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Designation Of A Neotype For Oeneis Melissa (Fabricius, 1775) And A Lectotype For Oeneis Polixenes (Fabricius, 1775) (Lepidoptera: Nymphalidae: Satyrinae), Crispin S. Guppy

The Taxonomic Report of the International Lepidoptera Survey

The Oeneis melissa (Fabricius, 1775) “neotype” designations of Lukhtanov and Eitschberger (2000; 2001) do not meet several critical requirements of the ICZN and are therefore invalid. In addition, the locality data is too vague to be taxonomically useful, their “neotype” specimen is a specimen of the taxon generally known as O. melissa semplei, and acceptance of their specimen as neotype would change the meaning of the taxon Oeneis melissa. Hence, I reject their neotype designation. A neotype Oeneis melissa (Fabricius, 1775) is designated from Nain, Labrador with careful consideration of the Articles of the International Code of Zoological …

Changes To North American Butterfly Names, Jing Zhang, Qian Cong, Jinhui Shen, Paul A. Opler, Nick V. Grishin

The Taxonomic Report of the International Lepidoptera Survey

We obtained and analyzed whole genome shotgun sequences of all 845 species of butterflies recorded from Canada and the United States. Genome-scale phylogenetic trees constructed from the data reveal several nonmonophyletic genera and suggest improved classification of species included in these genera. Here, these changes are formalized and 2 subgenera are described: Amblyteria Grishin, subgen. n. (type species Goniloba exoteria Herrich-Schäffer, 1869, parent genus Amblyscirtes Scudder, 1872), and Coa Grishin, subgen. n. (type species Hesperia baracoa Lucas, 1857, parent genus Polites Scudder, 1872). Furthermore, we resurrect 3 genera and 2 subgenera from synonymy, change the rank of 6 currently used …

Expanded Phenotypic Diagnoses For 24 Recently Named New Taxa Of Hesperiidae (Lepidoptera), Nick V. Grishin

The Taxonomic Report of the International Lepidoptera Survey

Expanded diagnoses by phenotypic characters for the 24 new taxa named in the article "Genomes of skipper butterflies reveal extensive convergence of wing patterns" by Li, W., Cong, Q., Shen, J., Zhang, J., Hallwachs, W., Janzen, D.H. and Grishin, N.V., 2019 and published in the Proceedings of the National Academy of Sciences of the United States of America on March 15, 2019 are provided and illustrated. More detailed diagnoses will help identifying these phylogenetic groups by their wing patterns and shapes and other morphological characters including the structures of antennae and genitalia using this single publication, instead of obtaining the …

Cercyonis Pegala Agawamensis (Satyridae): A New Butterfly Subspecies From The Coastal Salt Marshes Of The Northeastern United States Of America, Matthew D. Arey, Alex Grkovich

The Taxonomic Report of the International Lepidoptera Survey

A new subspecies of the Common Wood Nymph (Cercyonis pegala) from the New England coastline in the northeastern region of the United States is described on the basis of phenotype, behavior, habitat, seasonality, flight period, and phenology. Agawamensis is univoltine as in all pegala. The newly described subspecies occurs almost entirely in coastal salt marshes and estuarine meadows, quite atypical as understood for Cercyonis pegala. The newly described subspecies agawamensis predominately prefers and occurs within large open salt marshes but can also be found in similar habitat along smaller tidal rivers, coastal inlets and streams

A Case Of Sympatric Celastrina Ladon (Cramer), Celastrina Lucia (W. Kirby) And Celastrina Neglecta (Edwards) (Lycaenidae: Polyommatinae) In Northern Virginia, With Additional Records Of C. Lucia In Virginia, Harry Pavulaan

The Taxonomic Report of the International Lepidoptera Survey

A case of fully sympatric Celastrina ladon, C. lucia, and early spring brood C. neglecta is documented at a site in northern Virginia. Observations indicate that all three species occupy the same habitat, fly during the same flight period and utilize the same hostplant with no evidence of hybridization. C. ladon and C. lucia are obligate univoltines while C. neglecta is multivoltine. A later flight (second brood) of Celastrina neglecta at the same site utilizes eriophyid mite-induced leaf galls on the very same host tree species. Additional Virginia records of C. lucia are documented.

Larval Host Plants Of Enodia Anthedon, Satyrodes Appalachia And S. Eurydice In Vermont, Usa, David J. Hoag

The Taxonomic Report of the International Lepidoptera Survey

Field observation and captive rearing was used to clarify larval host plant use in Enodia anthedon, Satyrodes appalachia, and S. eurydice in Vermont, USA. In nature S. appalachia larvae were found on Carex lacustris, C. lupulina, C. gracillima, and C. tuckermanii. Enodia anthedon in nature was found to oviposit on grass and on Carex lacustris. Larvae of E. anthedon were found on C. lacustris and C. lupulina. In captivity, all but two E. anthedon larvae chose Carex over grass. Both E. appalachia and E. anthedon …

The Nomenclatural Status Of Ten Names In The Genus Pieris (Lepidoptera: Pieridae), Crispin S. Guppy, Norbert G. Kondla

The Taxonomic Report of the International Lepidoptera Survey

The nomenclatural status of ten names in the genus Pieris are reviewed. The dates and locations of publication of pseudobryoniae Verity, 1908 and adalwinda Fruhstorfer, 1909 are corrected; they are infrasubspecific unavailable names. The names pseudobryoniae Barnes and McDunnough, 1916 and arctica Barnes and McDunnough, 1916 are newly identified as available species-group names with at least four and six syntypes respectively. A lectotype of pseudobryoniae Barnes and McDunnough is selected, with the type locality being Nulato, Alaska; browni Eitschberger, 1983 is a subjective synonym. A lectotype of arctica Barnes and McDunnough is designated, with the type locality being northern Norway. …

Observations On Anthocharis Julia Browningi And Anthocharis Thoosa Thoosa Including Tension Zones Near Nephi, Juab County, Utah, Todd Stout

The Taxonomic Report of the International Lepidoptera Survey

Two visibly distinct taxa of the sara orangetip fly sympatrically and synchronically in a zone northeast of Nephi, Juab County, Utah. This paper discusses how these two taxa interact based upon six character sets—adult phenotype, adult male behavior, distribution, habitat, larval coloration, and pupal shape.

Tips On Collecting And Rearing Immatures Of 375 Butterfly And Skipper Taxa, Jacque Wolfe, Jack Harry, Todd Stout

The Taxonomic Report of the International Lepidoptera Survey

Rearing techniques are discussed for 375 different butterfly and skipper taxa from Utah and beyond.

Natural Life Histories Of Alaska Colias (Lepidoptera: Pieridae), Jack L. Harry

The Taxonomic Report of the International Lepidoptera Survey

Eight species of the genus Colias of Alaska were studied during the years of 2005, 2006, and 2007. Natural life histories were determined, immature stages described, measured, and photographed. Scanning Electron Microscope (SEM) pictures of the micropyle of five species are included.

Yosemite Butterflies (Text), 2nd Edition, Kenneth E. Davenport

The Taxonomic Report of the International Lepidoptera Survey

The author, Kenneth E, Davenport, has collected or observed butterflies extensively in the Yosemite region (at least 90 collecting or observation days) since August 1963. Information for this study began with Davenport obtaining a copy of Garth and Tilden’s 1963 Yosemite Butterflies in 1970. The annual Season Summaries for California published in the News of the Lepidopterists’ Society between 1975 and 2006 have been checked for records. The common English and scientific names used in this publication mostly follow the names used by the International Lepidoptera Survey (TILS) except in a few cases.

The following individuals contributed information: George T. …

A Description Of A New Subspecies Of Lycaena Phlaeas (Lycaenidae: Lycaeninae) From Montana, United States, With A Comparative Study Of Old And New World Populations, Steve Kohler

The Taxonomic Report of the International Lepidoptera Survey

The Palaearctic, Oriental, and Ethiopian Region subspecies of Lycaena phlaeas are briefly discussed. A more detailed account of the North American subspecies is presented, and a new subspecies, L. p. weberi, from the Sweet Grass Hills, Montana is described. The possibility that the eastern United States subspecies hypophlaeas was introduced from the Old World is discussed; however no conclusion can be reached with certainty. The relationship between Old World and New World subspecies of L. phlaeas is discussed. Evidence presented supports the treatment of New World populations as subspecies of L. phlaeas.

Bionomic Studies Of Papilio Demoleus Linnaeus, The Citrus Butterfly, (Lepidoptera: Papilionidae), From Lower Sindh, Pakistan, Arshad Munir, Nikhat Yasmin Siddiqui, Muhammad A. Rafi, Harry Pavulaan, David Wright

The Taxonomic Report of the International Lepidoptera Survey

Biological parameters of Papilio demoleus Linnaeus, the Citrus Butterfly, in Tando Muhammad Khan, lower Sindh, Pakistan are presented. This major agricultural pest was reared on Murraya koenigii (L.) Spreng. (Curry Leaf) and ten different varieties of Citrus sp. (Citrus) plants. The length of the adult mating period ranged 5 to 8 hours. Following mating, females laid 27.7 ± 3.3 eggs. Eggs hatched in 1.5 to 7 days. Mean durations of larval instars were 1.8 ± 0.1 (minimum) and 2.9 ± 0.1 (maximum) days respectively. Pupal duration was 5.5 to 20 days. Biotic factors including predators [praying mantis, ants, Common Garden …

A New Subspecies Of Colias Gigantea From Arctic Alaska (Pieridae), Jack L. Harry

The Taxonomic Report of the International Lepidoptera Survey

A new subspecies of Colias gigantea Strecker from the 'north slope' of Alaska is described.

The Yosemite Butterflies: Color Plates, Kenneth E. Davenport, Norbet G. Kondla, Charles Grisham, Howard Grisham

The Taxonomic Report of the International Lepidoptera Survey

This compilation of butterfly illustrations for the Yosemite region is intended primarily as a supplement to the text publication on Yosemite butterflies (Davenport, 2004, and Davenport, 2007, 2nd edition). Taxonomic interpretations and names follow these documents. Common names and authors of the scientific names are given in the text publication. This color plates publication also functions as a stand-alone document for naturalists and lepidopterists with need for reference illustrations only.

Butterflies were selected from Ken Davenport’s private collection to illustrate most species and subspecies for the Yosemite region. If good quality specimens were available from within the region, we often …

Celastrina Serotina (Lycaenidae: Polyommatinae): A New Butterfly Species From The Northeastern United States And Eastern Canada, Harry Pavulaan, David M. Wright

The Taxonomic Report of the International Lepidoptera Survey

A new light-venter Azure species, Celastrina serotina, is described from the northeastern United States and eastern Canada. The flight period of this univoltine spring species is interpolated (allochrony) between its sympatric congeners, C. lucia Auctorum (not Kirby) and C. neglecta (Edwards). In this regard it is similar to C. neglectamajor Opler & Krizek in the southern Appalachians and C. idella Wright & Pavulaan on the New Jersey coastal plain. In a significant portion of its range, C. serotina larvae feed on eriophyid mite-formed galls on the upper surface of Black Cherry Prunus serotina Ehrh. …

Taxonomic Analysis Of The Genus Megisto (Satyridae) In The Eastern United States, Part I: Types, Type Localities, And Taxonomic Relationships Of The Available Names, Ronald R. Gatrelle

The Taxonomic Report of the International Lepidoptera Survey

The genus Megisto Hübner, 1819 in the eastern United States and southeastern Canada is currently comprised of the original epithets Papilio eurytus Fabricius, 1775, Papilio cymela Cramer, 1777, Papilio eurytris, Fabricius, 1793, and Neonympha eurytris viola Maynard, 1891. Because the eastern Megisto is considered by some, including the present author, to contain two or more sibling species, the types and type localities of each of these names were studied. Where no type was found and no type locality fixed, typification was established by lectotypification or neotypification and a type locality was fixed. The taxonomic relationship of these taxa were …

Immature Stages Of Estigmene Acrea From Guatemala (Lepidoptera: Arctiidae), Jack Harry

The Taxonomic Report of the International Lepidoptera Survey

The immature stages of Estigmene acrea Drury are described and figured. The larval host utilized was Brassica oleracea L. var. botrytis (cauliflower).

Immature Stages Of Colias Johanseni From Arctic Canada (Lepidoptera: Pieridae), Jack Harry

The Taxonomic Report of the International Lepidoptera Survey

The immature stages of Colias johanseni are described and figured. The probable larval host plant is Hedysarum mackenziei, a common arctic legume.

Immature Stages Of Colias Occidentalis Sullivani From Oregon (Lepidoptera: Pieridae), Jack Harry

The Taxonomic Report of the International Lepidoptera Survey

The immature stages of Colias occidentalis sullivani Hammond and McCorkle are described and figured. The larval host is Lathyrus rigidus White.

The Yosemite Butterflies (Text), And The Yosemite Butterflies (Text): Supplement Of Additional Information, Collections And Corrections For Volume 5, No. 1, Kenneth E. Davenport

The Taxonomic Report of the International Lepidoptera Survey

The current checklist covers the 170 known species and additional 48 subspecies known to occur in the Yosemite region, California, United States. This represents a total of 218 taxa (some being undescribed and unnamed) now recognized in the study area. This compares with Garth and Tilden’s 1963 survey which recognized 134 species and 16 additional subspecies, a total of 150 taxa. In addition to those, six additional species reported from the region are “questionable or doubtful” and another six species occur just outside the defined region.

The supplement includes additional information for the benefit of those who obtained the first …

Description Of A Multilevel Cryptic New Species Of Phyciodes (Nymphalidae: Melitaeinae) From The Southern Appalachian Mountains, Ronald R. Gatrelle

The Taxonomic Report of the International Lepidoptera Survey

Phyciodes incognitus is described as a new species from the medium to high elevation hardwood forests of the southern Appalachian Mountains, type locality: Duncan Ridge Road, 3,700 ft., Union County, Georgia, United States. Phyciodes incognitus is verified from four colonies in Union County, Georgia, United States, and Clay and Macon counties, North Carolina, United States. It is projected to range, in suitable habitat, as far north as montane Pennsylvania, United States (indicated by photos). Phyciodes incognitus possesses a unique set of character traits that are virtually identical to both P. tharos and P. cocyta but at differing character …

A Concise Update Of The Information Provided In The Butterflies Of Southern California (1973) By Thomas C. Emmel And John F. Emmel, Kenneth E. Davenport

The Taxonomic Report of the International Lepidoptera Survey

This paper’s purpose is to bring together in one resource a concise, but thorough, report on the current status of butterflies discussed in the Butterflies of Southern California by Thomas C. Emmel and John F. Emmel, published by the Natural History Museum of Los Angeles County, Los Angeles, California, United States. 1973. This update deals primarily with new species or subspecies, taxonomic matters, and new distributional information. The family order follows A Catalogue / Checklist of the Butterflies of America North of Mexico by Clifford D. Ferris, editor, published by the Lepidopterists’ Society as Memoir No. 3, 1989. Species order …

A Taxonomic Examination Of Harkenclenus Titus (Lycaenidae: Theclinae) In The Eastern United States: Recognition And Resolution Of Taxonomic Problems By The Delineation Of Old Names, Naming Of A Long Recongnized (But Heretofore Undescribed) Subspecies, And Description Of A Wide Ranging New Subspecies, Ronald R. Gatrelle

The Taxonomic Report of the International Lepidoptera Survey

Harkenclenus is retained per dos Passos 1970 as a genus distinct from Satyrium. The names titus and mopsus have long been considered to apply to two subspecies. These are demonstrated to be synonymous as both names are based on southeastern United States populations. The holotype of titus (Natural History Museum London) is examined and its type locality established as Screven County, Georgia, United States. A neotype is established for the synonymic name mopsus, type locality, Screven County, Georgia, United States. This leaves the long recognized northern subspecies without a name; it is described as new subspecies Harkenclenus titus …

A Case For Taxonomic Recognition Of The Taxon Enodia Anthedon Borealis A. H. Clark (Satyridae), Alex Grkovich, Harry Pavulaan

The Taxonomic Report of the International Lepidoptera Survey

Clark (1936) described the taxa anthedon and borealis both as new subspecies of Enodia portlandia (Fabricius, 1781). Clark described borealis as the northernmost phenotypically different taxon in this group. Anthedon later became recognized as a distinct species with borealis as a subspecies of it. Masters (1971) characterized E. anthedon borealis as displaying several subtle phenotypic differences from nominotypical anthedon and also noted significant differences in behavior and habitat and reinforced the continued recognition of borealis as a valid subspecies. However, a number of publications after 1971 generally failed to recognize subspecific status for borealis, either ignoring borealis entirely …

A Taxonomic Review Of Chlosyne Ismeria With Description Of A New Subspecies From The Southern Appalachian Mountains, Ronald R. Gatrelle

The Taxonomic Report of the International Lepidoptera Survey

The taxon Boisduval and Le Conte described in 1833 as Melitaea ismeria is examined as a historical entity and confirmed as a sister subspecies of the taxon Chlosyne ismeria nycteis. The ismeria populations in the southern Appalachian mountains (United States) are described as a new subspecies: Chlosyne ismeria obsoleta. In facies, this new subspecies is most similar to the western subspecies Chlosyne ismeria drusius in that it is dark and contrasting in its dorsal markings. It is known to range from the mountain region of extreme northern Georgia north in the Appalachians to near southwestern Virginia, United States. …

A Subspecific Assessment Of The Genus Hesperia (Hesperiinae) In Eastern North America (Part I: The South): New Subspecies Of Hesperia Metea, Hesperia Sassacus And Hesperia Meskei, Ronald R. Gatrelle, Marc C. Minno, Alex Grkovish

The Taxonomic Report of the International Lepidoptera Survey

Hesperia meskei pinocayo Gatrelle and Minno is described from Big Pine Key, Monroe County, Florida, United States. This subspecies is thought to be nearly extinct. Its mainland status is uncertain. It is characterized by bi-colored ventral hindwings with more prominent ventral hindwing spots. Hesperia metea intermedia Gatrelle is described from Pickens County, South Carolina, United States. Its range is from northwestern South Carolina, United States west through Mississippi, United States and probably into Missouri, United States. Males have the least dorsal fulvous of the metea subspecies; ventrally they are well marked. Females are similar to subspecies licinus. Hesperia sassacus …

Butterflies Of The North Okanagan, British Columbia, Canada, David L. Threatful

The Taxonomic Report of the International Lepidoptera Survey

An annotated list of 106 species of butterflies known to occur in the north Okanagan area of British Columbia, Canada is presented. Information is provided on flight phenology, elevation and habitat for each species. Field work over the past 100 years shows that this is one of the premiere areas in Canada for butterfly species diversity. More than one third of all known Canadian butterflies have been found in this small area.

Fixation Of Type Locality For Lycaena Acmon Westwood And Characterization Of The Species And Its Distribution, Paul A. Opler

The Taxonomic Report of the International Lepidoptera Survey

Lycaena acmon Westwood, 1852 is based on a painting and plate legend in Westwood and Hewitson’s Genera of Diurnal Lepidoptera. The specimen illustrated was located in the British Museum Natural History and is the holotype by monotypy. The accompanying plate legend gives “California” as the type locality. Because the butterfly is a member of a complex of species, now considered in the genus Plebejus (Opler and Warren, 2003), that requires much systematic study, and some confusion exists on the identity of L. acmon, it is necessary to fix a more specific type locality, to characterize the species, …

Taxonomy And Distribution Of Butterflies (Papilionoidea) Of The Skardu Region, Pakistan, Muhammad Abbas, Muhammad Ather Rafi, Mian Inayatullah, Muhammad Rafique Khan, Harry Pavulaan

The Taxonomic Report of the International Lepidoptera Survey

A study was conducted in six different localities around Skardu, Pakistan to document the butterfly fauna of that region. The study revealed that 16 species in 5 families and 14 genera occur in the area. The families include: Papilionidae (represented only by the genus Parnassius); Pieridae (genera Pieris, Pontia, and Colias are represented); Lycaenidae (genera Lycaena, Everes, Aricia, Plebejus, Zizeeria, and Zizina are represented); Nymphalidae (only two genera, Aglais and Cynthia are represented); and Satyridae (represented by the genera Pararge and Maniola).