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Behavior and Ethology Commons

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Theses/Dissertations

2020

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Full-Text Articles in Behavior and Ethology

Variation In Personality Among Semi-Wild Myanmar Timber Elephants, Sateesh Venkatesh Dec 2020

Variation In Personality Among Semi-Wild Myanmar Timber Elephants, Sateesh Venkatesh

Theses and Dissertations

This study examines two personality traits: exploration and neophobia, which could influence human-elephant conflicts. Thirty-one semi-wild elephants were tested over two trials using a custom novel puzzle tube containing three tasks and three rewards. Our studies show that elephants do vary significantly between individuals in both exploration and neophobia.


Influence Of Testosterone On Male-Male Competition In The Red-Sided Garter Snake, Thamnophis Sirtalis Parietalis, Isabella M. G. Bukovich Dec 2020

Influence Of Testosterone On Male-Male Competition In The Red-Sided Garter Snake, Thamnophis Sirtalis Parietalis, Isabella M. G. Bukovich

Senior Honors Projects, 2020-current

Female mimicry in the red-sided garter snake, Thamnophis sirtalis parietalis, is described in several well-studied evolutionary advantages, but an aspect that remains elusive is how reproductive traits are linked to the 3-fold higher circulating androgens in males with this phenotype. In this study, we implanted male garter snakes (n=15 per group) with either a blank implant (SHAM; control) or a T implant, the latter simulating the female mimic phenotype. Following simulated low-temperature dormancy, males were scored for courtship intensity over three days of behavioral trials with females housed in the same facility. Males were sacrificed, and sperm were collected …


Predator-Avoidance Of Larval Black-Bellied Salamanders (Desmognathus Quadramaculatus) In Response To Cues From Native And Nonnative Salmonids, Brian L. Dempsey Dec 2020

Predator-Avoidance Of Larval Black-Bellied Salamanders (Desmognathus Quadramaculatus) In Response To Cues From Native And Nonnative Salmonids, Brian L. Dempsey

Electronic Theses and Dissertations

The introduction of nonnative salmonids into Southern Appalachia may pose a threat to resident salamander populations. In recent years, the stocking and encroachment of rainbow trout (Oncorhynchus mykiss) into headwaters where the black-bellied salamander (Desmognathus quadramaculatus) and brook trout (Salvelinus fontinalis) naturally coexist has raised concerns. In aquatic prey, predator-avoidance responses are primarily influenced through the detection of chemical cues released from predators. The objective of this study was to determine how co-occurrence with a predator influences black-bellied salamander predator recognition behavior. To evaluate this, salamander activity metrics (general activity, number of movements, and latency to move) were recorded before …


Responses Of Juvenile Atlantic Salmon To Competition And Environmental Change: Implications For Performance In Maine Streams, Nicole C. Ramberg-Pihl Dec 2020

Responses Of Juvenile Atlantic Salmon To Competition And Environmental Change: Implications For Performance In Maine Streams, Nicole C. Ramberg-Pihl

Electronic Theses and Dissertations

New England’s climate is changing faster than that of any other region in the continental United States. Over the last century, Maine has experienced an increase in annual temperature of approximately 1.48oC along with a 15 percent increase in annual precipitation. Temperature and precipitation play vital roles in shaping the ecology of freshwater environments. Therefore, changes in regional climate could undermine the structure and stability of Maine’s freshwater systems as they currently exist.

Maine currently harbors the last wild populations of Atlantic salmon (Salmo salar) in the United States. Atlantic salmon were once abundant in Maine …


Effects Of Long-Term Variation In Temperature On Reproductive Phenology In A Population Of Eastern Bluebirds (Sialia Sialis), Paul Pleiman Dec 2020

Effects Of Long-Term Variation In Temperature On Reproductive Phenology In A Population Of Eastern Bluebirds (Sialia Sialis), Paul Pleiman

Graduate Theses and Dissertations

This study investigates the relationship between multiple temperature variables, to include annual and pre-lay date temperatures with first-egg and mean first-egg lay dates of the eastern bluebird at the Warner Parks in Nashville, Tennessee, USA. Data is collected by citizen scientists for the Eastern Bluebird Nesting Box Project while visiting artificial nest boxes throughout the park and recording observations made during the breeding season. Temperature data is retrieved from the Northwest Alliance for Computational Science and Engineering’s Parameter-elevation Regressions on Independent Slopes Model (PRISM) Climate Group, based at Oregon State University. The analyses showed no correlation between annual or pre-lay …


Determining The Physiological And Behavioral Responses To Elevated Salinity In The Freshwater Bivalves, Corbicula Fluminea And Lampsilis Ovata, John W. Roden Dec 2020

Determining The Physiological And Behavioral Responses To Elevated Salinity In The Freshwater Bivalves, Corbicula Fluminea And Lampsilis Ovata, John W. Roden

Electronic Theses and Dissertations

Salinization has been identified as an increasing threat to freshwater mussel diversity in recent years. Native mussels have been observed to display reduced salinity tolerance in comparison to some invasive bivalve species, but methods by which organismal tolerance is achieved are not well understood. This study was designed to compare the behavioral and physiological responses of the native Lampsilis ovata to that of the invasive Corbicula fluminea. Lampsilis were found to exhibit strong behavioral avoidance to salinity exposure, whereas Corbicula displayed very weak avoidance to comparable salinity concentrations followed by indications of osmotic conformation through physiological mechanisms. Prolonged valve …


Dietary Development And Nutritional Ontogeny In Gorilla Beringei : A Multi-Layered, -Omics Approach, Emma C. Cancelliere Sep 2020

Dietary Development And Nutritional Ontogeny In Gorilla Beringei : A Multi-Layered, -Omics Approach, Emma C. Cancelliere

Dissertations, Theses, and Capstone Projects

In species who consume folivorous diets, immature individuals must contend with the challenges of extracting nutrients from fibrous foods before dietary adaptations and strategies are fully developed. Additionally, immatures have distinct nutritional needs to support their stage-specific metabolic and biophysiological requirements. To meet these stage-specific needs, while constrained by underdeveloped feeding strategies and digestive capacities, immatures may adopt distinct diets better suited to their specific developmental context. However, where dietary modification is constrained by low dietary diversity or landscape homogeneity, it is unclear how immature individuals compensate through alternative strategies. In turn, little is known about the nutritional and life …


Hind Wing Eyespots Of Brassolini Butterflies (Lepidoptera, Nymphalidae): Evolutionary Diversification And Functions In Anti-Predator Defense And Mating Behavior, Logan Crees Aug 2020

Hind Wing Eyespots Of Brassolini Butterflies (Lepidoptera, Nymphalidae): Evolutionary Diversification And Functions In Anti-Predator Defense And Mating Behavior, Logan Crees

University of New Orleans Theses and Dissertations

Ventral hind wing eyespots are prominent pattern elements in Brassolini butterflies, likely functioning in predator-prey interactions and reproductive activities. Caligo and Opsiphanes differ in male mate-seeking behaviors and it has been suggested that Caligo females use the male cua1 eyespot as a mate-locating cue, but Opsiphanes females do not seem to do so. We predict Caligo males should have larger eyespots than congeneric females, but the sexes would not differ in eyespot size in Opsiphanes. Our analyses supported both these predictions. Displacing the eyespot to the center of the wing makes eyespots more conspicuous, we asked if eyespot position …


Connecting The Social And Spatial Behaviors Of A Territorial Species (Anolis Carolinensis), Jordan M. Bush Aug 2020

Connecting The Social And Spatial Behaviors Of A Territorial Species (Anolis Carolinensis), Jordan M. Bush

Doctoral Dissertations

Why animals live where they do is a key question in ecology and evolution. An individual’s home range determines the resources they have access to, conspecifics they encounter, and predators and pitfalls they must avoid. Home range behaviors also have an inherently social component; where animals live affects the rivals they compete with and the mates they have access to. This is especially true in territorial species, as defensive displays make up a large portion of their social behaviors. In this dissertation, I sought to understand how territorial behaviors affect the social lives of the green anole lizard (Anolis …


Comparative Spatial And Behavioral Ecology Of The Prairie Lizard Sceloporus Consobrinus In The Arkansas River Valley, E. A. Free Kashon Aug 2020

Comparative Spatial And Behavioral Ecology Of The Prairie Lizard Sceloporus Consobrinus In The Arkansas River Valley, E. A. Free Kashon

Theses and Dissertations from 2020

Habitat-driven differences in reptile life-histories have been observed in many species. Prairie lizards, Sceloporus consobrinus, in the Arkansas River Valley inhabit rocky, forested, and intermediate habitat types that exhibit different thermal resource availabilities and habitat structures. I studied prairie lizard space usage and antipredator responses to examine whether lizards exhibit habitat-driven differences in ecology, and whether these differences are influenced, in part, by individual personality. I utilized radio telemetry to track lizards in each of these habitat types and established estimates of space usage and daily linear movements. I used behavioral approaches to quantify lizard antipredator responses to a simulated …


Influences Of Anti-Bd Bacteria On Amphibians And Their Microbiomes, Emme L. Schmidt May 2020

Influences Of Anti-Bd Bacteria On Amphibians And Their Microbiomes, Emme L. Schmidt

Honors Thesis

The fungal pathogen Batrochochytrium dendrobatidis (Bd) has been threatening amphibian populations across the globe. Recent work has discovered that bacteria isolated from certain populations of amphibians protect hosts from this pathogen. This project focuses on the anti-fungal mechanisms of Serratia marcescens, a species of bacteria isolated from a Costa Rican frog that dramatically inhibited Bd in culture. Wild-type and recombinant S. marcescens was introduced to the microbiomes of a live amphibian host, Acris blanchardi (Blanchard’s Cricket Frog) and then challenged with Bd to examine the protective effects of this bacteria. The experiment includes recombinant S. marcescens with …


Behavioral Ecology Of Territorial Aggression In Uca Pugilator And Uca Pugnax, Abbey N. Thomas May 2020

Behavioral Ecology Of Territorial Aggression In Uca Pugilator And Uca Pugnax, Abbey N. Thomas

Honors Theses

The nature of animal aggression has long been a research interest in many different scientific fields. Resources, including food, shelter, and mates are all common assets for which animals compete. Two species of fiddler crabs, the Atlantic Sand Fiddler Crab (Uca pugilator) and the Atlantic Marsh Fiddler Crab (U. pugnax) were studied in regards to their aggression shown when competing for shelter. One crab was placed in a contained shelter for two days, and then a second of the same species was introduced. Aggressive interactions were scored and compared with claw and carapace size to ascertain …


Determining The Impacts Of Urbanization On Song Structure And Its Function In Territorial Defense For Gray Catbirds (Dumetella Carolinensis), Morgan L. Rhodes May 2020

Determining The Impacts Of Urbanization On Song Structure And Its Function In Territorial Defense For Gray Catbirds (Dumetella Carolinensis), Morgan L. Rhodes

Masters Theses, 2020-current

In urbanized habitats, animals are faced with novel selection pressures such as differences in community structure and increased urban noise. Urban noise pollution can negatively impact songbirds as low-frequency noise often masks portions of birds’ mating signal and reduces signal transmission. Previous research has demonstrated that songs of birds in more urban habitats have structural differences that enhance signal transmission when noise is present. These studies have focused on species that deliver short, stereotyped songs with limited repertoires. Gray catbirds (Dumetella carolinensis, family: Mimidae) sing long bouts containing imitated, improvised, and invented song elements, and therefore may have …


Habitat Selection And Host Detection In The Salamander Mussel, Simpsonaias Ambigua, Eric A. Stegmann May 2020

Habitat Selection And Host Detection In The Salamander Mussel, Simpsonaias Ambigua, Eric A. Stegmann

MSU Graduate Theses

The native freshwater mussels, Order Unionida, have suffered many species extinctions and loss of abundance. Two important threats to native mussels are loss of habitat and loss of access to the vertebrate hosts of the parasitic mussel larvae. The Salamander mussel, Simpsonaias ambigua, is a habitat specialist, living under flat rocks. It is often found in direct association with its only known host, the common mudpuppy, Necturus maculosus. This association could result from movement and habitat selection by the mussels themselves. Alternatively, it might result from the deposition of juveniles by a resident host. Habitat selection and host …


Temporal Factors Affecting Foraging Patterns Of A Diurnal Orb-Weaving Spider, Micrathena Gracilis (Araneae: Araneidae), Mitchell Davis Long May 2020

Temporal Factors Affecting Foraging Patterns Of A Diurnal Orb-Weaving Spider, Micrathena Gracilis (Araneae: Araneidae), Mitchell Davis Long

Undergraduate Honors Theses

Several studies have investigated the ecological factors that affect behavior in Micrathena gracilis, a diurnal orb-weaving spider that forages on flying insects during the day. However, none yet have considered how the temporal distributions of prey and predator occurrences shape their daily behavioral rhythms, especially web construction, which involves a heavy energetic investment well in advance of potential nutritional benefit. Recently, several orb-weaving spider species have been shown to exhibit a variety of abnormal rhythms, suggesting that circadian clock-controlled rhythms may play an unexpected role in behavioral evolution. Despite the appearance of significant insect abundance in the evenings, M. gracilis …


Effects Of Non-Photic Zeitgebers On The Circadian Clock In The Common House Spider, Parasteatoda Tepidariorum (Araneae: Theridiidae), Mattea Garmany, Darrell Moore, Thomas C. Jones May 2020

Effects Of Non-Photic Zeitgebers On The Circadian Clock In The Common House Spider, Parasteatoda Tepidariorum (Araneae: Theridiidae), Mattea Garmany, Darrell Moore, Thomas C. Jones

Undergraduate Honors Theses

Circadian rhythms are endogenous cycles that control physiological and behavioral changes that can be affected by environmental factors which allow most eukaryotic organisms to synchronize their daily activities with the 24-hour day. Parasteatoda tepidariorum,the common house spider, demonstrates a short-period circadian clock averaging 21.6 hours when left in constant darkness, yet they are able to entrain to a 24-hour light cycle. We tested whether these spiders were able to use non-photic Zeitgebers to entrain to the 24-hour day. Periodic presentation of food and disturbance were not found to be effective cues for the spiders’ entrainment. A few individuals were clearly …


Encephalization In Commensal Raccoons: A Unique Test Of The Cognitive Buffer Hypothesis, Peter M. Anderson May 2020

Encephalization In Commensal Raccoons: A Unique Test Of The Cognitive Buffer Hypothesis, Peter M. Anderson

Electronic Theses and Dissertations

This study investigated selective pressures associated with encephalization in mammals and discussed broader implications. Relative brain size as measured by EQ (Encephalization Quotient) was compared between ecological categories. Omnivores had higher average EQ than ecological specialists. Since specialists are disproportionately affected by extinction events, selection for ecological generalism is proposed as encephalization mechanism. This mechanism may reinforce the more widely known Cognitive Buffer Hypothesis (CBH)—the idea that possessing relatively large brains has buffered lineages against environmental change. CBH is tested here by comparing EQs in Procyon lotor (raccoon) in urban and rural environments. CBH predicts that raccoons in the most …


Glare Reduction By Dark Facial Markings And Bills In Birds, Clara Lebow May 2020

Glare Reduction By Dark Facial Markings And Bills In Birds, Clara Lebow

Electronic Theses and Dissertations

Avian facial plumage, bill coloration, and feather microstructure may serve one or more adaptive functions. Several researchers have proposed that dark eyestripes, bills, and facial masks aid in reducing glare, however, there have been relatively few tests of this hypothesis. Dark facial markings have been shown to have an adaptive glare-reduction function in recent field studies of a few species, but this hypothesis has never been tested in a broad multispecies analysis. It is likely that feather microstructure influences feather brightness and has an effect on the efficacy of glare reduction properties of feathers. I examined the link between dark …


Chemical Communication In Songbirds, Leanne A. Grieves Apr 2020

Chemical Communication In Songbirds, Leanne A. Grieves

Electronic Thesis and Dissertation Repository

Avian chemical communication has been understudied due to the misconception that olfaction is unimportant or even lacking in birds. Early work focused on the olfactory foraging capabilities of seabirds because of their ecology (open ocean foraging) and large olfactory bulbs. In contrast, olfaction in passerine birds, comprising over half of all extant avian taxa, was long overlooked due to their relatively small olfactory bulbs. It is now well established that passerines can smell, and their olfactory acuity is comparable to that of macrosmatic mammals such as rats. Much of our theory on communication and mate choice has involved studying visual …


Personality In Stomatopod Crustaceans, Jillian Page Blenderman Apr 2020

Personality In Stomatopod Crustaceans, Jillian Page Blenderman

Electronic Theses and Dissertations

Stomatopods, or mantis shrimp, are small, predatory crustaceans of tropical and subtropical waters. Mantis shrimp research has focused largely on their agonistic interactions and complex visual systems. Despite their broad behavioral repertoire, stomatopods have not yet been investigated for consistent behavioral differences at the individual level, referred to as animal personality. Fifteen individual Neogonodactylus oerstedii were tested in three scenarios designed to measure potential differences in behaviors: 1) exploration of a novel environment, 2) response to and recovery from a startling event, and 3) response to a novel object. Each individual went through the series of tests twice, with a …


Evaluation Of Space Use And Movement By Wild Turkey (Meleagris Gallopavo) During Extreme Climatic Disturbances And Annual Phenological States, David J. Moscicki Jan 2020

Evaluation Of Space Use And Movement By Wild Turkey (Meleagris Gallopavo) During Extreme Climatic Disturbances And Annual Phenological States, David J. Moscicki

LSU Master's Theses

Space use is driven by both intrinsic and extrinsic factors across space and time. Meaning a species demography and habitat requirements may vary across landscapes. Recent studies on wild turkeys (Meleagris gallopavo spp., hereafter turkey) have shown both direct and indirect demographic impacts of climatically driven events on turkey populations. Similarly, there is an abundance of information documenting turkey demographic parameters and space use, but few studies have addressed breeding phenology in great detail. We evaluated the impact on space use and movement patterns of Eastern wild turkeys (M. g. silvestris; n = 20) and Rio Grande wild …


Foraging Strategy Plasticity In Fiordland Penguins (Eudyptes Pachyrhynchus): A Stable Isotope Approach, Jeffrey Wayne White Jan 2020

Foraging Strategy Plasticity In Fiordland Penguins (Eudyptes Pachyrhynchus): A Stable Isotope Approach, Jeffrey Wayne White

Theses, Dissertations and Capstones

Foraging ecology in the crested penguins (genus: Eudyptes) is an active area of research, with new techniques rapidly increasing our understanding of these charismatic species. The most common techniques to assess foraging ecology include stomach content analysis, fecal DNA analysis, stable isotope analysis, tracking, and video loggers. Here we review dietary research on all 8 taxa within the genus Eudyptes to identify gaps in our current knowledge. However, foraging studies that assess dietary segregation require a method for quickly and accurately sexing penguins in the field. Obvious sexual dimorphism in plumage is largely absent in penguins leaving behavioral cues …


Eastern Diamondback Rattlesnake (Crotalus Adamanteus) Ambush Site Selection In Coastal Saltwater Marshes, Emily Rebecca Mausteller Jan 2020

Eastern Diamondback Rattlesnake (Crotalus Adamanteus) Ambush Site Selection In Coastal Saltwater Marshes, Emily Rebecca Mausteller

Theses, Dissertations and Capstones

The eastern diamondback rattlesnake (Crotalus adamanteus; EDB) is a species of conservation concern associated with the imperiled longleaf pine-grassland ecosystem. The longleaf pine ecosystem is characterized by an open canopy and rich ground cover. Researchers have speculated that the vegetation structure of salt marshes may serve as a surrogate habitat for longleaf pine savannas. Although these marshes have little topography, they provide a heterogeneous landscape with patches of mud flats, sandy hard marsh along upper tidal areas, and salt marsh hummocks throughout. I used radio telemetry to monitor free-ranging EDBs on a South Carolina sea island. The goal of my …


Southern Fox Squirrel (Sciurus Niger Niger) Foraging Behavior And Population Dynamics, John Huang Jan 2020

Southern Fox Squirrel (Sciurus Niger Niger) Foraging Behavior And Population Dynamics, John Huang

Theses, Dissertations and Capstones

The change in the interaction between southern fox squirrel (Sciurus niger niger) and eastern gray squirrels (Sciurus carolinensis) provided an opportunity to examine sublethal effects of predation as it relates to a species’ risk perception and habitat structure at two scales. The interactions allowed us to examine the evolutionary responses of the southern fox squirrel and eastern gray squirrel to predation risk reflective of their historically associated landscapes and habitat structures. Second, the succession of savanna woodlands into closed-canopy mixed forest provided an opportunity to assess the flexible behaviors of both species in response to changes …


Mapping Floral Resources For Bees Using Drone Imagery, Arya Massarat Jan 2020

Mapping Floral Resources For Bees Using Drone Imagery, Arya Massarat

HMC Senior Theses

Poor nutrition among modern day honey bee colonies is contributing to their decline. Yet understanding how the diversity and abundance of flowering species around a colony affects its health remains difficult because of the manual labor required to analyze these large foraging landscapes. We describe a procedure for automatically mapping the species of flowering plants around a colony from overlapping drone images. We developed a pipeline for stitching the images together, identifying plants within them, and classifying each plant by its species. The resulting map of the flowering species surrounding a colony could be used in future experiments that aim …


Effects Of Shelterwood And Patch Cut Harvests On A Post White-Nose Syndrome Bat Community In The Cumberland Plateau In Eastern Kentucky, Phillip Lee Arant Jan 2020

Effects Of Shelterwood And Patch Cut Harvests On A Post White-Nose Syndrome Bat Community In The Cumberland Plateau In Eastern Kentucky, Phillip Lee Arant

Theses and Dissertations--Forestry and Natural Resources

The impact of shelterwood and patch cuts harvests on bat communities was tested at three sites in Eastern Kentucky. Shelterwood harvests had 50% of the basal area and understory removed to create a uniform spacing of residual trees. Patch cuts had 1-hectare circular openings created to remove 50% of the basal area creating an aggregated spacing of residual trees. Acoustic detectors were deployed to assess activity levels pre-harvest. Sites were then sampled from 1 – 2 years post-harvest to determine differences. Pre-harvest data revealed little acoustic activity for the Myotis spp. at two sites. The remaining site had high activity …


A Troop, A Raft, A Bed, Hanna Jane Guendel Jan 2020

A Troop, A Raft, A Bed, Hanna Jane Guendel

Senior Projects Spring 2020

A Troop, a Raft, a Bed tells the interwoven fictional stories of three major animals (the mountain gorilla, the Adélie penguin, and the American eel) and four transitional animals (the white stork, the humpback whale, the common octopus, and the great white shark). The stories are told from the animals' perspectives, and are written with language that considers each animal's unique intelligence, mind, and behavior. These stories seek to communicate how animals around the world may be experiencing the various effects of climate change and global warming.


Implementing Welfare Science: Case Studies In Evidence-Based Zoo Management, Marisa Suzanne Spain Jan 2020

Implementing Welfare Science: Case Studies In Evidence-Based Zoo Management, Marisa Suzanne Spain

UNF Graduate Theses and Dissertations

Animal welfare science is a field that focuses on how we can improve the lives and well-being of animals in human care. Modern welfare science has moved away from simply preventing suffering and on to promoting positive welfare states, a concept that has been coined animal “wellness”. A process called evidence-based zoo management has been implemented in many zoos as a way to promote and ensure wellness. This is the idea that husbandry and housing standards should be evaluated and tested for their efficacy using data rather than relying on traditional best practices. In this manuscript we discuss an informal …


The Cardiovascular Response To Acute Hypoxic Conditions In Danio Rerio, Bryce Fetterman Jan 2020

The Cardiovascular Response To Acute Hypoxic Conditions In Danio Rerio, Bryce Fetterman

Williams Honors College, Honors Research Projects

There are many fish that can withstand hypoxia levels. One in particular is the zebrafish (Danio rerio). The zebrafish is a small-sized, Cyprinid teleost fish, and has been used in a laboratory setting for many reasons. A few reasons being that they are widely available, easy to care for, and the cost for these fish are very low. There are various reasons why these fish are used as a model. One main reason is because their cardiovascular system is one of the first developing organs. This allows for better resolution when studying the cardiovascular system. The purpose of …


Possible Role Of Breeding Phenology, Brood Size, And Male Parental Care On The Number Of Nesting Attempts By Female Eastern Bluebirds, A Facultatively Multibrooded Species, Michael Daniel Patton Jan 2020

Possible Role Of Breeding Phenology, Brood Size, And Male Parental Care On The Number Of Nesting Attempts By Female Eastern Bluebirds, A Facultatively Multibrooded Species, Michael Daniel Patton

Online Theses and Dissertations

To maximize reproductive output, several species of songbirds attempt to raise two or more broods in a single breeding season. The results of previous studies have revealed much variation among species in the factors that influence the likelihood of female songbirds attempting to raise multiple broods during a breeding season. As such, additional studies are needed to better understand the roles of early breeding, brood sizes, and, especially, male parental care in determining the likelihood of having multiple broods. My objective, therefore, was to examine the possible effects of breeding initiation date, brood size, and male provisioning behavior on the …