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Behavior and Ethology Commons

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Theses/Dissertations

2014

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Full-Text Articles in Behavior and Ethology

The Role Of Mate Preference And Personality On Reproductive Performance In An Ex-Situ Conservation Breeding Program For The Giant Panda (Ailuropoda Melanoleuca), Meghan S. Martin Dec 2014

The Role Of Mate Preference And Personality On Reproductive Performance In An Ex-Situ Conservation Breeding Program For The Giant Panda (Ailuropoda Melanoleuca), Meghan S. Martin

Dissertations and Theses

Successful captive-breeding and re-introduction programs must have the ability to breed a surplus of genetically suitable animals for release into the wild. Unfortunately, many individuals in captive breeding programs often do not reproduce even when they are apparently healthy and presented with genetically appropriate mates. Mate choice can affect multiple parameters of reproductive sperformance, including mating success, offspring production, survival, and fecundity. We investigated the role of mate preference and personality on the reproductive performance of male and female giant pandas (Ailuropoda melanoleuca) as measured by intromission success and litter production. We conducted these studies on giant pandas at the …


Relationship Between Relative Hive Entrance Position And Dance Floor Location, Chelsea E. Corrigan Dec 2014

Relationship Between Relative Hive Entrance Position And Dance Floor Location, Chelsea E. Corrigan

Undergraduate Honors Theses

It has been observed that returning honey bee foragers congregate with unemployed foragers and food receiver bees in a localized region of the hive known as the dance floor. Here, the returning foragers advertise food sources via the waggle dance. It was hypothesized that the close proximity of the dance floor to the hive entrance was related to foragers minimizing time and travel inside the hive. The hive entrance is conventionally located at the bottom of the hive. It was suggested that this location was ideal for easy removal of debris. This correlation between dance floor location and hive entrance …


The Effects Of Suburbanization On Nest Ectoparasites And Nest Defense Behavior In The Wood Thrush, Evan N. Dalton Nov 2014

The Effects Of Suburbanization On Nest Ectoparasites And Nest Defense Behavior In The Wood Thrush, Evan N. Dalton

Masters Theses

The Wood Thrush (Hylocichla mustelina) is declining throughout its range, yet is capable of persisting in both contiguous forests and small forest patches surrounded by human suburban development. Thus, it is an ideal species for gaining insight into the effects of suburbanization on migrant songbirds. I investigated two aspects of Wood Thrush nesting ecology: nest ectoparasites and nest defense behavior in order to determine if suburbanization influences either aspect. Nests from suburban forests had fewer haematophagous mites, though the abundance of haematophagous blowfly larvae did not differ between suburban and contiguous forests. There was no relationship between the …


Spatio-Temporal Factors Affecting Human-Black Bear Interactions In Great Smoky Mountains National Park, Nathan Buckhout Nov 2014

Spatio-Temporal Factors Affecting Human-Black Bear Interactions In Great Smoky Mountains National Park, Nathan Buckhout

Masters Theses

Wildlife managers use models to aid in predicting high risk areas for human and black bear (Ursus americanus) interactions (HBI). These tools help managers implement management strategies to minimize HBI. Over 3,000 incidents of HBI were compiled from management reports at Great Smoky Mountains National Park (GSMNP) during 1998-2011, a park with 9-10.2 million visitors per year and a black bear population of about 1,600 bears.

We used data from bear management reports along with annual visitor use, mast and bear abundance data to develop a series of generalized linear models to assess the spatial and temporal factors …


The Role Of Chemical Attractants In The Use Of Tick Traps For Life Stages Of Tick Species Ixodes Scapularis, Dermacentor Variabilis And Amblyomma Maculatum, Pamela C. Kelman Oct 2014

The Role Of Chemical Attractants In The Use Of Tick Traps For Life Stages Of Tick Species Ixodes Scapularis, Dermacentor Variabilis And Amblyomma Maculatum, Pamela C. Kelman

Biological Sciences Theses & Dissertations

Amblyomma maculatum, Ixodes scapularis and Dermacentor variabilis are three of the hard-bodied ticks found in the United States. This study explored three aspects of the appetence process of these tick species: attraction, desiccation survival, and locomotive activity. The first portion of this work was to improve one of the standard methods used for collecting ticks from the environment, the tick trap. To determine the attractants needed to capture all life stages of the three chosen species of ticks in the field using tick traps, a series of experiments were conducted using laboratory-reared larval, nymphal and adult A. maculatum, I. scapularis …


Age-Related Aspects Of Mirror-Use By Bottlenose Dolphins (Tursiops Truncatus), Rachel A. Morrison Oct 2014

Age-Related Aspects Of Mirror-Use By Bottlenose Dolphins (Tursiops Truncatus), Rachel A. Morrison

Dissertations, Theses, and Capstone Projects

Bottlenose dolphins are neuroanatomically different and evolutionarily divergent from primates yet they exhibit mirror self-recognition (MSR), a rare cognitive ability in non-human animals. This research investigated the developmental and age-related aspects of MSR in this species. During a longitudinal study, a social group of bottlenose dolphins at the National Aquarium, Baltimore, MD were exposed to a mirror and their behavioral responses were recorded to: 1) further confirm the presence of MSR in this species, 2) determine the age of emergence of MSR and 3) draw comparisons with data documenting the emergence of this ability in humans and great ape species. …


Behavioral Responses Of Brazilian Free-Tailed Bats (Tadarida Brasiliensis) To Noctuid Moth Migrations, Jennifer Joy Krauel Aug 2014

Behavioral Responses Of Brazilian Free-Tailed Bats (Tadarida Brasiliensis) To Noctuid Moth Migrations, Jennifer Joy Krauel

Doctoral Dissertations

Animal migrations involve significant movement of biomass across landscapes and are likely to have cascading effects on animal and plant communities. However, most studies on migration address the behavior and ecology of single taxa, such as birds or insects. Few consider more than one trophic level or predator/prey interaction within the overall migration context. I studied the migration ecology of noctuid moths and of Brazilian free-tailed bats in Texas. Noctuid moth migrations during the 2010-2012 fall seasons were driven significantly by weather at the regional and local levels. Bats also responded to the same weather patterns, with changes in body …


Behavioral Responses Of Burrowing Owls To Experimental Clutch Manipulations: An Investigation Of Conspecific Brood Parasitism And Laying Determinacy, Jamie Lynn Groves Aug 2014

Behavioral Responses Of Burrowing Owls To Experimental Clutch Manipulations: An Investigation Of Conspecific Brood Parasitism And Laying Determinacy, Jamie Lynn Groves

Boise State University Theses and Dissertations

During 2012-2013, I investigated the behavioral responses of western burrowing owls (Athene cunicularia hypugaea) in southwestern Idaho to experimental conspecific brood parasitism to determine if they were capable of egg discrimination, and if they would accept or reject a parasitic egg introduced into their nest. I experimentally added one egg derived from source nests to host nests either during the egg-laying period or just after adult females initiated incubation and examined acceptance or rejection of the parasitic egg in relation to control nests where eggs were handled but not exchanged. Burrowing owl responses to parasitic eggs varied with …


Effects Of Water Parameters On Container Mosquito (Diptera: Culicidae) Oviposition And Performance, Stephanie Sue Schelble Aug 2014

Effects Of Water Parameters On Container Mosquito (Diptera: Culicidae) Oviposition And Performance, Stephanie Sue Schelble

Master's Theses

Water body parameters have a considerable effect on the communities that develop within them. In small container habitats like tires, the depth, surface area, and volume effect the development and success of the mosquito communities. This study investigated the choices of the adult female mosquitoes, Aedes albopictus and Culex quinquefasciatus, between different depths and surface areas. In addition, larval performance was determined under differing depth and larvae densities. Oviposition studies showed that Ae. albopictus had a preference for deeper habitats (χ2= 14.2902, p= 0.0139) but did not prefer any surface areas (χ2= 7.2321, p= 0.0649) though …


An Agent-Based Model Of Ant Colony Energy And Population Dynamics: Effects Of Temperature And Food Fluctuation, Guo Xiaohui Aug 2014

An Agent-Based Model Of Ant Colony Energy And Population Dynamics: Effects Of Temperature And Food Fluctuation, Guo Xiaohui

Electronic Theses and Dissertations

The ant colony, known as a self-organized system, can adapt to the environment by a series of negative and positive feedbacks. There is still a lack of mechanistic understanding of how the factors, such as temperature and food, coordinate the labor of ants. According to the Metabolic Theory of Ecology (MTE), the metabolic rate could control ecological process at all levels. To analyze self-organized process of ant colony, we constructed an agent-based model to simulate the energy and population dynamics of ant colony. After parameterizing the model, we ran 20 parallel simulations for each experiment and parameter sweeps to find …


A Description Of Vocalizations And Their Association With Mouthing Behaviors And Social Context In Bottlenose Dolphins, Tursiops Truncatus, Christina Elyse Perazio Aug 2014

A Description Of Vocalizations And Their Association With Mouthing Behaviors And Social Context In Bottlenose Dolphins, Tursiops Truncatus, Christina Elyse Perazio

Master's Theses

Dolphin communication is multimodal and incorporates physical behaviors and vocalizations. Dolphins often exchange information with conspecifics using different types of vocalizations, and these vocalizations are sometimes associated with specific behaviors. However, the relationship of vocalization type and mouthing behavior type has not been investigated. This thesis examines simultaneous acoustic and visual recordings of bottlenose dolphins (Tursiops truncatus) to determine the relationship between type of mouthing behavior and type of vocalization (whistle, whistle-squawk, chirp, moan, burst-pulse type A, burst-pulse type B, and click trains). The role of the social context of a mouthing behavior is also evaluated. Data were obtained opportunistically …


Behavioral And Hormonal Flexibility Across Light Environments In Guppies (Poecilia Reticulata), Julia C. Walz Aug 2014

Behavioral And Hormonal Flexibility Across Light Environments In Guppies (Poecilia Reticulata), Julia C. Walz

Master's Theses

Behavior may be dramatically influenced by changing environments, and differences in light intensity environments may have important behavioral consequences. One approach to understanding changes in behavior is by studying behavioral syndromes, suites of correlated behaviors reflecting between individual consistencies in behavior expressed within a behavioral situation (e.g., correlations between antipredator behaviors in different habitats), or across behavioral contexts (e.g., correlations among feeding, antipredator, or mating behavior) (Sih et al. 2004a). Guppies (Poecilia reticulata) provide a great model system to study behavior. Guppies are small, freshwater tropical fish that inhabit still pools in swift-flowing streams, and the backwaters of small rivers …


The Ecological Impact Of Casitas On Caribbean Spiny Lobster (Panulirus Argus) Populations In The Florida Keys, Fl, (Usa), Benjamin C. Gutzler Jul 2014

The Ecological Impact Of Casitas On Caribbean Spiny Lobster (Panulirus Argus) Populations In The Florida Keys, Fl, (Usa), Benjamin C. Gutzler

Biological Sciences Theses & Dissertations

Casitas are artificial structures placed on the seabed by fishermen to aggregate Caribbean spiny lobsters (Panulirus argus) for ease of capture. Some researchers suggest that they may also enhance lobster populations in shelter-limited environments. Conversely, aggregation of lobsters within casitas may be detrimental to the population if the nutritional condition or mortality of lobsters is poorer in casitas than in natural shelters. Small juvenile lobsters may be at particular risk because their foraging range is smaller and they are more readily preyed upon than larger lobsters. If so, then casitas placed in lobster nurseries may function as "ecological traps"; wherein …


Monarch Butterfly (Danaus Plexippus) Tree Preference And Intersite Movement At California Overwintering Sites, Jessica Lynn Griffiths Jun 2014

Monarch Butterfly (Danaus Plexippus) Tree Preference And Intersite Movement At California Overwintering Sites, Jessica Lynn Griffiths

Master's Theses

Managing Monarch butterfly overwintering groves: making room among the eucalyptus

Proper management and conservation of the coastal California overwintering sites used by western Monarch butterflies (Danaus plexippus L.) is critical for continued use of these sites by monarchs. Many management efforts are currently concentrating on eucalyptus-only sites because of the prevailing notion that monarchs prefer eucalyptus over native tree species. Yet, whether a preference exists or not has never been tested. Herein, we test the “eucalyptus preference” hypothesis with data from five overwintering sites comprised of blue gum eucalyptus (Eucalyptus globulus) and at least one other native tree …


Role Model Fathers Or Deadbeat Dads? A Study Of Peromyscus Paternal Behavior, Taylor Wapshott May 2014

Role Model Fathers Or Deadbeat Dads? A Study Of Peromyscus Paternal Behavior, Taylor Wapshott

Senior Theses

Paternal behavior is a largely understudied and poorly understood topic, especially in mammalian species. Many current mammalian models for paternal behavior use a comparative approach, taking advantage of natural differences in behavior between closely related species. This study compared paternal behavior in two rodent species, namely Peromyscus maniculatus (BW) and Peromyscus polionotus (PO). PO rodents have been shown to be monogamous, but there have been no studies of their paternal behavior at this time. 10 PO males and 12 BW males were filmed in their home cage for a 10 minute period following initial disturbance of their nest and removal …


Adaptive Strategies For Foraging And Their Implications For Flower Constancy, Or: Do Honey Bees Multitask?, Ashley E. Wagner May 2014

Adaptive Strategies For Foraging And Their Implications For Flower Constancy, Or: Do Honey Bees Multitask?, Ashley E. Wagner

Electronic Theses and Dissertations

Classical experiments on honey bee time-memory showed that foragers trained to collect food at a fixed time of day return the following day with remarkable time-accuracy. Previous field experiments revealed that not all foragers return to a food source on unrewarded test days. Rather, there exist 2 subgroups: “persistent” foragers reconnoiter the source; “reticent” foragers wait in the hive for confirmation of source availability. To examine how these foragers contribute to a colony’s ability to reallocate foragers across sources with rapidly changing availabilities, foragers were trained to collect sucrose during a restricted window for several days and observed over 3 …


Diel Patterns Of Foraging Aggression And Antipredator Behavior In The Trashline Orb-Weaving Spider, Cyclosa Turbinata, James C. Watts May 2014

Diel Patterns Of Foraging Aggression And Antipredator Behavior In The Trashline Orb-Weaving Spider, Cyclosa Turbinata, James C. Watts

Electronic Theses and Dissertations

Few studies have rigorously assessed the adaptive value of diel rhythms in animals. We laid the groundwork for assessing the adaptive rhythm hypothesis by assaying diel rhythms of foraging and antipredator behavior in the orb-weaving spider Cyclosa turbinata. When confronted with a predator stimulus in experimental arenas, C. turbinata exhibited thanatosis behavior more frequently and for longer durations during the day. However, assays of antipredator response within webs revealed more complex diel patterns of avoidance behaviors and no pattern of avoidance behavior duration. Assays of prey capture behavior found that the likelihood of exhibiting prey capture behavior varied significantly …


Is There Variation In The Effects Of Primate Size As Seed Dispersers?: Seed And Seedling Performance After Gut Simulation Treatments In Hydrochloric Acid, Denise Chac May 2014

Is There Variation In The Effects Of Primate Size As Seed Dispersers?: Seed And Seedling Performance After Gut Simulation Treatments In Hydrochloric Acid, Denise Chac

Chancellor’s Honors Program Projects

No abstract provided.


Testing For Positive Edge Responses In A Fragmented Landscape In The Eastern Tiger (Papilio Glaucus) And The Spicebush (P. Troilus) Swallowtail Butterflies, Jenna C. Siu Apr 2014

Testing For Positive Edge Responses In A Fragmented Landscape In The Eastern Tiger (Papilio Glaucus) And The Spicebush (P. Troilus) Swallowtail Butterflies, Jenna C. Siu

Electronic Thesis and Dissertation Repository

Landscape changes such as habitat fragmentation and habitat loss are contributing to a global decline in biodiversity. While habitat fragmentation research has mainly focused on species that avoid edges, or the boundaries between different landcover types (negative edge response), a hypothesized resource distribution model predicts that species that require complementary resources in different landcovers will be most abundant at edges (positive edge response). Adults of Eastern Tiger (Papilio glaucus) and Spicebush (P. troilus) swallowtail butterflies require forests for oviposition sites and meadows for nectar resources. I examined the relative abundance and flight orientation of both species …


The Culture Of Observation The Native Story Tradition In The Pacific Northwest Between 1850 And 1930, Rob Black Apr 2014

The Culture Of Observation The Native Story Tradition In The Pacific Northwest Between 1850 And 1930, Rob Black

Senior Theses and Projects

No abstract provided.


The Effects Of Predation Risk And Thermoregulation Cost On The Foraging Behavior Of The Eastern Grey Squirrel (Sciurus Carolinensis), Anne E. Collier Apr 2014

The Effects Of Predation Risk And Thermoregulation Cost On The Foraging Behavior Of The Eastern Grey Squirrel (Sciurus Carolinensis), Anne E. Collier

Senior Theses and Projects

In this study, the effects of predation risk and thermoregulation cost on the foraging behavior of the urban eastern gray squirrel (Sciurus carolinensis) were examined. There were two objectives. The first was to determine if either thermoregulation cost or predation risk had a more significant effect on foraging behavior. The second was to find out if foliar cover or distance to refuge was a more important cue of predation risk. To accomplish these goals, giving-up density and temperature data were collected at feeding trays both under the canopy and outside of the canopy at two deciduous trees and …


Testing Assumptions Of Coevolution In An Egg-Rejecting Brood Parasite Host: Uncovering Sensory, Cognitive, And Evolutionary Drivers Of Responses To Parasitism In American Robins (Turdus Migratorius), Rebecca Croston Feb 2014

Testing Assumptions Of Coevolution In An Egg-Rejecting Brood Parasite Host: Uncovering Sensory, Cognitive, And Evolutionary Drivers Of Responses To Parasitism In American Robins (Turdus Migratorius), Rebecca Croston

Dissertations, Theses, and Capstone Projects

Hosts of brood parasitic birds face fitness costs associated with rearing unrelated offspring. In response, the recognition and rejection of parasitic eggs is a common host defense. Brown-headed cowbirds (Molothrus ater) challenge coevolutionary theory, because although they exploit over 200 host species, they lay non-mimetic eggs, and most hosts do not combat cowbird parasitism with egg rejection. American robins (Turdus migratorius) are one of a handful of cowbird hosts known to recognize and remove cowbird eggs from the nest. I addressed the mechanistic and evolutionary drivers of egg rejection in this host species, by disentangling the roles of spectral tuning …


How Important Is Land-Based Foraging To Polar Bears (Ursus Maritimus) During The Ice-Free Season In Western Hudson Bay? An Examination Of Dietary Shifts, Compositional Patterns, Behavioral Observations And Energetic Contributions, Linda J. Gormezano Feb 2014

How Important Is Land-Based Foraging To Polar Bears (Ursus Maritimus) During The Ice-Free Season In Western Hudson Bay? An Examination Of Dietary Shifts, Compositional Patterns, Behavioral Observations And Energetic Contributions, Linda J. Gormezano

Dissertations, Theses, and Capstone Projects

Trophic mismatches between predators and their prey are increasing as climate change causes decoupling of phenological relationships. Predators linked to the life histories of a particular prey will have a more difficult time persisting through environmental change unless they can alter their behavior to maintain the historical match or possess the ability to pursue alternate prey. Arctic predators typically possess flexible foraging strategies to survive in the labile environment, however, quantifying the limits of those strategies can be difficult when life history information is incomplete. In such cases, piecing together different aspects of a predator's foraging behavior, particularly when environmental …


Environmental Factors Influencing Spring Migration Chronology Of Lesser Scaup (Aythya Affinis), Taylor A. Finger Jan 2014

Environmental Factors Influencing Spring Migration Chronology Of Lesser Scaup (Aythya Affinis), Taylor A. Finger

Electronic Thesis and Dissertation Repository

Weather likely affects the timing and rate of migration by waterfowl to their breeding grounds. I hypothesized that timing of migration by lesser scaup during spring is affected by annual variation in temperature, precipitation and ice cover. I used satellite telemetry data, waterfowl survey data and corresponding weather data to evaluate competing models that explained variation in timing and rate of migration by lesser scaup. Timing of spring migration occurred earlier and faster when lesser scaup encountered warmer temperatures and greater precipitation, both of which are known to influence thermoregulation and habitat availability for waterfowl. Migration chronology of lesser scaup …


Using Ailuropoda Melanoleuca As A Model Species For Studying The Ecomorphology Of Paranthropus, Ryan Andrew King Jan 2014

Using Ailuropoda Melanoleuca As A Model Species For Studying The Ecomorphology Of Paranthropus, Ryan Andrew King

Theses, Dissertations and Capstones

Giant pandas (Ailuropoda melanoleuca) have cranial morphology similar to the extinct hominin genus Paranthropus which makes them an excellent model species when studying Paranthropus diet. Both species have wide skulls with flared zygomatic arches adapted for chewing. To gain insight into possible food sources of Paranthropus, I investigated the giant panda’s specialized diet of bamboo. The toughness, hardness, and stiffness of various bamboo species was determined to assess mechanical challenges facing giant pandas during feeding. Bamboo is thought to be tough, but studies on such properties and how they apply to mastication of giant pandas are largely …


A Quantitative Analysis Of The Effect Of Cash-4-Grass Programs On Water Consumption, Cynthia Allen Jan 2014

A Quantitative Analysis Of The Effect Of Cash-4-Grass Programs On Water Consumption, Cynthia Allen

Walden Dissertations and Doctoral Studies

Water supplies in California are stretched to critical levels as a result of population growth, periodic drought, and climate change. The California legislature recognized that the best way to increase supply is to decrease demand so the Water Use Efficiency Senate Bill 7 (SBx7-7) was signed into law in 2009. The law requires water purveyors to reduce per capita water usage by 20% by the year 2020. To comply, water purveyors are searching for innovative ways to increase water conservation. A review of the literature has shown that many factors influence water consumption. However, the majority of household water consumption …


Habitat Quality Influences Migratory Strategy Of Female White-Tailed Deer, Charles R. Henderson Jr. Jan 2014

Habitat Quality Influences Migratory Strategy Of Female White-Tailed Deer, Charles R. Henderson Jr.

Graduate Student Theses, Dissertations, & Professional Papers

Partial migration is a life history strategy that is common for ungulate species living in seasonal environments. One factor that influences the decision to migrate by ungulates is access to high quality habitat. We evaluated the influence of access to winter habitat of high quality on the probability of an individual migrating, the differences in seasonal habitat use between and within migratory and resident classes of deer, and the effects of this decision on the survival of female white-tailed deer. We hypothesized that deer with home ranges of relatively low quality in winter would have a relatively high probability of …


Effects Of Audible Human Disturbance On Koala (Phascolarctos Cinereus) Behavior In Queensland, Australia And Implications For Management, Galina Eugenia Kinsella Jan 2014

Effects Of Audible Human Disturbance On Koala (Phascolarctos Cinereus) Behavior In Queensland, Australia And Implications For Management, Galina Eugenia Kinsella

Honors Theses and Capstones

As the growing human population continues to encroach on wildlife habitat, species are forced to adapt in order to survive. In addition to causing habitat loss, human presence can create more subtle disturbances, such as noise pollution, that disrupt wildlife behavior. Adapting to human presence is particularly difficult for species with specialized resource needs or low mobility. The koala (Phascolarctos cinereus), a national icon of Australia, meets both these criteria. Koalas were once abundant throughout Australia, but are now classified as “threatened.” Habitat loss is at least partly responsible for this decline, but anthropogenic noise may also be …


Pcb Effects On Development Of Behavior In Rats, Robert Witte Jan 2014

Pcb Effects On Development Of Behavior In Rats, Robert Witte

Honors Projects

Abstract

PCBs were originally used as industrial chemicals in many devices and applications, including electrical capacitors and transformers, lubricants, and sealants in buildings. PCBs have been investigated and their use has been slowly phased out starting in the U.S. in the 1970s. A worldwide ban was put on the use and manufacturing of PCB in 2001, however they remain in the food chain because of their lipophilic properties and their long half-life. Major sources of PCB include wild-life, fish, and other seafood. Individuals exposed to PCB during development can experience negative effects on neurobiological, cognitive, and behavior functioning in both …


Tactical Escape Behavior And Performance Of A Small Sceloporine Lizard In Two High Risk Habitats, Seth Parker Jan 2014

Tactical Escape Behavior And Performance Of A Small Sceloporine Lizard In Two High Risk Habitats, Seth Parker

Electronic Theses and Dissertations

The negotiation of uneven and complex terrain has implications for many small terrestrial vertebrates. Variation in the running surface due to obstacles like woody debris or vegetation can alter escape paths and running performance. Additionally, these microhabitat features can influence behavioral tactics in complex environments. The ability to negotiate physical barriers in dense environments likely influences survivorship through important ecological tasks, such as finding mates, foraging, and evading predators. The Florida scrub lizard (Sceloporus woodi) is a small, rare species endemic to two distinct and structurally complex environments in Florida, i.e. sand-pine scrub and longleaf pine-wiregrass sand-hills. The …