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Revision Of The Central Asian Scorpion Genus Anomalobuthus Kraepelin, 1900, With Descriptions Of Three New Species And A Generic Synonymy (Scorpiones: Buthidae), Rolando Teruel, František Kovařík, Victor Fet

Euscorpius

We revise the Central Asian endemic genus Anomalobuthus Kraepelin, 1900, which was considered monotypic for more than 100 years until the recent addition of a second species from Iran (Teruel et al., 2014). We redefine the generic diagnosis of Anomalobuthus and reveal that it is composed of no less than six species, three of which are described as new: A. krivochatskyi, sp. n. (central Uzbekistan and extreme southern Kazakhstan), A. lowei, sp. n. (southeastern Kazakhstan), and A. pavlovskyi, sp. n. (south-central Kazakhstan and extreme northern Turkmenistan). The monotypic genus Psammobuthus Birula, 1911 (described from the Ferghana Valley …

Notes On The Genera Buthacus, Compsobuthus, And Lanzatus With Several Synonymies And Corrections Of Published Characters (Scorpiones: Buthidae), František Kovařík

Euscorpius

The taxonomic position of Buthacus armasi Lourenço, 2013, B. clevai Lourenço, 2001, B. huberi Lourenço, 2001, B. maliensis Lourenço & Qi, 2007, B. nigerianus Lourenço & Qi, 2006, Compsobuthus andresi Lourenço, 2004, C. simoni Lourenço, 1999, C. tassili Lourenço, 2010, C. tofti Lourenço, 2001, C. williamsi Lourenço, 1999, and Sabinebuthus elegans Lourenço, 2001 is revised and fictitious characters in their original descriptions are discussed and corrected. Buthacus armasi Lourenço, 2013 is synonymized with Buthacus leptochelys (Ehrenberg, 1829) syn. n., B. huberi Lourenço, 2001 is confirmed to be a synonym of Buthacus occidentalis Vachon, 1953, B. maliensis Lourenço & Qi, …

Three New Chaerilus From Malaysia (Tioman Island) And Thailand (Scorpiones: Chaerilidae), With A Review Of C. Cimrmani, C. Sejnai, And C. Tichyi, František Kovařík, Graeme Lowe, František Šťáhlavský

Euscorpius

Chaerilus majkusi sp. n. from Malaysia (Tioman Island), C. neradorum sp. n. and C. stockmannorum sp. n. from Thailand are described and fully illustrated with color photographs of live and preserved specimens, as well as of their habitat. They are compared to the species C. cimrmani Kovařík, 2012, C. sejnai Kovařík, 2005, and C. tichyi Kovařík, 2000, which we also illustrate with color photographs of live unpublished specimens. Hemispermatophores of C. cimrmani, C. majkusi sp .n., C. stockmannorum sp. n., and C. tichyi are illustrated and compared, and we also describe the karyotypes of C. cimrmani, C. …

Scorpions Of The Horn Of Africa (Arachnida: Scorpiones). Part Xv. Review Of The Genus Gint Kovařík Et Al., 2013, With Description Of Three New Species From Somaliland (Scorpiones, Buthidae), František Kovařík, Graeme Lowe, Pavel Just, Ahmed Ibrahim Awale, Hassan Sh Abdirahman Elmi, František Šťáhlavský

Euscorpius

We describe herein three new species of Buthidae: Gint amoudensis sp. n., G. gubanensis sp. n., and G. maidensis sp. n. from Somaliland. Additional information is provided on the taxonomy and distribution of other species of the genus Gint, fully complemented with color photos of live and preserved specimens, as well as of their habitat. The hemispermatophores of most Gint species are illustrated and described for the first time. In addition to the analyses of external morphology and hemispermatophores, we also describe the karyotype of four Gint species. The number of chromosomes is different for every one …

A New Species Of The Genus Thaicharmus Kovařík, 1995 (Scorpiones: Buthidae) From Northeast India, Zeeshan A. Mirza, Rajesh V. Sanap, Krushnamegh Kunte

Euscorpius

A new species Thaicharmus guptai sp. nov. is described from the northeast Indian state of Tripura. The new species differs from the known members of the genus in the following set of morphological characters: large size (total length 45.6 mm); trichobothrium d₂ is distal to i₁ on femur; trichobothria est, em and et on the external surface of the patella do not form a straight line. Subaculear tubercle is absent. The movable finger of pedipalp chela has 12 cutting rows of denticles. The movable finger is much longer than the pedipalp patella. Number of pectinal teeth 20/19; pectines …

Pseudouroctonus Peccatum Tate Et Al., 2013, Rediscovered, Richard F. Ayrey

Euscorpius

Shortly after the discovery of Pseudouroctonus peccatum Tate, Riddle, Soleglad et Graham, 2013 (Scorpiones: Vaejovidae) there was a large wildfire in the Spring Mountains of Nevada. At the time the species description was published, it was noted that the only known population of this species may have been extirpated by the fire and subsequent flooding of the type locality habitat (Tate et al. 2013). Today it is certain that the species has survived after the collection of 12 new specimens. Additional data on males are also presented.

Dna Markers Confirm Presence Of Euscorpius Avcii Tropea Et Al., 2012 (Scorpiones: Euscorpiidae) On Samos Island, Greece, Aristeidis Parmakelis, Panayiota Kotsakiozi, Gioele Tropea, Ersen Aydın Yağmur, Iasmi Stathi, Victor Fet, Michael E. Soleglad

Euscorpius

Euscorpius avcii Tropea et al., 2012 has been recently described from Dilek Peninsula in western Anatolia (Turkey, Aydın Province). The population from Samos Island in eastern part of the Aegean Sea is found to match closely the Anatolian E. avcii, making it a new, rare species for the Greek fauna, confirmed by two DNA markers as well as morphology. Samos also shares with western Anatolia two other local recently described scorpion species, Iurus kinzelbachi and Neocalchas gruberi (family Iuridae).

Courtship And Mating In Heterometrus Petersii (Thorell, 1876) (Scorpiones: Scorpionidae), Guo-Bin Jiao, Ming-Sheng Zhu

Euscorpius

Courtship and mating in Heterometrus petersii (Thorell, 1876) (Scorpiones: Scorpionidae) was observed in the laboratory. In this paper the behavior components displayed in courtship and mating are identified, analyzed and discussed.

A New Species Of Tityus C. L. Koch, 1836 (Subgenus Brazilotityus Lourenço, 2006) From The Dominican Amber (Scorpiones: Buthidae), Wilson R. Lourenço

Euscorpius

Tityus (Brazilotiyus) hartkorni sp. n., a new species of fossil scorpion belonging to the genus Tityus C. L. Koch and to the subgenus Brazilotityus Lourenço, is described from a specimen in amber from the Dominican Republic. The new species is clearly associated with the extant fauna of the Neotropical region. This discovery attests to a considerable degree of diversity in the Dominican amber-producing forests.

Etudes On Iurids, I. The Orthobothriotaxic Pattern Of Iuridae, With Observations On Neobothriotaxy In Genus Iurus (Scorpiones: Iuroidea), Michael E. Soleglad, František Kovařík, Victor Fet

Euscorpius

A unique orthobothriotaxic pattern of family Iuridae is examined and illustrated. Significant positional differences in this pattern between two monotypic iurid genera, Calchas and Iurus, are presented. With the analysis of over 100 specimens of genus Iurus, representing many localities in Greece and Turkey, several unique occurrences of neobothriotaxy involving both the chela and patella are discussed and illustrated, representing the first report of neobothriotaxy in family Iuridae.

Scorpions Of Iran (Arachnida, Scorpiones). Part V. Chahar Mahal & Bakhtiyari Province, Khodadad Pirali-Kheirabadi, Shahrokh Navidpour, Victor Fet, František Kovařík, Michael E. Soleglad

Euscorpius

Ten species of scorpions belonging to three families are reported from the montane Chahar Mahal & Bakhtiyari Province of Iran. Of these, only Razianus zarudnyi (Birula, 1903) was formerly reported for this province, while nine species are reported for the first time: Androctonus crassicauda (Olivier, 1807), Compsobuthus matthiesseni (Birula, 1905), Hottentotta saulcyi (Simon, 1880), Hottentotta zagrosensis Kovařík, 1997, Mesobuthus eupeus phillipsii (Pocock, 1889), Odontobuthus doriae (Thorell, 1876), Orthochirus zagrosensis Kovařík, 2004, Hemi- scorpius lepturus Peters, 1861, and Scorpio maurus townsendi (Pocock, 1900). Also presented is a key to all species of scorpions found in the Chahar Mahal & Bakhtiyari Province.

Tityus Ythieri Lourenço, 2007 Is A Synonym Of Tityus Magnimanus Pocock, 1897 (Scorpiones: Buthidae): A Combined Approach Using Morphology, Hybridization Experiments, Chromosomes, And Mitochondrial Dna, František Kovařík, František Šťáhlavský, Tereza Kořínková, Jiří Král, Tom Van Der Ende

Euscorpius

Diagnostic morphological differences to distinguish between two scorpion species, Tityus ythieri Lourenço, 2007 and T. magnimanus Pocock, 1897 do not exist; moreover, the two easily hybridize to produce fertile offspring. Therefore, we suggest that T. ythieri is a synonym of T. magnimanus. This conclusion is further supported by (1) very similar distribution of relative sizes of chromosome bivalents in both species, (2) regular pairing of homologous chromosomes in meiosis of their hybrids, and (3) identical sequences of mitochondrial DNA (COI) markers.

Distribution Of Hottentotta Saulcyi (Simon, 1880) (Scorpiones: Buthidae) In Turkey, Ersen Aydın Yağmur, Halil Koç, Mehmet Yalçın

Euscorpius

Distribution of Hottentotta saulcyi (Simon, 1880) (Scorpiones: Buthidae) in Turkey is investigated, and new distributional records are given. Formerly, this species was known only from Mardin Province; here, it is recorded for the first time from Batman, Şırnak, and Hakkâri Provinces.

The True Identity Of Scorpio (Atreus) Obscurus Gervais, 1843 (Scorpiones, Buthidae), Wilson R. Lourenço, Elise-Anne Leguin

Euscorpius

The true identity of the scorpion Scorpio (Atreus) obscurus Gervais, 1843 is discussed based on new data. The original type material was recently discovered in the collections of the Muséum national d’Histoire naturelle in Paris, which confirms that this old species, originally described by Gervais based on two female specimens from French Guiana, does indeed belong to the genus Tityus C. L. Koch, 1836. Furthermore, an analysis of all morphological characteristics of Tityus (Atreus) obscurus verifies its position as a senior synonym of both Tityus paraensis Kraepelin, 1896 and Tityus cambridgei Pocock, 1897. In this paper, we designate the two …

Scorpions Of Iran (Arachnida, Scorpiones). Part Iv. Kohgilouyeh & Boyer Ahmad Province, Shahrokh Navidpour, František Kovařík, Michael E. Soleglad, Victor Fet

Euscorpius

According to previous surveys, only Mesobuthus eupeus phillipsii (Pocock, 1889), Orthochirus zagrosensis Kovařík, 2004, and Hemiscorpius lepturus Peters, 1861 have been recorded from Kohgilouyeh & Boyer Ahmad Province. Collections made by a team under Shahrokh Navidpour (Razi Reference Laboratory of Scorpion Research, Razi Vaccine and Serum Research Institute, Ahvaz, Khoozestan, Iran) reveal eight other species belonging to two families recorded from the province for the first time: Androctonus crassicauda (Olivier, 1807); Compsobuthus matthiesseni (Birula, 1905); Hottentotta saulcyi (Simon, 1880); Hottentotta zagrosensis Kovařík, 1997; Odontobuthus bidentatus Lourenço et Pézier, 2002; Orthochirus iranus Kovařík, 2004; Razianus zarudnyi (Birula, 1903); and Scorpio maurus …

The Geographic Pattern Of Distribution Of The Genus Rhopalurus Thorell, 1876 In The Guayana-Amazon Region (Scorpiones: Buthidae), Wilson R. Lourenço

Euscorpius

A new discussion is proposed on the geographic patterns of distribution presented by the species of the genus Rhopalurus Thorell, inhabiting the savannah formations of the northern South America. Two lineages of Rhopalurus species are defined for this region, each one containing forms with very weak morphological differences, a consequence of only a minor process of differentiation. The observed pattern of distribution and differentiation is suggested to be directly associated with more or less recent palaeoclimatic vicissitudes, which took place in tropical South America during Pleistocene. Two new subspecies are also described, representing new records of Rhopalurus for Brazil and …

A New Species Of Ananteris Thorell, 1891 (Scorpiones: Buthidae) From The Caribbean Region Of Colombia, Rolando Teruel, César A. Roncallo

Euscorpius

A new species of Ananteris Thorell, 1891 is herein described from a single locality in the Caribbean lowlands of Colombia, in northern Bolívar Department. This new taxon shares several unusual features with Ananteris tolimana Teruel et García, 2007, from the Andean region of Tolima Department. With this addition, the number of Colombian species of Ananteris is raised to seven.

Contributions To Scorpion Systematics. Iii. Subfamilies Smeringurinae And Syntropinae (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet

Euscorpius

Stockwell (1989), in an unpublished revision, suggested splitting the family Vaejovidae in two subfamilies, introducing also four tribes and four new genera. These results have not been reevaluated for 20 years. Here, we establish a new subfamily Smeringurinae, subfam. nov., with two tribes, four genera, and 40 species and subspecies. The subfamily Smeringurinae includes two tribes: Smeringurini, trib. nov. and Paravaejovini, trib. nov. The tribe Smeringurini includes three genera: Paruroctonus Werner, 1934; Smeringurus Haradon, 1983; and Vejovoidus Stahnke, 1974. The tribe Paravaejovini includes the genus Paravaejovis Williams, 1980. We restore from synonymy the subfamily Syntropinae Kraepelin, 1905, and …

Scorpions Of Iran (Arachnida, Scorpiones). Part Iii. Ilam Province, Shahrokh Navidpour, Victor Fet, František Kovařík, Michael E. Soleglad

Euscorpius

Seven species of scorpions belonging to three families have been previously recorded from the Ilam Province of Iran: Androctonus crassicauda (Olivier, 1807), Compsobuthus matthiesseni (Birula, 1905), Hottentotta saulcyi (Simon, 1880), Mesobuthus eupeus phillipsii (Pocock, 1889), Razianus zarudnyi (Birula, 1903), Scorpio maurus townsendi (Pocock, 1900), and Hemiscorpius lepturus Peters, 1861. Collections made by a team organized by Shahrokh Navidpour (Razi Reference Laboratory of Scorpion Research, Razi Vaccine and Serum Research Institute, Ahvaz, Khoozestan, Iran) reveal seven other species recorded from the province for the first time: Apistobuthus susanae Lourenço, 1998, Buthacus macrocentrus (Ehrenberg, 1828), Compsobuthus jakesi Kovařík, 2003, Odontobuthus bidentatus …

Rare Or Poorly Known Scorpions From Colombia. Iii. On The Taxonomy And Distribution Of Rhopalurus Laticauda Thorell, 1876 (Scorpiones: Buthidae), With Description Of A New Species Of The Genus, Rolando Teruel, César A. Roncallo

Euscorpius

A new species of the genus Rhopalurus Thorell, 1876 is herein described from northeastern Colombia; the new species is closely related to (and has been previously confused with) Rhopalurus laticauda Thorell, 1876. Also, some comments on the taxonomy and distribution of the latter taxon are included.

Scorpions Of Iran (Arachnida, Scorpiones). Part Ii. Bushehr Province, Shahrokh Navidpour, Michael E. Soleglad, Victor Fet, František Kovařík

Euscorpius

The Bushehr Province of Iran contains the type localities of three scorpion taxa, Mesobuthus eupeus phillipsii (Pocock, 1889), Orthochirus iranus Kovařík, 2004, and Scorpio maurus townsendi (Pocock, 1900). Apart from them, Androctonus crassicauda (Olivier, 1807), Compsobuthus matthiesseni (Birula, 1905), Hottentotta saulcyi (Simon, 1880), Odontobuthus bidentatus Lourenço & Pézier, 2002, Razianus zarudnyi (Birula, 1903), and Hemiscorpius lepturus Peters, 1862 have also been known to occur in the province. Collections made by a team under Shahrokh Navidpour (Razi Reference Laboratory of Scorpion Research, Razi Vaccine and Serum Research Institute, Ahvaz, Khoozestan, Iran) reveal four additional species recorded from the province for the …

A New Scorpion Genus Representing A Primitive Taxon Of Tribe Stahnkeini, With A Description Of A New Species From Sonora, Mexico (Scorpiones: Vaejovidae), Matthew R. Graham, Michael E. Soleglad

Euscorpius

A new scorpion genus, Gertschius gen. nov., and species, Gertschius crassicorpus sp. nov., (Scorpiones: Vaejovidae) from Sonora, Mexico is described. Gertschius exhibits a reduced modification of the basal pectinal teeth in the female holotype and serration of the chelal finger denticles is limited to the median denticles (MD), thus we consider it a relict member of tribe Stahnkeini. Serradigitus agilis Sissom et Stockwell, 1991, is transferred to this new genus; Gertschius agilis, comb. nov.

Identity Of Planidium Larvae (Hymenoptera: Chalcidoidea) Previously Recorded On Antillean Scorpions, Luis F. De Armas

Euscorpius

The insect planidium larvae found on some Antillean scorpions of the family Buthidae, and previously recorded as belonging to Perilampidae wasps (Hymenoptera: Chalcidoidea), are herein identified as members of Eucharitidae, possibly genus Kapala. Their occurrence on scorpions is interpreted as accidental or perhaps phoretic.

Resumen

Las larvas tipo planidio halladas sobre escorpiones antillanos de la familia Buthidae y previamente registradas como pertenecientes a avispas Perilampidae (Hymenoptera: Chalcidoidea), son aquí reconocidas como miembros de la familia Eucharitidae. Su presencia en escorpiones pudiera ser accidental o tratarse de foresis.

Scorpions Of Iran (Arachnida, Scorpiones). Part I. Khoozestan Province, Shahrokh Navidpour, František Kovařík, Michael E. Soleglad, Victor Fet

Euscorpius

Collections made by a team of Shahrokh Navidpour (Razi Reference Laboratory of Scorpion Research, Razi Vaccine and Serum Research Institute, Ahvaz, Khoozestan, Iran) include 17 of the 19 species known to inhabit Khoozestan Province, and form the basis of this paper. Among them are two new species (Hottentotta khoozestanus sp. n. and Vachoniolus iranus sp. n.), Compsobuthus jakesi Kovařík, 2003 previously known only from Iraq, and five species representing first records for the province: Buthacus macrocentrus (Ehrenberg, 1828); Odontobuthus bidentatus Lourenço & Pézier, 2002; Orthochirus farzanpayi (Vachon et Farzanpay, 1987); Orthochirus stockwelli (Lourenço et Vachon, 1995) comb. n. …

Rare Or Poorly Known Scorpions From Colombia. Ii. Redescription Of Tityus Columbianus (Thorell, 1876) (Scorpiones: Buthidae), Rolando Teruel, Luis F. Garcia H

Euscorpius

A fully illustrated redescription is herein presented of Tityus columbianus (Thorell, 1876). It includes a description of the adult male, an update of its geographical distribution, and detailed information about age-related individual variation, with comments on the taxonomy of the “Tityus clathratus” group.

Rare Or Poorly Known Scorpions From Colombia. I. Redescription Of Tityus Macrochirus Pocock, 1897 (Scorpiones: Buthidae), Rolando Teruel, Luis F. Garcia H

Euscorpius

A fully illustrated redescription is herein presented of Tityus macrochirus Pocock, 1897, based upon a series of specimens of both sexes recently collected in a new location near the type locality. We include an update of its geographical distribution and detailed information about age-related individual variation.

A New Species Of Tarsoporosus Francke, 1978 (Scorpiones: Scorpionidae: Diplocentrinae) From Northeastern Colombia, Rolando Teruel, César A. Roncallo

Euscorpius

A new species of Tarsoporosus Francke, 1978 is described herein from two localities in La Guajira Peninsula, northeastern Colombia. This new taxon inhabits arid environments, and is morphologically close to Tarsoporosus flavus González-Sponga, 1984, from northwestern Venezuela. With this addition, the total number of described species in Tarsoporosus is raised to five, with at least two occurring in Colombia.

A New Species Of Ananteris Thorell, 1891 From Cordillera Central In Colombia, With Some Notes On The Taxonomy Of The Genus (Scorpiones: Buthidae), Rolando Teruel, Luis F. García H

Euscorpius

A new species of Ananteris Thorell, 1891 is herein described from a single locality placed in the Cordillera Central of Colombia, which represents the first record of this genus for Tolima department. This new taxon exhibits several singular features, and apparently represents a relict population inside an urbanized area. Also, some aspects on the taxonomy of the genus are briefly discussed, and its diagnosis is emended. With this addition, the total of Ananteris species described from Colombia is raised to six.

A Revision Of The Genus Hottentotta Birula, 1908, With Descriptions Of Four New Species (Scorpiones, Buthidae), František Kovařík

Euscorpius

The genus Hottentotta is revised and a key to the species is provided. Subgenera Balfourianus Vachon, 1979 and Deccanobuthus Lourenço, 2000 are synonymized with the subgenus Hottentotta Birula, 1908. Four new species of Hottentotta are described: H. finneganae sp. n. from Pakistan, H. jabalpurensis sp. n. from India (Madhya Pradesh), H. jalalabadensis sp. n. from Afghanistan, and H. stockwelli sp. n. from India (Andhra Pradesh and Maharashtra). The following species and subspecies are synonymized: Buthotus alticola kabulensis (Vachon, 1958) with H. buchariensis (Birula, 1897) comb. n.; Hottentotta caboverdensis Lourenço & Ythier, 2006 with H. hottentotta (Fabricius, 1787); H. acostai …

Mesobuthus Nigrocinctus (Ehrenberg, 1828) (Scorpiones: Buthidae) In Turkey: Distribution And Morphological Variation, Ayşegül Karataş

Euscorpius

Distribution of Mesobuthus nigrocinctus is studied and new geographical records are given. As a result of field studies, M. nigrocinctus is recorded from 17 confirmed localities from five provinces in Turkey. Comparative statistical analysis of morphology of M. nigrocinctus and M. gibbosus indicated highly significant differences between these two species. Student’s t-test analysis and Discriminant Function Analysis (DFA) showed that: (a) while all morphometric ratios demonstrated significant differences between M. nigrocinctus and M. gibbosus females, only selected ratios show significant difference in the males (Ca_L/W, Ch_L/W, Met-II_L/H, and Met- V_L/H); (b) among M. nigrocinctus, Adıyaman population was partly separated; …