practice

Tylosaurus nepaeolicus is a basal, medium-sized tylosaurine mosasaur that lived during the late Coniacian and early Santonian in the Western Interior Seaway. Despite being a well-studied species, there is debate over whether Tylosaurus kansasensis represents the juvenile form of T. nepaeolicus, or if the two species are distinct. Osteohistology can determine organism age at time of death and can be used to help resolve this debate and provide a better understanding of the growth of T. nepaeolicus. Thin-sections were taken from a humerus, rib, and vertebra of a single individual (FHSM VP-2209). All thin-sections show primarily parallel-fibered cortical …

A Paleobiological And Paleoecological Analysis Of Mississippian Trilobites With Segmentation Abnormalities, April Catherine Cerami

Graduate Theses, Dissertations, and Problem Reports (ETD)

Trilobites are a well-preserved group of arthropods, with a fossil record extending from the Cambrian to the end of the Permian. Abnormalities, such as injuries or teratological defects, have been observed and described in multiple individuals, although investigating how these abnormalities occur in populations has been largely overlooked.

One possible cause of abnormalities in marine arthropods is heavy metal pollution. Such metals can leave individuals more susceptible to teratological deformities and injuries, interfering with the molting and healing process. Here, methods for analyzing elemental composition and preservation of the trilobite cuticle were refined, finding that 30 μm thin sections are …

Invertebrate Functional Diversity Of Fossils From Arrow Canyon, Nevada, Nathaniel Valentine

Undergraduate Research Symposium Podium Presentations

Paleontology can be used to study how ecosystems evolve and respond to changes in the environment. Here we examine invertebrate fossils from the Pennsylvanian, a time period that ranged from 323.2 to 298.9 million years ago, which are preserved along the walls of Arrow Canyon (Nevada) in the Bird Spring Formation. We surveyed three sites within the canyon, and used data from the Paleobiology Database to create a list of taxa found on the canyon from this time interval. Although ecosystems have traditionally been studied using the composition of the ecosystem (the number of species), recent work has shown that …

Tales In The Teeth: Using Isotopic Analysis To Understand The Paleoecology And Social Behavior Of Brontotheres Of The Intermountain Basins During The Middle Eocene, John Michael Michalski

All Graduate Reports and Creative Projects, Fall 2023 to Present

Isotopic analysis has been a common practice used in paleontology to study ancient ecosystems, and the lives of animals therein. Oxygen isotopic analysis is of particular importance not only because of its use in studying paleoecology, diet, and migration, but also its use in studying mountain elevation, and its impact on ancient ecosystems. I use stable oxygen isotopic analysis to study the habits and ecology of brontotheres from intermountain basins surrounding the Uinta Mountains of Utah and Wyoming, USA. I studied the oxygen isotopes from 16 fossil specimens of teeth, most of which belonged to brontotheres, including the genera Palaeosyops …

Biostratigraphic Revision Of Esthonyx (Tillidontia, Mammalia) In The Contect Of Climate Change In The Lower Eocene Of The Bighorn Basin, Wyoming, John Colter Johnson

Department of Earth and Atmospheric Sciences: Dissertations, Theses, and Student Research

Esthonyx is an early Eocene tillodont found in North America, Europe, and India. The genus was named by Edward Drinker (E. D.) Cope in 1874 based on specimens from the San Jose Formation in New Mexico. Since then, several species of Esthonyx have been described from North America alone. The best record of Esthonyx comes from the central Bighorn Basin (BHB) in Wyoming, where it appears in 220 localities in a 640 m-thick stratigraphic succession. Since the last summary of Esthonyx in the BHB, several hundred new specimens have been collected and curated at the Denver Museum of Nature and …

Controls On Soft Tissue And Cellular Preservation In Late Eocene And Oligocene Vertebrate Fossils From The White River And Arikaree Groups Of Nebraska, South Dakota, And Wyoming, John E. Gallucci, Grace Woolslayer, Kelsey Barker, Brian Kibelstis, Allison R Tumarkin-Deratzian, Paul V. Ullmann, David E. Grandstaff, Dennis O. Terry

School of Earth & Environment Departmental Research

Previous studies on microtaphonomy have identified multiple types of organic microstructures in fossil vertebrates from a variety of time periods and past environmental settings. This study investigates potential taphonomic, paleoenvironmental, and paleoclimatic controls on soft tissue and cellular preservation in fossil bone. To this end, fifteen vertebrate fossils were studied: eight fossils collected from the Oligocene Sharps Formation of the Arikaree Group in Badlands National Park, South Dakota, and seven fossils from formations in the underlying White River Group, including the Oligocene Brule Formation of Badlands National Park, and the Eocene Chadron Formation of Flagstaff Rim, Wyoming; Toadstool Geologic Park, …

Pathology And Osteological Observations Of Early Pliocene Rhinoceros, Teleoceras Aepysoma (Perissodactyla, Rhinocerotidae) From Gray Fossil Site, Tennessee, Thomas Scaife

Electronic Theses and Dissertations

Rhinoceroses were an important part of North America’s Paleogene and Neogene ecosystems, with Teleoceras aepysoma being one of the last representatives of this family. Specimens of T. aepysoma from the Gray Fossil Site (GFS) possess distinct/peculiar pathologies: including a pair of fused ribs and ankylosed phalanges. A qualitative description of the pathologies in the GFS T. aepysoma, including new material, was conducted to accurately identify pathologies and make interpretations about the life history of the GFS rhinos. Analysis suggests that rheumatoid arthritis is common in the lower limb bones of GFS rhinos. Additionally, the rib and toe pathologies are more …

Descriptions Of Interglacial Mastodons From Snowmass, Colorado, Connor White

Electronic Theses and Dissertations

The Ziegler Reservoir fossil site (ZRFS) in Colorado contains over 4000 mastodon bones that date from 140,000 to 100,000 years ago. At an elevation of ~2705 meters above sea level, ZRFS represents an alpine ecosystem dated to Marine Isotope Stage (MIS) 5. Formal descriptions of cheek teeth, mandibles, crania, and femora were completed. Statistical analyses of the upper and lower third molars, including a novel measurement of interloph(id) distances, indicate significant differences between ZRFS mastodons and Mammut pacificus, while falling within the ranges for Mammut americanum. This study agrees with the taxonomic assignment of ZRFS mastodons to Mammut …

Consequences Of The Megafauna Extinction: Changes In Food Web Networks On The Edwards Plateau Across The Pleistocene–Holocene Transition, Quentin A. Smith Jr.

School of Biological Sciences: Dissertations, Theses, and Student Research

We are experiencing biodiversity loss due to climate change and human impacts, which is not only harmful to the environment but can also alter the composition of communities and interactions among species. The late Pleistocene experienced a loss of large-bodied mammals which resulted in significant changes in community structure due to changes in body size, diet, and species associations. The impact of these changes on species interactions and community structure across the Pleistocene-Holocene transition remains poorly understood. Using a robust data set of species composition, stable isotopes, body size, and climate variables, we constructed and compared ecological networks of mammal …

Establishing “The Fossil Record”: A Database Of Vertebrate Paleontological Sites Across The State Of Tennessee, Sarah Mclaurine

Electronic Theses and Dissertations

Fossil localities across the state of Tennessee and the data related to those sites were compiled from Tennessee Division of Geology Bulletin 84, titled “Tennessee’s Prehistoric Vertebrates,” and stored in a Microsoft Access geodatabase housed by the Department of Collections at the East Tennessee State University Museum of Natural History located at the Gray Fossil Site. Included in the database are forms to enter new site localities, view information about those already entered, view and add data to a master faunal list for the state, view sites repository information and store and add documents that are key-word searchable from the …

Assessing Unstable Fossils For Long-Term Storage, Carson Cope, Alex Landwehr, Kale Link, Israel Rivera-Molina, Laura E. Wilson

SACAD: Scholarly Activities

In 2022, Fort Hays State University’s Sternberg Museum of Natural History (FHSM) began a project to address the long-term preservation of a Late Miocene mammal collection. Many of the fossils from the collection were poorly consolidated, uncurated, and improperly stored, leaving them unstable and prone to significant degradation. To organize our stabilization efforts, we developed a new assessment workflow consisting of two evaluation tools. These tools have helped us categorize the risk factors for specimens, the priority in which they should be addressed, and how to store the fossils long-term. With these newly developed workflows, we are in a better …

Analyzing The Shark Paleoecology Of Coastal Georgia From The Miocene And Pliocene Epochs, Joshua Lee Clark, Benjamin Angalet

Georgia Journal of Science

The field of shark paleoecology often yields indecisive conclusions based on the limited fossilization of their anatomical structures, with the exception of their teeth. The majority of the Atlantic coast has been studied regarding the presence of certain prehistoric shark species from the Miocene, Pliocene, and Pleistocene epochs. However, information pertaining to the Georgia coast and understanding its potential community structure is relatively understudied. This study was conducted in which thousands of fossil shark specimens and subsequent marine fauna were collected from dredge spoils created by the US Army Corps of Engineers (USACE): Savannah District. A total of 5,127 fossil …

Field Guide To Big Bone Lick, Kentucky: Birthplace Of American Vertebrate Paleontology, Glenn W. Storrs, H Gregory Mcdonald, Eric Scott, Robert A. Genheimer, Stanley E. Hedeen, Cameron E. Schwalbach

Special Publication--KGS

Big Bone Lick is the birthplace of vertebrate paleontology in the Western Hemisphere and has a long and celebrated history in the exploration of the American colonial frontier and of the early United States. Notable European scientists of the 18th century such as Buffon, Cuvier, and Hunter discussed the fossils found there. Prominent Americans of the time, such as Boone, Washington, Franklin, and Jefferson are also part of the site’s history. It is the type locality for several extinct late Pleistocene megafaunal mammals, most notably the iconic American Mastodon, who were attracted to the area by salt licks dictated by …

New Species Of Dryolestoid From The Late Cretaceous Allen Formation And Implications For South American Faunal Diversity., Brigid Erin Connelly

Electronic Theses and Dissertations

Dryolestoids are extinct cladotherians mammals from the Jurassic and Cretaceous periods. I describe a collection of dryolestoid specimens from the Late Cretaceous localities of Cerro Tortuga (Allen Formation), Anfiteatro 1, and Shining (both La Colonia Formation) from Patagonia, Argentina. Using comparative morphology, I identify a new species of meridiolestidan dryolestoid based on eleven specimens across both formations. The new species’ recovery from La Colonia Formation represents the first dryolestoid connection between the two approximately contemporaneous formations. The species’ morphology may represent an ecological shift within Meridiolestida from insectivory to herbivory, showing a transition in characters between the plesiomorphic sharp-toothed meridiolestidans …