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Some Information On The Ecology Of Khulan (Equus Hemionus Pallas, 1775) In The Western Part Of The South Gobi Province, Mongolia, D. Tsendjav, S. Purevsuren

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

This short study provides information on the density and distribution of khulans in September 2003 and speculates about factors that negatively influence the khulan population in the sums of Khurmen, Bayandalai, Noyon, Sevrei, and Gurvantes in the South Gobi province.

An Overview On The State Of Equus Hemionus In Whole China, Weikang Yang

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The wild ass in China is divided into two species: Gobi khulan (Equus hemionus) and kiang (Equus kiang). Maps and charts of the distribution and population sizes in China and Mongolia of each species are shown from the 1980s through the early 2000s.

Territorial Behaviour Of Kiang (Equus Kiang Moorcroft, 1841) In Ladakh (India), Natalia V. Paklina, Chris Van Orden

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The observations of kiang behavior were made in Navokar Valley northeast of Tso Kar Lake (Ladakh, India) between July 30 and November 22, 2001. In the breeding season (end of July until the end of August) adult kiang males kept not overlapping, protected territories (about 10 km²), and marked by single defecation and urination marks. There were adult females with and without offspring on the territories (up to 12 animals, including the male). The distance between male and females on the territories was usually hundred times bigger, then a distance between stallion and his harem in horses. Females …

Assessment Of Non-Metric Skull Characters And Age Determination In The Asiatic Wild Ass Equus Hemionus: A Methodological Approach, Hermann Ansorge, Annegret Stubbe, Nyamsuren Batsaikhan, Ravčigijn Samjaa, Michael Stubbe

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The report presents the first information on the use of non-metric skull characters and age determination by tooth section for Asiatic wild asses. It was initiated by a collection of more than 400 skulls of the Asiatic wild ass from the Bordzongijn-Gobi 2002-2004. The methodological overview gives a first research prospect in Asiatic Wild Ass concerning the use of non-metric skull characters as a morphological tool for assessing genetic variability, determining epigenetic divergences and evaluating fluctuating asymmetry. The time course of tooth eruption and replacement in the Asiatic wild ass is described with regard to its use for age determination. …

Remarks On The Social System Of The Mongolian Wild Ass (Equus Hemionus Hemionus), Gertrud Neumann-Denzau, Helmut Denzau

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

So far the social system of Mongolian wild asses is non-uniformly interpreted. The authors describe their own observations from different parts of Mongolia and review data from available publications. The arguments support a territorial social system, which agrees with the social system of all other wild asses in Asia and Africa.

Population Assessment Of Khulan (Equus Hemionus) In Mongolia, Badamjav Lkhagvasuren

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The data presented here suggest that 18,411 (± 224) khulan inhabit southern Mongolia with a density of 1.4 individuals per 1,000 km² within the 157,525 km² territory of its actual distribution. The Dornogobi province contains the highest number of khulan with a mere 67%, while 20% are found in the Ömnögobi province, 12% are in Djungarian Gobi and only 1% in the Gobi-Altay and Bayankhongor provinces. According to the proportion of foals and yearlings within each aimag, the Dornogobi, Ömnögobi east and Khovd populations have an average reproduction rate.

Semi-Wild Population Of Kulans In The Bukhara Breeding Centre And Their Co-Habitation With Przewalski’S Horses, Karim Bahloul, Olga B. Pereladova, Natalia V. Soldatova, Ekaterina V. Sidorenko, Antoine J. Sempere

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Asiatic wild asses and Przewalski's horses initially inhabited steppe, semi-desert and desert areas, but Przewalski's horses became extinct in the wild, and kulans are under threat of disappearance. The Bukhara Breeding Centre (Uzbekistan) was created in 1976 for conservation and reintroduction of wild ungulate species. In 1977-1978, five kulans (two males and three females) from Barsa-Kelmes Island on the Aral lake were introduced to the reserve. The group increased to 25-30 animals in 1995-1998, when five Przewalski's horses from Moscow and St. Petersburg zoos were introduced to the same territory. We analyzed the home ranges, preferred habitats and social interactions …

Up-Date On The Behaviour And Status Of The Critically Endangered Onager, Equus Hemionus Onager, From Iran, Laurent Tatin, Bijan F. Darreh-Shoori, Christophe Tourenq, David Tatin, Bijan Azmayesh

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The onager, Equus hemionus onager, an Asiatic wild ass endemic to Iran, is categorized as Critically Endangered on the IUCN Red List. Its biology and conservation requirements are poorly documented. We report some observations made in 1997 and 2000 on the behavior and ecology of the two remaining populations, located in the Touran Protected Area and the Bahram-e-Goor Reserve. Recent population counts by the Department of Environment of Iran (471 in the Protected Area, and 96 in the Reserve) are markedly lower than the count of 600-770 made in the 1970s in the Touran Protected Area. We observed social …

Asiatic Wild Asses In The Literature: What Do We Need To Know Now?, Sarah R.B. King

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Asiatic wild asses were once found across central Asia, but over time their range has been reduced until today all subspecies are under threat. This paper examined the literature about wild asses with an aim to illustrating gaps in our knowledge as a target for future research.

A search was made on BIOSIS ISI for the following strings: Equus hemionus, wild ass, khulan, kulan, kiang, khur, and onager. In total 61 relevant references were found, published between 1900 and 2005. Most of these references were published in the 1990s, with only one published between 1900 and 1970. Khur and …

Kulan (Equus Hemionus Pallas 1775) In Turkmenistan, Victor S. Lukarevskiy, Yu. K. Gorelov

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The kulan (Equus hemionus kulan) is the only odd-toed ungulate left in the wild in northern Eurasia. In the 1930s, the kulan survived in southern Turkmenistan only. In the 1940s, kulans were found only in an area named Badkhyz in Turkmenistan. To conserve the last natural population of this subspecies of kulan the Badkhyz Natural Reserve was established in 1941. The historical and contemporary distribution, fluctuation in numbers, conservation and sci research of kulan populations in the Badkhyz Natural reserve are reviewed in this paper.

Turkmenian Kulan (Equus Onager Kulan) Captive Population, Anna Mekarska

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The Turkmenian kulan (Equus onager kulan) is critically endangered, fewer then 1,500 animals are left in the wild. In 2004, there were additional 570 Turkmenian kulans living in captivity: 156 specimens in Euro-Asian zoos, 83 in Askania Nova Reserve and 331 in Canyon Colorado Equid Sanctuary. Population size in EAZA region is decreasing both in terms of numbers and genetic parameters. Those facts underline the urgent need for coordinating development of Eurasian kulan’s captive population. From 2004 onwards, large efforts to create healthy breeding groups in zoos raised bigger interest in this species in European zoos.

Asiatic Wild Ass In Israel, David A. Blank

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The Syrian wild ass (Equus hemionus hemippus) lived in Israel in historic times until the 1930s, when it disappeared from the entire Middle East region. Khulans from other subspecies (E. h. onager - six individulas and E. h. kulan - five individuals) were brought to Israel from European zoos during 1968-69 (the authors of project considered that onager and kulan were the same subspecies) and these animals bred together in the semi-captivity conditions of the Haibar Yotvata Reserve. Later, during a 12-year period from 1982 through 1993, six groups totalling 38 khulans (17 males and 21 females) …

Identification Of Herder-Wild Equid Conflicts In The Great Gobi B Strictly Protected Area In Sw Mongolia, Petra Kaczensky, N. Enkhsaihan, O. Ganbaatar, Christian Walzer

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In protected areas of Mongolian overgrazing, competition with wild ungulates and poaching are important management concerns. The Great Gobi B Strictly Protected Area (SPA) in SW Mongolia is a re-introduction site for the Przewalski’s horse (Equus ferus przewalskii), a stronghold of the Asiatic wild ass (Equus hemionus), and remains an important grazing area for seminomadic herders. We show the power of simple inventory and monitoring methods to assess herder-wildlife conflicts, by combining data of: (1) human and livestock demographic data, (2) migration patterns, of semi-nomadic herders, (3) monthly surveys of wild- and domestic ungulates, and (4) …

Investigation Of The Kiang (Equus Kiang, Equidae) Skull From Ladakh, India, Natalia Spasskaya

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The species-rank of Equus kiang was recently supported by molecular and genetic studies (RYDER & CHEMNICK 1990). Three subspecies are currently recognized in this species: eastern - E. k. holdereri, southern - E. k. polyodon and western - E. k. kiang, differing in some cranial and external features (GROVES & MAZAK 1967). Kiang populations are scarce and widely separated at present; which was a reason for a reassessment of the taxonomic status and distribution of at least one or two of these three races.

An interesting skull of kiang was recovered by N. Paklina and C. van Orden …

Surviving A Drought: Population Dynamics Of Ochotona Pallasi Pricei In A Dry Steppe, Gobi Altai, Mongolia, Karin Nadrowski, Georg Miehe

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Two distinct life history traits are described from the genus Ochotona, the group of “burrowing” species exhibiting high but fluctuating population densities and the “non-burrowing” species exhibiting relatively stable low population densities. The life history traits are linked to ecosystem dynamics with climatically variable steppe environments hosting “burrowing” species and relatively stable mountainous and rocky habitats hosting “non-burrowing” species. There are few intermediate species, living in both steppe and rocky environments. This study presents survival rates, reproductive rates and a tentative model of population dynamics for Ochotona pallasi pricei, an intermediate species with respect to habitat preference. We …

The Birth Of A Wild Ass (Equus Hemionus Khur) In India’S Little Rann Of Kutch, Gertrud Neumann-Denzau

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The birth of a wild ass was completely described and photographed in the wild for the first time. It happened in the daytime, in the open desert, and in the vicinity of other wild asses. The mother isolated her foal afterwards for an entire day. After birthing she drove away her male yearling, which did not join her any longer from this moment on.

Biodiversity In Mongolia, Photos, Annegret Stubbe, Michael Stubbe

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Pictures of wildlife in Mongolia in 2007.

The Emerging Dziggetai (Equidae: Equus Hemionus Pallas): An Illustrated History Of Taxonomic Concepts For The Identification, Classification, And Distribution Of Hemiones From Central Asia, Arnd Schreiber

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The history of the discovery and the exploration of the hemione populations (Equus hemionus PALLAS) from Central Asia (Mongolia, northwest China, northeast Tibet, Kazakhstan, and Russian Siberia) from the 13th century to approximately 1950 are compiled with the aim to provide materials for a taxonomic and population genetic revision of the dziggetais. Data on their phenotypic differentiation, historical distribution status, and their husbandry history in European zoological gardens, are reviewed, as is the gradual emergence of concepts of how to identify and to classify these populations. Hemiones were an important historical case in the discussion of how to represent …

Zum Vorkommen Ausgewählter Bembidion-Arten In Der Mongolei (Coleoptera: Carabidae, Bembidiinae): 2. Beitrag Zur Carabidenfauna Der Mongolei, Peer Haro Schnitter

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

To the occurrence of some Bembidion-species in Mongolia (Coleoptera: Carabidae, Bembidiinae) - Second contribution to the carabid fauna of Mongolia.

The distribution of four Bembidion-species, Bembidion (Asioperyphus) infuscatum DEJEAN, 1831, B. (Asioperyphus) ovale MOTSCHULSKY, 1844, B. (Bracteon) conicolle MOTSCHULSKY, 1844 und B. (Bracteon) lapponicum ZETTERSTEDT, 1828, in Mongolia are discussed, and new data are given.

Feeding Ecology Of Asiatic Wild Ass Equus Hemionus, Jochen Lengger, Frieda Tataruch, Christian Walzer

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The Mongolian wild ass or khulan (Equus hemionus) is a potential competitor for forage to reintroduced Przewalski’s horses. To evaluate the major foraging plants of khulan we chose the alkane method that was first described by Mayes in 1984. Different plant species contain varying amounts and proportions of n-alkanes. This fact allows the determination of diet composition by comparing the plant alkane to the fecal alkane pattern. The major advantage is that the method is non-invasive and has been shown to be a reliable method in captive trials. Fecal and plant samples were collected simultaneously and preserved by …

Social Networks In Wild Asses: Comparing Patterns And Processes Among Populations, Daniel I. Rubenstein, Siva Sundaresan, Ilya Fischhoff, David Saltz

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Asiatic wild asses inhabit some of the most arid environments in the world. All live in fissionfusion societies, but demography varies and the deserts in which they live often differ in subtle ways. Characterizing details of social structure of wild ass populations has been a challenge and has made it difficult to determine causes and consequences of any differences that might exist. We use network theory to compare the social structures of two populations of Asiatic asses/ onagers inhabiting the Negev desert, Israel and khur of the Little Rann of Kuch, India and show that populations differ in important structural …

Numbers, Distribution And Social Structure Of Kiang (Equus Kiang Moorcroft 1841) Population In The Southwestern Part Of Tibet, China, Natalia V. Paklina, Chris Van Orden

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In September - October 1998 we made an extensive survey in southwest Tibet to study numbers, distribution and social structure of the kiang population in this remote part of its range. Kiangs were sighted between 29º 40’ and 32º 50’ N, and 81º- 86º E. They preferred broad flat valleys with lakes at an altitude from 4,000 up to 5,000 m. Small kiang groups and solitary animals were sighted in Transhimalaya. There were no kiangs in Tsangpo (Brahmaputra) River valley, in the sand desert near Ali and in the Clay Mountains near Tsada. Seven old kiang corpses were found along …

Winter Pasture Conditions And Forage Use By Argali (Ovis Ammon) In Gobi Gurvan Saykhan National Park, Bayart Mandakh, Ganchimeg J. Wingard, Richard P. Reading

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Conservation of Argali sheep (Ovis ammon) presents a significant challenge to biologists and conservation managers in Mongolia. Argali are endangered and available data suggest that their numbers are declining. However, the mechanisms causing this decline are poorly understood. Recent research suggests that declines in Argali sheep numbers may be due in part to competition with domestic livestock for forage.

Following privatization of herd ownership in 1991, livestock numbers increased dramatically across Mongolia. With this increase, many herders expanded grazing onto more marginal rangeland resulting in increased competition with wild ungulates and displacement of Argali from their former population …

Wild Bactrian Camel Conservation, Richard P. Reading, Evan S. Blumer, Henry Mix, Jadamsuren Adiya

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The wild Bactrian camel (Camelus bactrianus ferus) is critically endangered throughout its range in China and Mongolia. Yet, wild camels remain poorly understood, with knowledge derived primarily from a few short studies and anecdotal information. We initiated a wild camel conservation project to determine the reasons for camel decline and to develop a program to address those problems.

We are employing satellite telemetry to gather data on wild camel movement patterns, home ranges, habitat use, and sources of mortality. We are also collecting feces from camels and wolves to determine important forage plants and to begin to assess …

Beaver Research In The Uvs Nuur Region, Michael Stubbe, N. Dawaa, R. Samjaa, Annegret Stubbe, A. P. Saveljev, D. Heidecke, D. Sumjaa, H. Ansorge, S. Shar, J.-F. Ducroz

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In 1985, 1988, and 2002 Castor fiber birulai was introduced to the Tes Gol of the Uvs Nuur basin in North-western Mongolia. The beavers migrated through the Republic of Tyva and settled in the middle part of Tes Gol near the Tyvinian-Mongolian border. About 10 colonies were recorded in this region in 2002. Strict protection of Castor fiber birulai has to be ensured in Mongolia and the Republic of Tyva in future.

Facts From A Year Of Drought: Forage Competition Between Livestock And The Mongolian Pika (Ochotona Pallasi) And Its Effects On Livestock Densities And Body Condition, Vroni Retzer

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Burrowing small mammals in grasslands have long been regarded as pests because they compete for forage with livestock and reduce the forage availability for livestock by destroying pastures through their intensive digging activity.

In order to investigate forage competition between the Mongolian Pika (Ochotona pallasi) and livestock an exclosure experiment consisting of four different treatments was set up. The treatments were: 1) accessible only for pikas, (only pika) 2) accessible only for livestock, (only livestock) 3) accessible for both herbivore groups (pika & livestock) and 4) no grazing (no grazing). During the investigation period all requirements for forage …

Ecology Of Argali In Ikh Nartiin Chuluu, Dornogobi Aymag, Richard P. Reading, Sukhiin Amgalanbaatar, Ganchimeg J. Wingard, Danny Kenny, Anthony Denicola

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Argali sheep (Ovis ammon) are listed as threatened in both Mongolia and internationally. Yet, little is known about the biology and ecology of this species. Available data suggests that Argali in Mongolia are declining due to direct poaching and competition with domestic livestock. We initiated several research projects to better understand and conserve the species. In this report we discuss Argali ecology using radio telemetry.

We captured and radio-collared 36 Argali using drive-nets, lamb captures, and dating from 2000–2004. Fifteen collared animals have died: 2 due to capture techniques, 8 from predation, 1 from starvation and exposure, 1 …

Reproductive Ecology Of Two Common Woody Species, Juniperus Sabina And Artemisia Santolinifolia, In Mountain Steppes Of Southern Mongolia, Katrin Ronnenberg

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The present study describes the reproductive ecology of the prostrate shrub Juniperus sabina and the dwarf shrub Artemisia santolinifolia in dry mountain steppes of southern Mongolia. Whereas stands of the juniper are located at the drought limit of the genus’ distributional range, the genus Artemisia is typical of central Asian drylands. Both species produce large numbers of reproductive organs. For Juniperus sabina, however, only 2.5% of the morphologically intact seeds were found to be viable. Correspondingly, neither seedlings nor saplings were found in the field, and recruitment by reseeding happens at most rarely under the present climatic conditions.

Instead, …

Zur Populationsökologie Und Raumnutzung Der Steppenwühlmaus Microtus Brandti (Radde, 1861) In Der Fortpflanzungsperiode, Ulrich Zöphel, N. Dawaa

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

During the growing season between 1988 and 1990 the population biology of Brandt’s vole (Microtus brandti) was observed in a population near Ulaanbaatar. Mark-recapture techniques (1,438 markings, 19,606 recaptures in total) and complementary methods were used. The density maximum was observed in July with 98 (1988); 169 (1989) and 94 (1990) individuals/ha and the population fluctuated by a factor of 7.4.

At the beginning of reproductive period the males disperse over a minimum distance of one female home range whereas the females do not migrate after winter. The typical mating system is promiscuity. Youngsters have been recorded from …

Beiträge Zur Ökologie Und Verbreitung Der Steppenwühlmaus Microtus Brandti Radde, 1861 In Der Mongolei = Ecology And Distribution Of The Steppe Vole Microtus Brandti Radde, 1861 In Mongolia, N. Dawaa, Michael Stubbe, Annegret Stubbe

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Microtus brandti is one of the most important pest species of Mongolian rodents. Since the publication of Bannikov (1954) the area of expansion is increased, in the Archangaj-Aimag 200 km westward, in the Central Aimag 70 km to the North and in the north-eastern part of Dornogov’- Aimag about 110 km to the South. In years of gradation the plant cover was exterminated up to 100%. We can distinguish a zone of exposed harmfulness with a damage of 70-100%. In this region 16 great centers of infestation were listed. Between 1925 and 1985 six important calamities were noticed, in intervals …