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Comparison Of Enrichment Interaction Duration Between Male And Female Panthera Leo Leo, Maegen G. Meldrum

Undergraduate Honors Thesis Projects

African lion (Panthera leo leo) populations have declined significantly over the past century within their natural habitats, increasing the importance of effective management strategies for individuals in human care. Environment enrichment is a critical tool for promoting species typical behavior in human care; however, enrichment practices are often generalized and may not account for sex-specific differences. This study examined how sex, enrichment types, behaviors, and time of day influenced enrichment interaction duration in a small pride of African lions (n = 4) housed at the Columbus Zoo and Aquarium. Two enrichment types were evaluated: cognitive/occupational (CO, a manipulable barrel) …

Taxonomy, Distribution And Ecology, Biology, Nutritional Composition, And Conservation Status Of Critically Endangered Lake Chala Tilapia Oreochromis Hunteri (Günther, 1889): Implications For Aquaculture Development, Jonathan Munguti, Mavindu Muthoka, Domitila Kyule, Esther Magondu, Jacob O. Iteba, Kevin Obiero, Paul Orina, Menaga Meenakshisundaram, Tanga M. Chrysantus

All Peer-Reviewed Publications

Oreochromis hunteri Günther, 1889, commonly known as Lake Chala tilapia, is an endemic cichlid restricted to the small crater lake, Lake Chala, on the border of Kenya and Tanzania. This review synthesizes existing knowledge on the taxonomy, and phylogenetic placement, distribution and ecology, and biology including nutritional composition, and conservation status of O. hunteri and identifies knowledge gaps to assess its potential for aquaculture development. The knowledge gaps include its reproductive biology, population structure, and adaptive capacity to environmental and climatic variability. As the type species of the genus Oreochromis, O. hunteri is a unique, genetically isolated lineage within a …

Insect Farming Innovations: A Pathway To Self-Reliance In Agrifood Systems In Africa, Zewdu Abro, Menale Kassie, Dennis Beesigamukama, Shaphan Yong Chia, Ermias Engida Legesse, Subramanian Sevgan, Julius Ecuru, Chrysantus M. Tanga

All Peer-Reviewed Publications

Insect farming as an innovative approach to transforming agrifood systems

Insect farming has emerged as a transformative and sustainable approach to addressing the interconnected challenges of food insecurity, environmental degradation, and economic vulnerability in Africa’s agrifood systems. By converting organic waste into high-value products (protein-rich feed, frass fertilizers, and bioactive compounds), insect farming offers an efficient circular economy solution that bridges nutrition, livelihoods, and ecological resilience (Lalander et al. 2025; Mei et al. 2024; Barragán-Fonseca et al. 2022; Xia et al. 2021; Van Huis 2020; Barragán-Fonseca et al. 2017; Van Huis and Oonincx 2017; Yi et al. 2014). Almost all …

Ecological Intensification Of Maize-Kale Systems In Kenya By Biofertilization And Push-Pull Management Shifts Nematode Communities, Agnes Kiriga, Johnnie Van Den Berg, Frank Chidawanyika, Solveig Haukeland, Bianca Greyvenstein, Dennis Beesigamukama, Chrysantus M. Tanga, Sevgan Subramanian, Daniel M. Mutyambai

All Peer-Reviewed Publications

Plant-parasitic nematodes (PPNs) pose a persistent threat to vegetable production in Kenya, contributing to reduced yields and soil degradation. The management of PPNs in vegetables in Kenya has largely remained unaddressed. The integration of ecological intensification strategies such as the vegetable integrated push-pull (VIPP) system and the use of black soldier fly frass fertilizer (BSFF) is gaining traction in sustainable agriculture. The aim of this study was to determine the effects of integrating VIPP and BSFF on nematode abundance and diversity in maize-kale cropping systems. Field trials were conducted in two different agroecological zones in Kenya. Field treatments involved plots …

Brief Therapy Dog Interactions May Increase Self-Efficacy And Lower Perceived Stress And Cortisol In High School Students: A Preliminary Study, Dawn Melzer, Deirdre Yeater, Madison Derubis, Christiana Rose Kohlroser, Barbara J. Pierce

Psychology Faculty Publications

The present study examined changes in high schoolers’ perceived stress, general self-efficacy and cortisol levels after a brief interaction with a certified therapy dog. High school students completed self-report measures of perceived stress and general self-efficacy and provided salivary cortisol samples both before and after a 10-min interaction with a certified therapy dog. Following the interaction, students reported reductions in perceived stress scores, exhibited lower salivary cortisol concentrations and demonstrated increases in self-efficacy scores compared to before the interaction. Although higher self-efficacy scores were observed, general self-efficacy is generally regarded as a stable construct; therefore, these findings may reflect a …

Poly(A) Probe Hcr Rna-Fish Specifically Marks Pyriform Nurse Cells In The Brown Anole Lizard Ovary, Zoe B Griffin, Bonnie K Kircher, Richard R Behringer

Faculty, Staff and Student Publications

Hybridization chain reaction RNA-fluorescent in situ hybridization (HCR RNA-FISH) is a powerful and increasingly used method for visualizing gene expression in cells and tissues. A probe set against polyadenylated RNA (poly(A)) is often used as a positive control for RNA integrity and staining quality. While optimizing this technique in the ovary of the brown anole lizard (Anolis sagrei), we found that the poly(A) probe produced a strikingly specific and intense signal in pyriform cells, a specialized lizard-specific nurse cell type. This staining pattern was found in both whole-mount samples and paraffin sections, suggesting that poly(A) signal intensity can serve as …

Host Modulation Therapy In Periodontitis: From Established Therapies To Emerging Technologies, Aonjittra Phanrungsuwan, Jiaqi Huang, Neeraja Dharmaraj, Alejandra Cobos Perez, Omid Veiseh, Simon Young, Chun-Teh Lee

Faculty, Staff and Student Publications

Periodontitis is a biofilm-induced chronic inflammatory disease, characterized by gingival inflammation and alveolar bone loss. According to a national survey, approximately half of the U.S. adults are affected by periodontal disease. To effectively prevent and treat periodontitis, it is essential to address its underlying causes. The primary etiological factors include polymicrobial synergy and dysbiosis of the oral microbiota, and a dysregulated immune response. The standard therapeutic approach, mechanical removal of biofilm through debridement, sometimes demonstrates limited efficacy, particularly in cases of severe periodontitis, which may require adjunctive or additional therapy. Emerging evidence indicates that periodontal tissue destruction is initiated by …

Integrating Multi-Method Monitoring To Characterize Seasonal Habitat Use Of Green Sturgeon (Acipenser Medirostris) In Humboldt County, California, Olivia S. Boeberitz

Cal Poly Humboldt theses and projects

Green sturgeon (Acipenser medirostris) are a long-lived, anadromous, iteroparous species that spawns in large West Coast North American rivers but ranges widely in the coastal ocean. For coastal California Native American Tribes, green sturgeon are a vital part of their cultural heritage and traditions, and central to their food security. Two genetically distinct population segments (DPSs) of green sturgeon are recognized in the United States, Canada, and Mexico; the southern DPS is listed as “threatened” under the Endangered Species Act and the northern DPS is considered of “conservation concern.” Although green sturgeon are known to use large estuaries …

Factors Influencing Tree Vigor And Structural Failure Along Powerline Corridors In Northern California: A Multi-Species Analysis, David Justin Dorval

Cal Poly Humboldt theses and projects

Tree failures (e.g., stem breakage, uprooting, or branch drop) can pose significant risks to public safety, particularly along powerline corridors for the risk of power outages and wildfire ignitions. In California, tree-related powerline failures have led to severe ecological and human costs in recent years. Despite extensive research on tree failure in urban environments and post-disturbance contexts, comparatively little empirical work has examined where powerlines intersect largely unmanaged stands. This study evaluates tree failure trends across multiple forest types in Northern California by examining how tree-level characteristics influence recent growth and failure occurrence in regionally dominant tree species. Specifically, the …

Do California Rangeland Healthy Soil Practices Offer Co-Benefits For Grassland Birds?, Ximena O. Gil

Cal Poly Humboldt theses and projects

Rangelands are important landscapes in California that support a robust cattle industry and serve as vital grassland habitat for birds. Climate change is projected to impact rangelands, leading to economic and ecological deterioration. In California, the Healthy Soils Program (HSP) incentivizes climate-smart solutions to increase working land resiliency. While the HSP targets soil and range health for livestock, it may also benefit birds. We surveyed rangeland operators to examine whether the implementation and duration of specific range management practices, such as range seeding, prescribed grazing, and silvopasture, are associated with bird community responses, using avian point count data from Point …

Forest Management Options For The Wildland Urban Interface: Comparative Analysis Of Stand Development, Economics, And Forest Carbon Over 100 Years In Sierra Nevada Ponderosa Pine, Kassandra A. Mcdaniel

Cal Poly Humboldt theses and projects

This thesis presents a simulation study of various forest stand treatments and their long-term effects on ponderosa pine-dominated forests in the Sierra Nevada Mountain Range of the western USA. The goal was to create recommendations for forest management in the wildland urban interface (WUI) of the Tahoe National Forest and other locations with similar conditions. This study simulates current United States Forest Service (USFS) - Truckee Ranger District silvicultural treatment alternatives repeated over a 100-year period using the USFS Forest Vegetation Simulator (FVS). Proposed treatment alternatives include restoration, resilience, transition, conservation, and no management. Treatments are to be compared in …

Coho Salmon (Oncorhynchus Kisutch) Response To Habitat Restoration Through Space And Time: Monitoring Growth, Survival, And Movement In A Restored Habitat Network, Carter Perez Adamson

Cal Poly Humboldt theses and projects

Despite the decline of coho salmon (Oncorhynchus kisutch), Prairie Creek, a tributary of Redwood Creek in northern California, remains a coho salmon stronghold. The restoration of juvenile coho rearing habitat was identified as a recovery objective in this watershed by the National Marine Fisheries Service. Among the restoration projects addressing this objective are an ongoing project on Prairie Creek and its tributary Skunk Cabbage Creek, and a prior effort on Strawberry Creek (a tributary of Redwood Creek’s estuary). Standard monitoring methods for assessing project success have limitations and may be improved by measuring salmon use of restored habitats …

Relational Approaches To Fire Resilience: Designing A Living Shaded Fuel Break With Indigenous Knowledge Systems, Natalee Nicole Lopes

Cal Poly Humboldt theses and projects

Escalating wildfire severity across California reflects not only changing climatic conditions but also the cumulative impacts of colonial land management systems that have disrupted Indigenous fire stewardship and altered forest structure and function. This project examines how wildfire mitigation infrastructure, specifically linear fuel treatments, such as fuel breaks, can be reimagined through Indigenous relational frameworks of care-based land stewardship. Grounded in Happy Camp (Athithúfvuunupma), California, within the ancestral homelands of the Karuk Tribe along the mid-Klamath River, this research is carried out in collaboration with Karuk knowledge holders, centering Indigenous knowledge systems and priorities.

Using a mixed-methods approach that integrates …

Are There Enough Bees? Assessing Pollen Limitation In Native Eastern Washington Prairie Forbs, Alejandro Torres

EWU Masters Thesis Collection

Prairie ecosystems in eastern Washington have been severely degraded due to agricultural intensification, urbanization, and habitat fragmentation, leaving less than 0.1% of native prairie intact. As a result, these ecosystems have become some of the most endangered in the United States. One group of essential pollinators is the bees, which facilitate plant reproduction and contribute to the long-term persistence of restored plant communities. However, despite the presence of at least 257 native bee species in the region, the impacts of habitat fragmentation and restoration on pollinator activity and plant-pollinator networks remain poorly understood. For this study, we worked across Eastern …

Moth Assemblages Of Fort Leonard Wood, Missouri, Kimberly Anne Steese

Masters Theses

"Moths (Lepidoptera) are ecologically important insects that function as herbivores, pollinators, prey resources, and indicators of ecosystem integrity. Despite their ecological significance, moth communities in the Missouri Ozarks remain poorly documented, particularly within military-managed landscapes. This thesis examined patterns of moth diversity, assemblage structure, and environmental drivers at Fort Leonard Wood (FLW), Missouri, while also assessing the distribution and genetic diversity of species within the genus Papaipema.

Field surveys were conducted from 2019 through 2021 across prairie, savanna, forest, and restoration habitats at FLW using UV bucket traps, sheet trapping, and targeted larval surveys. Across all sampling efforts, 29,116 individual …

A Note On The Novel Passive Sampling In Environmental Dna (Edna) Collection Technique To Detect Yellow Mud Turtle (Kinosternon Flavescens): Field And Laboratory Approaches In Turtle Edna, Sakib Tahmid Rishan, Richard J. Kline, Md. Saydur Rahman

School of Integrative Biological & Chemical Sciences Faculty Publications

Environmental DNA (eDNA) is a powerful and sensitive method for monitoring aquatic and terrestrial organisms, offering insights into their distribution. Usually, eDNA is collected using a filter membrane through water filtration. However, the passive eDNA method can simplify sampling and broaden its scope, offering new insights for ecological surveillance. This study examined the efficacy of six different membrane filters in detecting the yellow mud turtle (Kinosternon flavescens) through a novel passive eDNA method. After being immersed in water for 12 h, only the mixed cellulose ester (MCE) membrane filter consistently demonstrated specific polymerase chain reaction (PCR) amplification using …

Reinterpreting Stochastic Optimal Control Under Ecological Uncertainty: Inferring Decision Urgency From Vegetation Biomass Dynamics, Komi Mensah Agboka, Tobias Landmann, Elfatih M. Abdel-Rahman

All Peer-Reviewed Publications

Stochastic optimal control provides a rigorous framework for systems subject to uncertainty, yet its operational use in ecological crisis contexts remains limited by interpretability. We reinterpreted a stochastic control formulation in which selected parameters emerged as indicators of decision urgency rather than normative preferences. Vegetation biomass was modeled as a stochastic stock subject to nonlinear loss driven by feeding pressure (e.g., desert locust activity) and multiplicative noise, with uncertainty represented by a time-varying volatility term that integrated extreme rainfall anomalies and conflict-related disruption. Rather than prescribing an optimal policy, we inverted the closed-form solution of the control problem to infer …

Wattage- And Composition-Dependent Ocular Surface Toxicity: An In Vitro Study On E-Cigarette Aerosol Exposure In Human Corneal Epithelial Cells, Tzu-Ling Lin

Masters Theses

As electronic cigarette consumption continues to escalate globally, its impact on non-respiratory mucosal surfaces, particularly the ocular surface, remains largely understudied. This study investigated the toxicological effects of e-cigarette aerosols using a human corneal epithelial cell (HCE-2) model. Our findings reveal that aerosol exposure induces a significant, dose-dependent reduction in cell viability, exacerbated by higher device wattage and specific flavor agents such as watermelon. Exposure was found to trigger intracellular oxidative stress and the activation of apoptotic pathways, leading to programmed cell death. Furthermore, the upregulation of proinflammatory cytokines suggests that vaping may predispose users to chronic inflammatory conditions of …

Investigation Of Perennial Clover Living Mulch And Reduced Tillage Effects On Weed Suppression, Crop Metrics, And Yield Of Organic Cabbage And Sweet Corn In The Northern Great Plains, Kristina Harms

Electronic Theses and Dissertations

Vegetable farmers often face weed management and soil health challenges, such as nitrate leaching, soil compaction, and top-soil loss. Sustainable agriculture is a forefront of innovation compared to conventional agriculture. Living mulches, non-cash crops grown alongside cash crops, have demonstrated weed suppression and impact on the promotion of sustainable agriculture. Yet studies have not evaluated established living mulch systems on cash crop quality and yield. This study conducted in 2023, 2024, and 2025 at SDSU Southeast Research Farm, Beresford, SD, USA evaluates the impact of established ‘Dynamite’ red clover (Trifolium pratense), ‘Domino’ white clover (T. repens), and ‘Aberlasting’ white × …

Biomechanical Integrity Of The Female Pelvic Floor: A Comparison Of Military And Civilian Women, Vladimir Egorov, Peter Takacs, Vincent Lucente, Heather Van Raalte, Joy A. Greer, Noune Sarvazyan

Department of Obstetrics & Gynecology Faculty Publications

Introduction

The health of the pelvic floor depends on the integrity of its muscles, connective tissue, ligaments, and their innervation. The female pelvic floor may be more vulnerable to repetitive heavy loads in a military environment than the male pelvic floor because of anatomical and physiological differences. The study aim was to identify and explore possible differences in the female pelvic floor conditions between active military and civilian women with similar demographics.To identify and explore possible differences in the conditions of the female pelvic floor between active military women and civilian women with similar demographics.

Materials and Methods

Military active …

Prepregnancy Diet Quality Index And Adverse Pregnancy Outcomes, Ummayhany Bharmal, George Saade, Philip Greenland, Brenda J. Smith, Andrea C. Kozai, Stephanie A. Fisher, Lauren Theilen, William Grobman, Noel Bairey Merz, Uma Reddy, Tetsuya Kawakita

Department of Obstetrics & Gynecology Faculty Publications

Introduction

Adverse pregnancy outcomes (APOs) are a major cause of maternal and neonatal morbidity.¹ Preconception diet quality is associated with APO risk,² but prior pregnancy-related indices had notable limitations.³ We developed the Prepregnancy Dietary Quality Index (PreP-DQI) to assess its association with APOs and cardiometabolic health.

Methods

This cohort study used data from the Nulliparous Pregnancy Outcomes Study: Monitoring Mothers-to-Be (NuMoM2b) and the NuMoM2b-Heart Health Study (NuMoM2b-HHS),⁴,⁵ excluding individuals who delivered before 23 weeks of gestation, had fetal anomalies, missing dietary data, or had implausible caloric intake (below 1st percentile or above 99th percentile). All participants in the original studies …

Selenium And Liver Steatosis And Fibrosis: Opposing Direct And Steatosis-Mediated Associations In A Large Cohort, Neda Rehan, Jubilee Benedict, Rehan Qayyum

Department of Medicine Faculty Publications

Background and Aims

Epidemiologic studies have reported conflicting associations between selenium exposure and liver disease. Whether selenium exerts opposing direct and steatosis-mediated effects on liver fibrosis is unknown. To evaluate the associations of blood selenium with elastography-defined hepatic steatosis and fibrosis and to determine whether hepatic steatosis mediates the relationship between selenium and fibrosis.

Methods

We conducted a cross-sectional analysis of adults aged >= 18 years in NHANES 2017-2023. Hepatic steatosis was assessed using the controlled attenuation parameter (CAP) and fibrosis using liver stiffness measurement (LSM). Blood selenium was categorized into quartiles. Survey-weighted multivariable linear regression and structural equation modeling …

Microplastics, Nanoplastics, And Obesity: Current Evidence And Biological Mechanisms, Maram Albandak, Sanam Rabeeah, Ahmad Mahdi Zahidi, David A. Johnson

Department of Medicine Faculty Publications

Obesity is a rapidly growing global health concern. The pathogenesis is complex and cannot be fully explained by lifestyle factors alone. The increasing attention towards this concern has finally been directed towards environmental contributors which may influence metabolic regulation. Microplastics and nano-plastics (MNPs) are ubiquitous environmental pollutants that have recently been detected in food, drinking water, air, and human biological samples, raising concerns about their potential role in metabolic disorders, including obesity. Chronic exposure to MNPs may interfere with metabolic homeostasis through multiple biological pathways. Emerging evidence, including both experimental and animal studies, reports alterations in lipid metabolism, body weight, …

Study Of Fire Regimes In Southwestern Australia Using Geospatial Techniques, Ana Do Carmo Carvalho

Theses: Doctorates and Masters

The Northern Jarrah Forest in Southwestern Australia (SWA), part of a global biodiversity hotspot, is home to fire-sensitive tree species such as marri (Corymbia calophylla) and jarrah (Eucalyptus marginata). As climate change intensifies drought and extreme fire events, understanding the drivers of fire severity is increasingly critical. This thesis analyses fire regimes within the Mundaring drinking water catchment to better understand the spatiotemporal complexity of fire severity in relation to fire history and environmental factors.

The thesis commences with a review of three Fire History Databases (FHDs) used in SWA, highlighting significant inconsistencies and data gaps across time and space. …

Limited Site-Specific Blue Carbon Recovery Following Tropical Seagrass Restoration In Thailand, Ratchanee Kaewsrikhaw, Milica Stankovic, Pere Masqué, Mathew A. Vanderklift, Tipamat Upanoi, Anchana Prathep

Research outputs 2022 to 2026

Seagrass restoration is gaining popularity as a nature-based method for climate change mitigation, owing to its effectiveness in capturing and storing organic carbon (Corg). However, demonstrating blue carbon additionality — defined as a measurable increase in carbon stocks directly attributable to restoration activities — remains challenging. This research assessed sedimentary Corg stocks and carbon accumulation rates in restored tropical seagrass meadows throughout Thailand. Despite successful restoration at multiple sites, sediment analysis showed little Corg accumulation, low Corg and organic matter content, with non-significant differences among restored seagrass areas, patchy natural meadows, and nearby bare sand areas. Limited sediment accumulation was …

Decline In Offspring Quantity But Not Quality From Successive Matings In Male Rainforest Drosophila, With No Evidence For Genetic Divergence In Male Mating Behaviour Along Elevational And Density Gradients, Andrew D. Saxon, Eleanor K. O'Brien, Natalie E. Jones, Jon Bridle

Research outputs 2022 to 2026

Fertilisation success is a major component of male fitness, meaning males should capitalise on all opportunities for mating. However, other sources of variation in fitness mean that males often evolve life histories that limit their ability to mate frequently. We quantified mating latency, mating duration, and offspring production in males of the tropical fly Drosophila birchii when presented with up to 4 females consecutively. Males were sourced from isofemale lines from the extremes of 2 elevational gradients (20–1100 m) that show substantial differences in population density, temperature, and humidity. Total offspring sired increased with number of matings achieved, demonstrating substantial …

Insights Into Site Fidelity Of A Low-Density Dugong Population Using Small-Drone Imagery And Photo-Identification, Roxana Newman, Ashwag Asseri, William David Paterson, Deni Porej, Harry Clark, Mishari Alghrair, Winston Cowie, David Charles Wells, Amanda Jane Hodgson

Research outputs 2022 to 2026

The feasibility of using unoccupied aerial vehicles (UAVs, ‘drones’) to collect high-resolution still and video imagery of dugongs (Dugong dugon) for photo-identification (photo-ID) in the Sindalah Archipelago, NEOM (area ~ 84 km²), within the northeast Red Sea, Saudi Arabia, was evaluated during opportunistic marine megafauna searches. Drone searches were conducted whenever possible over one year (January 2024 to January 2025). A total of 48 dugong observations were made during 91 days of drone flights, covering a cumulative distance of 5,962 km of flying. Of these, 35 (72.9%) were repeat sightings (range = 2 to 8 resights) of nine …

The Conservation Value Of Silvicultural Systems For Breeding And Post-Fledging Bird Communities In Northeastern North America, Christopher Andrew Liazos

Antioch University Dissertations & Theses

Forest bird community decline in the eastern United States is partly due to the lack of early-and-late-successional forests stemming from historic land use. Silviculture can create underrepresented forest age classes and structures to support avian communities during the breeding and post-fledging periods. In southwestern New Hampshire, we conducted a case study evaluating avian conservation value indices across silvicultural systems. Our goal was to observe how avian conservation value indexes and abundance differed between recently managed and unmanaged stands. We surveyed 215 breeding and 118 post-fledging points across 22 sites using point counts and vegetation surveys. Conservation values in the breeding …

Are There Ecological Consequences Of Urban Adaptation? A Test Of Eco-Evolutionary Dynamics In A Terrestrial Isopod (Oniscus Asellus), Aaron Yilmaz, Grace Bellino, Ryan A. Martin

Faculty Scholarship

Urbanization affects many aspects of the environment with the potential to alter both ecological and evolutionary processes. However, while examples of urban evolution are accumulating, tests of urban eco-evolutionary dynamics are rare. We carried out a reciprocal transplantation experiment using the terrestrial isopod Oniscus asellus to test how adaptation to the urban heat island influenced ecological processes. As isopods are important decomposers, we asked if leaf-litter decomposition differed among reciprocal transplant treatments and used a separate laboratory experiment to isolate the effects of temperature on rates of isopod leaf consumption. We found that leaf-litter decomposition was greater in urban habitats, …

Preliminary Analysis Of Soil Volatile Organic Compounds As Soil Health Indicators, Thorne Taul

Graduate Theses, Dissertations, and Problem Reports (ETD)

Over the past decade, soil health (SH) has gained popularity due to the novelty of sustainable soil use related to human wellbeing. Soil health describes the overall quality of soil and its capacity to function over time. As interest in SH grows, researchers continue to explore new soil health indicators that can improve knowledge on how soil health is assessed and managed. Some indicators directly reflect soil health status, and some indirectly relate to processes and functions occurring within the soil. Volatile organic compounds (VOCs) are naturally occurring compounds that play a role in multiple soil processes. They are associated …