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2017

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Articles 181 - 210 of 401

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Verification Of Echosounder Measurements Of Thickness And Spatial Distribution Of Kelp Forests, Huamei Shao, Kenji Minami, Hokuto Shirakawa, Takashi Maeda, Toshiaki Ohmura, Yoshikazu Fujikawa, Norishige Yotsukura, Masahiro Nakaoka, Kazushi Miyashita Jun 2017

Verification Of Echosounder Measurements Of Thickness And Spatial Distribution Of Kelp Forests, Huamei Shao, Kenji Minami, Hokuto Shirakawa, Takashi Maeda, Toshiaki Ohmura, Yoshikazu Fujikawa, Norishige Yotsukura, Masahiro Nakaoka, Kazushi Miyashita

Journal of Marine Science and Technology–Taiwan

Acoustic methods can be used to assess seaweed meadows. The accuracy of the methods is a key factor in the estimation of seaweed distribution and conditions. We obtained and verified thickness and spatial distribution values measured using an echosounder. We determined the thickness of seaweed growing in the coastal waters off Higashidoori-mura, Aomori, Japan on June 5-6, 2013. Acoustic data were collected using an on-board quantitative echosounder at 120 kHz. The thickness was also directly measured at 14 points. The root mean square error (RMSE) of the thickness determined by the acoustic and direct methods was calculated. A survey to …


Library And Information Services To The Visually Impaired Persons, Junaid Rayini Jun 2017

Library And Information Services To The Visually Impaired Persons, Junaid Rayini

Library Philosophy and Practice (e-journal)

Visually impaired people have the same information needs as sighted people. Just as sighted people might read a newspaper, listen to a CD or download electronic information from the Internet; visually impaired people also want access to relevant information in their chosen accessible format. Developing an efficient library service for print-disabled people is extremely important, because there are significantly fewer books available commercially in accessible formats compared to what is published in print for the general public. Libraries have a moral obligation to make information available to all categories of users regardless of their gender, age, race, political affiliation or …


Relative Selection Strength: Quantifying Effectsize In Habitat- And Step-Selection Inference, Tal Avgar, Subhash R. Lele, Jonah L. Keim, Mark S. Boyce Jun 2017

Relative Selection Strength: Quantifying Effectsize In Habitat- And Step-Selection Inference, Tal Avgar, Subhash R. Lele, Jonah L. Keim, Mark S. Boyce

Wildland Resources Faculty Publications

Habitat-selection analysis lacks an appropriate measure of the ecological significance of the statistical estimates-a practical interpretation of the magnitude of the selection coefficients. There is a need for a standard approach that allows relating the strength of selection to a change in habitat conditions across space, a quantification of the estimated effect size that can be compared both within and across studies. We offer a solution, based on the epidemiological risk ratio, which we term the relative selection strength (RSS). For a "used-available" design with an exponential selection function, the RSS provides an appropriate interpretation of the magnitude of the …


Wolbachia-Host Interactions And The Implications To Insect Conservation And Management, Amy Michelle Truitt Jun 2017

Wolbachia-Host Interactions And The Implications To Insect Conservation And Management, Amy Michelle Truitt

Dissertations and Theses

Parasitic reproductive endosymbionts are emerging as formidable threats to insect biodiversity. Wolbachia are prevalent maternally inherited intra-cellular bacteria found in >50% of arthropod species. These symbiotic bacteria interact with their hosts in diverse ways, most often they alter host reproduction causing four conditions that all selectively favor infected females: feminization, male killing, parthenogenesis, and cytoplasmic incompatibility (CI). Furthermore, depending on strain-type and host genetic background, Wolbachia are known to affect insect behavior, expand or shift host thermal tolerance ranges, and confer anti-viral protection to their hosts. Because Wolbachia both reside in and are transmitted with host cell cytoplasm, mitochondria and …


Meiobenthic Invertebrates Community Associated With Aquatic Plant Ceratophyllum Demersum Salamiyat Irrigation Canal / North Baghdad, Prof. Dr. Haifa J. Jaweir, Rana Sabah Hassan Jun 2017

Meiobenthic Invertebrates Community Associated With Aquatic Plant Ceratophyllum Demersum Salamiyat Irrigation Canal / North Baghdad, Prof. Dr. Haifa J. Jaweir, Rana Sabah Hassan

Baghdad Science Journal

The aim of the present study is to study the meiobenthic invertebrate's community associated with the aquatic plant Ceratophyllum demersum in Al-Salamiyat irrigation canal / north Baghdad, with the chemical and physical parameters of the canal water, during the study period from September 2015 to May 2016. Two sites were chosen for sample collection, the first site (S1) at the beginning of the canal near it's connection with Tigris river, and the second site (S2) after 10 km from the first site. The chemico-physical analysis results revealed that the water temperature ranged from 10-30oC, and pH values ranged between 6.9-7.8, …


The Association Of Dietary Quality With Colorectal Cancer Among Normal Weight, Overweight And Obese Men And Women: A Prospective Longitudinal Study In The Usa, Rosalie A. Torres Stone, Molly E. Waring, Sarah L. Cutrona, Catarina I. Kiefe, Jeroan Allison, Chyke A. Doubeni Jun 2017

The Association Of Dietary Quality With Colorectal Cancer Among Normal Weight, Overweight And Obese Men And Women: A Prospective Longitudinal Study In The Usa, Rosalie A. Torres Stone, Molly E. Waring, Sarah L. Cutrona, Catarina I. Kiefe, Jeroan Allison, Chyke A. Doubeni

Sociology

Objective: Lower body mass index (BMI) and higher dietary quality reduce the risk of colorectal cancer (CRC). A full understanding of how these associations vary by sex and weight is lacking. Methods: We used data from the National Institutes of Health - American Association of Retired Persons (NIH)-AARP) Diet and Health Study for 398 458 persons who were 50-71 years old in 1995-1996 and followed through 2006. Exposures were dietary quality as reflected by the Mediterranean Diet, the Healthy Eating Index-2010 and the Dietary Approaches to Stop Hypertension score, stratified by BMI category. The outcome was CRC diagnosis from cancer …


Incorporating Place And Space: A Hierarchical Spatial Approach To Exploring Preventable Congestive Heart Failure Hospitalizations In New York City, Rachael Weiss Riley Jun 2017

Incorporating Place And Space: A Hierarchical Spatial Approach To Exploring Preventable Congestive Heart Failure Hospitalizations In New York City, Rachael Weiss Riley

Dissertations and Theses

Background: Faced with rising medical care costs, increasing prevalence, and widening health disparities, preventing congestive heart failure (CHF) hospitalizations is a central public health concern. Despite evidence of geographical clustering in preventable CHF admissions, there is a lack of research designed to examine spatial patterning of CHF and the local area neighborhood determinants that contribute to this variability. This study sought to assess and evaluate the importance of both space and place in analyzing preventable CHF hospitalizations and readmissions by applying appropriate statistical techniques, clarifying the assumption inherent in each method, and interpreting the findings within the context of existing …


Correlates Of Immune Defenses In Golden Eagle Nestlings, Benjamin M. Dudek, Julie A. Heath Jun 2017

Correlates Of Immune Defenses In Golden Eagle Nestlings, Benjamin M. Dudek, Julie A. Heath

Biology Faculty Publications and Presentations

An individual’s investment in constitutive immune defenses depends on both intrinsic and extrinsic factors. We examined how Leucocytozoon parasite presence, body condition (scaled mass), heterophil-to-lymphocyte (H:L) ratio, sex, and age affected immune defenses in golden eagle (Aquila chrysaetos) nestlings from three regions: California, Oregon, and Idaho. We quantified hemolytic-complement activity and bacterial killing ability, two measures of constitutive immunity. Body condition and age did not affect immune defenses. However, eagles with lower H:L ratios had lower complement activity, corroborating other findings that animals in better condition sometimes invest less in constitutive immunity. In addition, eagles with Leucocytozoon infections had higher …


A Comparative History Of Resurrection Plants, John Charles Ryan Jun 2017

A Comparative History Of Resurrection Plants, John Charles Ryan

CLCWeb: Comparative Literature and Culture

In his article "A Comparative Literary History of Resurrection Plants" John Charles Ryan assembles a comparative history of resurrection plants through textual analysis of early botanical commentaries, herbal references, prose, poetry, and other sources. Resurrection plants include a diverse range of botanical species, typically of arid regions, that appear to come back to life after complete desiccation. Historical and contemporary observers—from sixteenth-century herbalist John Gerard to contemporary Australian poet John Kinsella—have expressed an abiding fascination for resurrection plants' capacity to survive harsh environmental conditions. The plants court their own deaths by paring down—then restoring—physiological processes in relation to shifting ecological …


The Impact Of The Medieval Climatic Anomaly On The Archaeology At Edwards Air Force Base, Jessica Amanda Porter-Rodriguez Jun 2017

The Impact Of The Medieval Climatic Anomaly On The Archaeology At Edwards Air Force Base, Jessica Amanda Porter-Rodriguez

Electronic Theses, Projects, and Dissertations

A series of severe and prolonged droughts occurred throughout the Northern Hemisphere between approximately 1150 BP to 600 BP. This phenomenon is referred to as the Medieval Climatic Anomaly and has been shown to have differentially impacted various regions of the world. Previous studies have suggested causal links between the Medieval Climatic Anomaly and observed culture change.

The goal of this study was to examine the Antelope Valley region of the Mojave Desert for evidence of impacts on human populations related to the Medieval Climatic Anomaly. To achieve this goal, a sample selection of archaeological sites was chosen from lands …


Revisioning Ecology: Oikos Logos And A Human Niche, James Inabinet May 2017

Revisioning Ecology: Oikos Logos And A Human Niche, James Inabinet

Coastlines

No abstract provided.


Acute Memory And Psychotomimetic Effects Of Cannabis And Tobacco Both ‘Joint’ And Individually: A Placebo-Controlled Trial, C. Hindocha, T. P. Freeman, J. X. Xia, N. D. C. Shaban, H. V. Curran May 2017

Acute Memory And Psychotomimetic Effects Of Cannabis And Tobacco Both ‘Joint’ And Individually: A Placebo-Controlled Trial, C. Hindocha, T. P. Freeman, J. X. Xia, N. D. C. Shaban, H. V. Curran

Publications and Research

Background. Cannabis and tobacco have contrasting cognitive effects. Smoking cannabis with tobacco is prevalent in many countries and although this may well influence cognitive and mental health outcomes, the possibility has rarely been investigated in human experimental psychopharmacological research.

Method. The individual and interactive effects of cannabis and tobacco were evaluated in 24 non-dependent cannabis and tobacco smokers in a randomized, placebo-controlled, double-blind, 2 (cannabis, placebo) × 2 (tobacco, placebo) crossover design. Verbal memory (prose recall), working memory (WM) performance including maintenance, manipulation and attention (N-back), psychotomimetic, subjective and cardiovascular measures were recorded on each of four sessions.

Results. Cannabis …


Flood Risk Assessment Using The Updated Fema Floodplain Standard In The Ellicott City, Maryland, United States, Ranjeet Thakali, Ranjit Bhandari, Giles-Arnaud Arif-Deen Kandissounon, Ajay Kalra, Sajjad Ahmad May 2017

Flood Risk Assessment Using The Updated Fema Floodplain Standard In The Ellicott City, Maryland, United States, Ranjeet Thakali, Ranjit Bhandari, Giles-Arnaud Arif-Deen Kandissounon, Ajay Kalra, Sajjad Ahmad

Civil and Environmental Engineering and Construction Faculty Research

Every Year, flooding causes a calamitous impact on the people, economy, and environment all over the world. In recent years, the flood-related damages have been increasing in the United States regardless of several investments in the flood control measures. Floodplain mapping is an important tool for management that aids in the planning of infrastructures within the floodplain zone. With the magnifying effects of climate change on the hydrological cycle the study of floodplain is becoming a key tool in the water management. Federal Emergency Management Agency has recently updated their floodplain standard as per the presidential executive order 2015 on …


Needle Exchange Programs For The Prevention Of Hepatitis C Virus Infection In People Who Inject Drugs: A Systematic Review With Meta-Analysis, Stephen M. Davis, Shay Daily, Alfgeir L. Kristjansson, George A. Kelley, Keith Zullig, Adam Baus, Danielle Davidov, Melanie Fisher May 2017

Needle Exchange Programs For The Prevention Of Hepatitis C Virus Infection In People Who Inject Drugs: A Systematic Review With Meta-Analysis, Stephen M. Davis, Shay Daily, Alfgeir L. Kristjansson, George A. Kelley, Keith Zullig, Adam Baus, Danielle Davidov, Melanie Fisher

Clinical and Translational Science Institute

Background: Previous research on the effectiveness of needle exchange programs (NEP) in preventing hepatitis C virus (HCV) in people who inject drugs (PWID) has shown mixed findings. The purpose of this study was to use the meta-analytic approach to examine the association between NEP use and HCV prevention in PWIDs. Methods: Study inclusion criteria were (1) observational studies, (2) PWIDs, (3) NEP use, (4) HCV status ascertained by serological testing, (5) studies published in any language since January 1, 1989, and (6) data available for measures of association. Studies were located by searching four electronic databases and cross-referencing. Study quality …


An Ecological-Physiology Perspective On Seabird Responses To Contemporary And Historic Environmental Change, Alexis P. Will May 2017

An Ecological-Physiology Perspective On Seabird Responses To Contemporary And Historic Environmental Change, Alexis P. Will

Biological Sciences

The chapters included in this dissertation implement an ecological-physiology approach to understanding how long-lived marine organisms, using seabirds as a model, respond to changes in the environment. Many seabird populations are governed by bottom-up processes, yet efforts to connect prey dynamics and parameters such as breeding performance often yield mixed results. Here I examined how individual foraging behavior and nutritional status change at the inter-annual, decadal, and multi-decadal scale. I validated that the concentration of the avian stress hormone in seabird feathers is indicative of their exposure to nutritional stress. I then used this technique to show that young seabirds …


Lake Trout (Salvelinus Namaycush) Otoliths As Indicators Of Past Climate Patterns And Growth In Arctic Lakes, Eric S. Torvinen May 2017

Lake Trout (Salvelinus Namaycush) Otoliths As Indicators Of Past Climate Patterns And Growth In Arctic Lakes, Eric S. Torvinen

Fisheries

No abstract provided.


Mitigating The Conflict Between Pitfall-Trap Sampling And Conservation Of Terrestrial Subterranean Communities In Caves, Peter Kozel, Tanja Pipan, Nina Šajna, Slavko Polak, Tone Novak May 2017

Mitigating The Conflict Between Pitfall-Trap Sampling And Conservation Of Terrestrial Subterranean Communities In Caves, Peter Kozel, Tanja Pipan, Nina Šajna, Slavko Polak, Tone Novak

International Journal of Speleology

Subterranean habitats are known for their rich endemic fauna and high vulnerability to disturbance. Many methods and techniques are used to sample the biodiversity of terrestrial invertebrate fauna in caves, among which pitfall trapping remains one of the most frequently used and effective ones. However, this method has turned out to be harmful to subterranean communities if applied inappropriately. Traditionally, pitfall traps have been placed in caves solely on the ground. Here we present an optimized technique of pitfall trapping to achieve a balance between sampling completeness and minimal disturbance of the fauna in the cave. Monthly we placed traps …


Assessing The Effects Of Ecotourism On The Developing World: A Galapagos Case Study, Ife Adelona May 2017

Assessing The Effects Of Ecotourism On The Developing World: A Galapagos Case Study, Ife Adelona

Student Theses 2015-Present

Over the past decade, ecological tourism in the Galapagos has gained popularity. The local population of people living in the Galapagos is around 30,000; this includes, permanent residents, temporary residents, and immigrants. Tourist visitation within the 5 human inhabited islands is 200,000 people annually. The romantic idea of a pristine natural environment entices humans from across the globe to visit this World Heritage site. However, the growing attraction of this natural landmark has led to resource exploitation and the commodification of natural landscapes. In recent years, the Galapagos has developed its infrastructure in order to sustain the influx of tourists …


Larval Rockfish Diversity Abundance Dynamics In Association With A Marine Protected Area In The Southern California Bight, Dustin Chen May 2017

Larval Rockfish Diversity Abundance Dynamics In Association With A Marine Protected Area In The Southern California Bight, Dustin Chen

Theses

Marine protected areas (MPAs) are areas aimed at protecting natural and cultural resources that are often proposed as a way to alleviate the effects of overfishing on populations of targeted species; these areas have varying degrees of fishing and recreational use restrictions. In order to assess MPA efficacy, it is important to determine the mechanisms by which the presence of MPAs affect reproductive output within and potentially even beyond their boundaries. I attempted to address this through studying the responses of rockfish (Sebastes spp.) larval abundances to the presence of the Cowcod Conservation Areas (CCAs) located within the Southern California …


Use Of First Pelvic, Second Pelvic, And Fourth Anal Rays For Estimating Age Of Muskellunge (Esox Masquinongy), Marinda R. Cornett May 2017

Use Of First Pelvic, Second Pelvic, And Fourth Anal Rays For Estimating Age Of Muskellunge (Esox Masquinongy), Marinda R. Cornett

Honors Theses

While cleithra are the most popularly used calcified structure used to estimate age of Muskellunge (Esox masquinongy), the use of fin rays as an age estimation structure has been investigated as a nonlethal alternative. My objectives were to: (1) to compare the accuracy and precision of estimating age using first pelvic, second pelvic, and fourth anal rays from 46 known age Muskellunge and (2) to analyze a larger sample (n = 120) of fourth anal rays to estimate age. Ages were estimated blindly by three readers independently from all available structures. Different analyses of precision and accuracy had conflicting results, …


Application Of Anthromes To Frame Scenario Planning For Landscape-Scale Conservation Decision Making, Dainee Gibson, John Quinn May 2017

Application Of Anthromes To Frame Scenario Planning For Landscape-Scale Conservation Decision Making, Dainee Gibson, John Quinn

Biology Publications

Complexities in the rates and patterns of change necessitate the consideration of alternate futures in planning processes. These scenarios, and the inputs and assumptions used to build them, should reflect both ecological and social contexts. Considering the regional landscape as an anthrome, a priori, assumes human needs and institutions have a fundamental role and place in these futures, but that institutions incorporate ecological limits in decision making. As a case study of conservation scenario planning under the anthrome paradigm, we used a suite of InVEST models to develop and explore land use and land cover scenarios and to measure the …


A Guide To The Literature Of The Southwest, Lyle Saunders, Theo Cravenna May 2017

A Guide To The Literature Of The Southwest, Lyle Saunders, Theo Cravenna

New Mexico Quarterly

No abstract provided.


Hanoi Vietnam’S Peri-Urban Space And Its Impact On People’S Livelihoods, Collin V. Sumera May 2017

Hanoi Vietnam’S Peri-Urban Space And Its Impact On People’S Livelihoods, Collin V. Sumera

Sustainability and Social Justice

ABSTRACT

Hanoi Vietnam’s Peri-urban Space and Its Impact on People’s Livelihoods

Collin V. Sumera

Peri-urban is both a space and a process, yet there is no cookie cutter definition that may be applied to developing cities encountering this phenomenon. Each space is constructed differently, and each process has different actors. This paper examines the role that the Communist Party of Vietnam plays in the creation of peri-urban space in Hanoi, and how this space affects residential and former agricultural Land Use Right holder’s livelihood opportunities. The notion of the peri-urban space has not fully been developed for Hanoi, thus I …


Causes And Consequences Of Individual Phenotypic Differences In Brachyuran Crabs With A Focus On Behavior, Benjamin A. Belgrad May 2017

Causes And Consequences Of Individual Phenotypic Differences In Brachyuran Crabs With A Focus On Behavior, Benjamin A. Belgrad

Theses and Dissertations

Individual variation defines almost every morphological, physiological, and behavioral aspect of populations and is a fundamental component of many ecosystem processes. Recent work indicates that accounting for these individual differences can enhance our ability to predict community responses to environmental disturbances which is becoming increasingly important in an era of extraordinary global change. However, our understanding of how different individual characteristics are connected to each other and governed by the environment remains limited.

This study sought to evaluate the relationship between individual behavior, physiological condition, and local habitat for Brachyuran crabs as well as the subsequent strength of their predator-prey …


Phenology, Habitat Use, And The Impacts Of Wetland Management On Autumn Migrating Rails In Missouri, Auriel M.V. Fournier May 2017

Phenology, Habitat Use, And The Impacts Of Wetland Management On Autumn Migrating Rails In Missouri, Auriel M.V. Fournier

Graduate Theses and Dissertations

Rails (Family: Rallidae) are among the least studied birds in North America, in large part due to their elusive nature. As a wetland-dependent species, understanding the timing of their migration and their habitat needs during migration is especially important since management needs to be timed to balance the needs of many species. I developed and verified a new distance sampling based nocturnal ATV spotlight survey because traditional call-broadcast surveys are not effective during autumn migration because of the drop off in call rate after the breeding season. These surveys allow us to ask point-level questions about what habitats rails select …


Heterogeneity Of Avian Breeding Habitat On Grazing Lands Of The Northern Great Plains, Maggi S. Sliwinski May 2017

Heterogeneity Of Avian Breeding Habitat On Grazing Lands Of The Northern Great Plains, Maggi S. Sliwinski

School of Natural Resources: Dissertations, Theses, and Student Research

Native rangelands in the Great Plains are largely privately owned and used for beef production. Vegetation heterogeneity is important for maintaining biodiversity, but private land may be more homogenous than desired. My research had two components: 1) to examine whether a variety of grazing strategies created vegetation heterogeneity in a large, intact rangeland, and 2) to understand beef producers’ attitudes about vegetation heterogeneity.

First, I sampled vegetation structure, composition, and bird abundance at multiple plots on eleven management units in Cherry County, Nebraska. Units were managed with commonly used grazing strategies (e.g., short-duration grazing and season-long continuous grazing). I examined …


Niche Modeling For The Giant Panda, Ailuropoda Melanoleuca, And The Original Panda, Ailurus Fulgens: Habitat Preferences And Evolutionary Consequences, Lauren M. Lyon May 2017

Niche Modeling For The Giant Panda, Ailuropoda Melanoleuca, And The Original Panda, Ailurus Fulgens: Habitat Preferences And Evolutionary Consequences, Lauren M. Lyon

Electronic Theses and Dissertations

The well-known symbol for conservation, the giant panda bear, and the original red panda have been forced into remote habitats due to anthropogenic disturbance, making ecological study difficult. Therefore the first known species distribution model was created to predict the most likely areas of occurrence within the known range of these elusive animals. These models were then projected onto North America and evaluated against existing breeding programs. Additionally, the close proximity of the Gray Fossil Site and the discovery of the most complete fossil red panda specimens in the world allowed ecomorphological comparisons between the modern red panda, Ailurus fulgens …


Cranial Morphological Distinctiveness Between Ursus Arctos And U. Americanus, Benjamin James Hillesheim May 2017

Cranial Morphological Distinctiveness Between Ursus Arctos And U. Americanus, Benjamin James Hillesheim

Electronic Theses and Dissertations

Despite being separated by millions of years of evolution, black bears (Ursus americanus) and brown bears (Ursus arctos) can be difficult to distinguish based on skeletal and dental material alone. Complicating matters, some Late Pleistocene U. americanus are significantly larger in size than their modern relatives, obscuring the identification of the two bears. In the past, fossil bears have been identified based on differences in dental morphology or size. This study used geometric morphometrics to look at overall differences in cranial shape and used step-wise discriminant analysis to identify specific characters that distinguish cranial morphology between …


Plant Community Responses To Bison Reintroduction Within Montana's Northern Great Plains, Nicholas A. Mcmillan May 2017

Plant Community Responses To Bison Reintroduction Within Montana's Northern Great Plains, Nicholas A. Mcmillan

All Theses

In western North America, the rapid loss of native grazers, abiotic disturbance processes (e.g., fire), and human-driven land conversion or degradation have resulted in steadily declining biodiversity and rangeland health. Keystone species restoration and their potential to increase rangeland health are common justifications for bison (Bison bison L.) reintroduction projects throughout North America's Great Plains. Similarly, many other conservation groups also justify the removal of livestock from traditionally cattle-grazed rangelands by citing potential increases in rangeland health or community dynamics following years of heavy use by cattle. Our objective was to assess how ten-years of bison reintroduction and livestock removal …


The Evolution Of Androgen Receptor Expression And Social Behavior In Anolis Lizards, Miguel Angel Webber May 2017

The Evolution Of Androgen Receptor Expression And Social Behavior In Anolis Lizards, Miguel Angel Webber

Biology Honors Theses

Steroid hormones have a well-studied influence on behavior, but circulating levels of testosterone alone cannot fully predict levels of social, androgenic behaviors. Androgen receptor (AR) expression may bridge the gap between circulating androgens and the muscles that control social behavior: species with higher rates of behavior should have higher levels of AR protein in the nuclei of the muscles that control these behaviors. In anole lizards, the ceratohyoid (CH) muscle extends the dewlap, a colorful throat fan used in social displays, and the retractor penis magnus (RPM) muscle retracts the intromittent organ after copulation. I observed social behavior in the …