Environmental Sciences Commons^™

Variable Helper Effects, Ecological Conditions, And The Evolution Of Cooperative Breeding In The Acorn Woodpecker, Walter D. Koenig, Eric L. Walters, Joseph Haydock

Biological Sciences Faculty Publications

The ecological conditions leading to delayed dispersal and helping behavior are generally thought to follow one of two contrasting scenarios: that conditions are stable and predictable, resulting in young being ecologically forced to remain as helpers (extrinsic constraints and the habitat saturation hypothesis), or that conditions are highly variable and unpredictable, leading to the need for helpers to raise young, at least when conditions are poor (intrinsic constraints and the hard life hypothesis). We investigated how variability in ecological conditions influences the degree to which helpers augment breeder fitness in the cooperatively breeding acorn woodpecker (Melanerpes formicivorus), a …